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<document id="445E30481926A2B92610C02B8B412DF7" ID-DOI="10.11646/zootaxa.3692.1.11" ID-GBIF-Dataset="8da33386-32c1-4ace-9b2c-6ddcb6a4459f" ID-ISSN="1175-5326" ID-Zenodo-Dep="247905" ID-ZooBank="A96187E4-6C35-4A64-BD39-4D29606653BF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460653410026" checkinUser="plazi" docAuthor="Schüller, Myriam &amp; Jirkov, Igor A." docDate="2013" docId="20155B09EF0EBF4BFF17FC60FE1FDD00" docLanguage="en" docName="zt03692p237.pdf" docOrigin="Zootaxa 3692 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anobothrus rubropaleatus Schüller &amp; Jirkov, 2013, n.sp." docType="treatment" docVersion="7" lastPageNumber="223" masterDocId="DC2C2371EF1FBF58FF80FF96FF98DA6A" masterDocTitle="New Ampharetidae (Polychaeta) from the deep Southern Ocean and shallow Patagonian waters" masterLastPageNumber="237" masterPageNumber="204" pageNumber="221" updateTime="1698344165278" updateUser="plazi">
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<mods:title id="2FE1AAABABE969459A10207115800A42">New Ampharetidae (Polychaeta) from the deep Southern Ocean and shallow Patagonian waters</mods:title>
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<mods:namePart id="8366E97AC165F5D12EE269B8B74C59F9">Jirkov, Igor A.</mods:namePart>
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<treatment id="20155B09EF0EBF4BFF17FC60FE1FDD00" ID-DOI="http://doi.org/10.5281/zenodo.6152272" ID-GBIF-Taxon="127601692" ID-Zenodo-Dep="6152272" LSID="urn:lsid:plazi:treatment:20155B09EF0EBF4BFF17FC60FE1FDD00" httpUri="http://treatment.plazi.org/id/20155B09EF0EBF4BFF17FC60FE1FDD00" lastPageId="19" lastPageNumber="223" pageId="17" pageNumber="221">
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<paragraph id="A803EA1FEF0EBF49FF17FC60FD81DE7A" blockId="17.[151,537,1014,1073]" box="[151,537,1014,1040]" pageId="17" pageNumber="221">
<heading id="F34B5D73EF0EBF49FF17FC60FD81DE7A" bold="true" box="[151,537,1014,1040]" fontSize="11" level="1" pageId="17" pageNumber="221" reason="1">
<emphasis id="9AC8360DEF0EBF49FF17FC60FD81DE7A" bold="true" box="[151,537,1014,1040]" pageId="17" pageNumber="221">
<taxonomicName id="6FBC919CEF0EBF49FF17FC60FE4FDE7A" ID-CoL="5V3XV" box="[151,471,1014,1040]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="17" pageNumber="221" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0EBF49FF17FC60FE4FDE7A" bold="true" box="[151,471,1014,1040]" italics="true" pageId="17" pageNumber="221">Anobothrus rubropaleatus</emphasis>
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<taxonomicNameLabel id="81FB8B76EF0EBF49FE5EFC60FD81DE7A" box="[478,537,1014,1040]" pageId="17" pageNumber="221" rank="species">n.sp.</taxonomicNameLabel>
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<paragraph id="A803EA1FEF0EBF49FF17FB8FFE8CDE5B" blockId="17.[151,537,1014,1073]" box="[151,276,1049,1073]" pageId="17" pageNumber="221">
<figureCitation id="3087F69AEF0EBF49FF17FB8FFE8CDE5B" box="[151,276,1049,1073]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="18.[151,250,1309,1332]" captionStartId-1="19.[151,250,1715,1738]" captionStartId-2="20.[151,250,1045,1068]" captionTargetBox-0="[154,1428,347,1274]" captionTargetBox-1="[151,1436,385,1667]" captionTargetBox-2="[417,1179,247,1009]" captionTargetId-0="figure@18.[151,1436,339,1277]" captionTargetId-1="figure@19.[151,1436,385,1668]" captionTargetId-2="figure@20.[417,1179,247,1009]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." captionText-1="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." captionText-2="FIGURE 12. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—notochaetae of TU- 8, Buncini of neuropodium of TU- 8, C—neuropodium of mid-body AU, D—pygidium." httpUri-0="https://zenodo.org/record/247915/files/figure.png" httpUri-1="https://zenodo.org/record/247916/files/figure.png" httpUri-2="https://zenodo.org/record/247917/files/figure.png" pageId="17" pageNumber="221">Figs 1012</figureCitation>
</paragraph>
<paragraph id="A803EA1FEF0EBF49FF17FBF7FC1FDE13" blockId="17.[151,1436,1121,2017]" box="[151,903,1121,1145]" pageId="17" pageNumber="221">
<typeStatus id="770754BDEF0EBF49FF17FBF7FF67DE13" box="[151,255,1121,1145]" pageId="17" pageNumber="221" type="holotype">Holotype</typeStatus>
: Knipovich 258,
<geoCoordinate id="CD888CD8EF0EBF49FE43FBF7FDBADE13" box="[451,546,1121,1145]" direction="south" orientation="latitude" pageId="17" pageNumber="221" precision="925" value="-34.566666">34°34S</geoCoordinate>
,
<geoCoordinate id="CD888CD8EF0EBF49FDADFBF7FD0CDE13" box="[557,660,1121,1145]" direction="west" orientation="longitude" pageId="17" pageNumber="221" precision="925" value="-53.233334">53°14W</geoCoordinate>
,
<quantity id="6F4447FAEF0EBF49FD1FFBF7FD40DE13" box="[671,728,1121,1145]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="221" unit="m" value="50.0">50 m</quantity>
[ZMH-26065].
</paragraph>
<paragraph id="A803EA1FEF0EBF49FF47FB10FE0BDEAB" blockId="17.[151,1436,1121,2017]" pageId="17" pageNumber="221">
<typeStatus id="770754BDEF0EBF49FF47FB10FEACDEF4" box="[199,308,1158,1182]" pageId="17" pageNumber="221" type="paratype">Paratypes</typeStatus>
: Knipovich 258,
<geoCoordinate id="CD888CD8EF0EBF49FD86FB13FDFDDEF7" box="[518,613,1157,1181]" direction="south" orientation="latitude" pageId="17" pageNumber="221" precision="925" value="-34.566666">34°34S</geoCoordinate>
,
<geoCoordinate id="CD888CD8EF0EBF49FDF5FB13FD45DEF7" box="[629,733,1157,1181]" direction="west" orientation="longitude" pageId="17" pageNumber="221" precision="925" value="-53.233334">53°14W</geoCoordinate>
,
<quantity id="6F4447FAEF0EBF49FD6DFB13FCB3DEF7" box="[749,811,1157,1181]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="221" unit="m" value="50.0">50 m</quantity>
(
<specimenCount id="BEBA2196EF0EBF49FCBEFB13FC47DEF7" box="[830,991,1157,1181]" pageId="17" pageNumber="221" type="generic">64 specimens</specimenCount>
,
<date id="DC02CCDFEF0EBF49FC70FB13FB8FDEF7" box="[1008,1047,1157,1181]" pageId="17" pageNumber="221">1 x</date>
SEM, complete and incomplete) [ZMH-2606626067].
</paragraph>
<paragraph id="A803EA1FEF0EBF49FF47FB5BFEF2DF63" blockId="17.[151,1436,1121,2017]" pageId="17" pageNumber="221">Diagnosis: The species is characterized by its bright red paleae even in preserved material, and the lack of notopodia in TS-3.</paragraph>
<paragraph id="A803EA1FEF0EBF49FF47FA83FB26DF3B" blockId="17.[151,1436,1121,2017]" pageId="17" pageNumber="221">
Description:
<typeStatus id="770754BDEF0EBF49FED8FA83FE58DF47" box="[344,448,1301,1325]" pageId="17" pageNumber="221" type="holotype">Holotype</typeStatus>
<quantity id="6F4447FAEF0EBF49FE46FA83FD9FDF47" box="[454,519,1301,1325]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="221" unit="mm" value="6.0">6 mm</quantity>
long and 0.5 wide, with 13 AUs, complete. Complete
<typeStatus id="770754BDEF0EBF49FBE7FA80FB4ADF44" box="[1127,1234,1302,1326]" pageId="17" pageNumber="221" type="paratype">paratypes</typeStatus>
<quantity id="6F4447FAEF0EBF49FB58FA83FAADDF47" box="[1240,1333,1301,1325]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="17" pageNumber="221" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">28 mm</quantity>
long and about
<quantity id="6F4447FAEF0EBF49FF5CFAAFFEFBDF3B" box="[220,355,1337,1361]" metricMagnitude="-4" metricUnit="m" metricValue="3.9999999999999996" metricValueMax="7.0" metricValueMin="1.0" pageId="17" pageNumber="221" unit="mm" value="0.39999999999999997" valueMax="0.7" valueMin="0.1">0.10.7 mm</quantity>
wide, with 1213 AUs (
<figureCitation id="3087F69AEF0EBF49FDF6FAAFFD5FDF3B" box="[630,711,1337,1361]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="17" pageNumber="221">Fig. 10</figureCitation>
A). Colour in ethanol white, paleae light red.
</paragraph>
<paragraph id="A803EA1FEF0EBF4AFF47FACBFE78DA9D" blockId="17.[151,1436,1121,2017]" lastBlockId="18.[151,1436,151,247]" lastPageId="18" lastPageNumber="222" pageId="17" pageNumber="221">
Prostomium trilobed, middle lobe anteriorly rounded, without ridges or eyes (
<figureCitation id="3087F69AEF0EBF49FBBFFACBFB0EDF1F" box="[1087,1174,1373,1397]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="17" pageNumber="221">Figs 10</figureCitation>
B, 11A). Lower lip not enlarged. Presence of nuchal organs unresolved. Buccal tentacles all retracted. TS-2 bearing about 812 reddish paleae, positioned laterally directly below branchiophore ridge (
<figureCitation id="3087F69AEF0EBF49FCE6FA33FC2DDFD7" box="[870,949,1445,1469]" captionStart="FIGURE 11" captionStartId="19.[151,250,1715,1738]" captionTargetBox="[151,1436,385,1667]" captionTargetId="figure@19.[151,1436,385,1668]" captionTargetPageId="19" captionText="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="17" pageNumber="221">Fig. 11</figureCitation>
A, B). TS-2 laterally visible as a prominent lobe from which the paleae originate, otherwise more or less fused to TS-3. TS-3 achaetous. Paleal base distinctly stouter than best developed notochaetae of subsequent thoracic segments. Paleae flat, saber-like, width of paleae abruptly decreasing in distal third, from there chaetae fine, tapering to a pointed tip (
<figureCitation id="3087F69AEF0EBF49FB01F987FB4EDC43" box="[1153,1238,1553,1577]" captionStart="FIGURE 11" captionStartId="19.[151,250,1715,1738]" captionTargetBox="[151,1436,385,1667]" captionTargetId="figure@19.[151,1436,385,1668]" captionTargetPageId="19" captionText="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="17" pageNumber="221">Fig. 11</figureCitation>
B). Four pairs of branchiae (
<figureCitation id="3087F69AEF0EBF49FE94F9A3FEFFDC27" box="[276,359,1589,1613]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="17" pageNumber="221">Fig. 10</figureCitation>
C). Anterior three pairs of branchiophores attached to each other across dorsum, arranged in an almost straight line forming a prominent dorsal ridge. Forth branchial pair attached behind the second outermost pair and connected to notopodia of TC-2 (TS-5). Outermost branchial pair connected with notopodia of TC-1 (TS- 4). Diameters of all branchial scars almost equal. All branchiae lost, branchial surface structure unknown. 14 TCs, 1st pair of notopodia present in TS-4, well developed. Notochaetae limbate (
<figureCitation id="3087F69AEF0EBF49FB81F95EFBCBDC8A" box="[1025,1107,1736,1760]" captionStart="FIGURE 11" captionStartId="19.[151,250,1715,1738]" captionTargetBox="[151,1436,385,1667]" captionTargetId="figure@19.[151,1436,385,1668]" captionTargetPageId="19" captionText="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="17" pageNumber="221">Fig. 11</figureCitation>
C). 12 TUs, starting in TC-3 (TS-6). Thoracic neuropodia large tori, slightly decreasing in size towards the posterior. [Thoracic uncini with a main fang surmounted by 4 to 5 transverse rows of 2 teeth (
<figureCitation id="3087F69AEF0EBF49FCABF886FCE3DD42" box="[811,891,1808,1832]" captionStart="FIGURE 11" captionStartId="19.[151,250,1715,1738]" captionTargetBox="[151,1436,385,1667]" captionTargetId="figure@19.[151,1436,385,1668]" captionTargetPageId="19" captionText="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="17" pageNumber="221">Fig. 11</figureCitation>
D).] Circular band present in anterior half of TU- 2 (TC-4, TS-7) [hardly visible in most specimens] (
<figureCitation id="3087F69AEF0EBF49FD6AF8A1FCA5DD25" box="[746,829,1847,1871]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="17" pageNumber="221">Fig. 10</figureCitation>
A, C, D). Notopodia of 5th to last TC slightly shifted dorsally and connected by a prominent ridge (
<figureCitation id="3087F69AEF0EBF49FD27F8CBFD63DD1F" box="[679,763,1885,1909]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="17" pageNumber="221">Fig. 10</figureCitation>
A). Chaetae and uncini of 5th to last TU not modified (
<figureCitation id="3087F69AEF0EBF49FAEEF8CBFF2BDDF3" captionStart="FIGURE 12" captionStartId="20.[151,250,1045,1068]" captionTargetBox="[417,1179,247,1009]" captionTargetId="figure@20.[417,1179,247,1009]" captionTargetPageId="20" captionText="FIGURE 12. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—notochaetae of TU- 8, Buncini of neuropodium of TU- 8, C—neuropodium of mid-body AU, D—pygidium." httpUri="https://zenodo.org/record/247917/files/figure.png" pageId="17" pageNumber="221">Fig. 12</figureCitation>
A, B). 13 AUs, neuropodia of AU-1 and AU-2 of similar
<typeStatus id="770754BDEF0EBF49FCCEF814FCE6DDF0" box="[846,894,1922,1946]" pageId="17" pageNumber="221">type</typeStatus>
to thoracic neuropodia, only slightly erect tori. [Uncini not observed in AU-1 and AU-2 under SEM.] Abdominal neuropodia from AU-3 distinct, prolonged erect pinnules with uncini in marginal position (
<figureCitation id="3087F69AEF0EBF49FD0FF85FFD7DDD8B" box="[655,741,1993,2017]" captionStart="FIGURE 12" captionStartId="20.[151,250,1045,1068]" captionTargetBox="[417,1179,247,1009]" captionTargetId="figure@20.[417,1179,247,1009]" captionTargetPageId="20" captionText="FIGURE 12. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—notochaetae of TU- 8, Buncini of neuropodium of TU- 8, C—neuropodium of mid-body AU, D—pygidium." httpUri="https://zenodo.org/record/247917/files/figure.png" pageId="17" pageNumber="221">Fig. 12</figureCitation>
C). [Abdominal uncini with dental formula with main fang surmounted by a transverse row of 78 teeth, surmounted by numerous small teeth.] Rudimentary abdominal notopodia absent. All thoracic and abdominal noto- and neuropodia without cirri. Pygidium with two lateral oval to cirriform papillae (
<figureCitation id="3087F69AEF0DBF4AFEEBFF49FE24DA9D" box="[363,444,223,247]" captionStart="FIGURE 12" captionStartId="20.[151,250,1045,1068]" captionTargetBox="[417,1179,247,1009]" captionTargetId="figure@20.[417,1179,247,1009]" captionTargetPageId="20" captionText="FIGURE 12. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—notochaetae of TU- 8, Buncini of neuropodium of TU- 8, C—neuropodium of mid-body AU, D—pygidium." httpUri="https://zenodo.org/record/247917/files/figure.png" pageId="18" pageNumber="222">Fig. 12</figureCitation>
D).
</paragraph>
<caption id="FCC3BA97EF0DBF4AFF17FA8BFBDCDF38" httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="18" pageNumber="222" targetBox="[154,1428,347,1274]" targetPageId="18">
<paragraph id="A803EA1FEF0DBF4AFF17FA8BFBDCDF38" blockId="18.[151,1436,1309,1362]" pageId="18" pageNumber="222">
<emphasis id="9AC8360DEF0DBF4AFF17FA8BFEB0DF59" bold="true" box="[151,296,1309,1332]" pageId="18" pageNumber="222">FIGURE 10.</emphasis>
<taxonomicName id="6FBC919CEF0DBF4AFEB7FA88FDD3DF5E" box="[311,587,1310,1332]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0DBF4AFEB7FA88FDD3DF5E" box="[311,587,1310,1332]" italics="true" pageId="18" pageNumber="222">Anobothrus rubropaleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0DBF4AFDDAFA89FD12DF5E" box="[602,650,1311,1332]" pageId="18" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
(paratype, Knipovich 258, ZMH-26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view.
</paragraph>
</caption>
<paragraph id="A803EA1FEF0DBF4AFF47FA16FAA0DFD1" blockId="18.[151,1437,1407,2008]" pageId="18" pageNumber="222">
A pair of nephridial papillae positioned slightly apart from each other behind branchial ridge dorsally on TS-5 (TC-2) (
<figureCitation id="3087F69AEF0DBF4AFF74FA35FEDCDFD1" box="[244,324,1443,1467]" captionStart="FIGURE 10" captionStartId="18.[151,250,1309,1332]" captionTargetBox="[154,1428,347,1274]" captionTargetId="figure@18.[151,1436,339,1277]" captionTargetPageId="18" captionText="FIGURE 10. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26067) A—habitus lateral view, Bprostomium, dorso-frontal view, C—anterior end, dorsal view, D—anterior end, lateral view." httpUri="https://zenodo.org/record/247915/files/figure.png" pageId="18" pageNumber="222">Fig. 10</figureCitation>
C) [under SEM, additional papillae observed dorsally associated with notopodia of TU-3].
</paragraph>
<paragraph id="A803EA1FEF0DBF4AFF47FA51FEA9DC6E" blockId="18.[151,1437,1407,2008]" pageId="18" pageNumber="222">Tube membranous, covered loosely with fine sediment and detritus, brightly stained in methylene blue and methyl green.</paragraph>
<paragraph id="A803EA1FEF0DBF4BFF47F999FD64DB76" blockId="18.[151,1437,1407,2008]" lastBlockId="19.[151,1436,151,284]" lastPageId="19" lastPageNumber="223" pageId="18" pageNumber="222">
Remarks: All
<typeStatus id="770754BDEF0DBF4AFEECF986FE4FDC42" box="[364,471,1552,1576]" pageId="18" pageNumber="222" type="paratype">paratypes</typeStatus>
are either minute in size or incomplete. In complete specimens the distinguishing characters of the species are difficult to observe due to the small size. Hence, a large but incomplete specimen in good condition is chosen as
<typeStatus id="770754BDEF0DBF4AFE7FF9C1FDF9DC05" box="[511,609,1623,1647]" pageId="18" pageNumber="222" type="holotype">holotype</typeStatus>
here to display all important characters for species recognition. A characteristic trait of
<taxonomicName id="6FBC919CEF0DBF4AFE0DF9EAFD88DCFE" box="[397,528,1660,1684]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="genus">
<emphasis id="9AC8360DEF0DBF4AFE0DF9EAFD88DCFE" box="[397,528,1660,1684]" italics="true" pageId="18" pageNumber="222">Anobothrus</emphasis>
</taxonomicName>
is a circular whitish band anterior of parapodia of TU-2 or TU-3. While most species bear this feature in TU-3, the newly described species,
<taxonomicName id="6FBC919CEF0DBF4AFCDEF909FB84DCDD" box="[862,1052,1695,1719]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0DBF4AFCDEF909FB84DCDD" box="[862,1052,1695,1719]" italics="true" pageId="18" pageNumber="222">A. rubropaleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0DBF4AFBA4F936FBC2DCDD" box="[1060,1114,1696,1719]" pageId="18" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
, bears it in TU-2, which has only been reported for
<taxonomicName id="6FBC919CEF0DBF4AFE2DF952FC13DCB1" authority="Desbruyeres, 1978" authorityName="Desbruyeres" authorityYear="1978" box="[429,907,1731,1756]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="laubieri">
<emphasis id="9AC8360DEF0DBF4AFE2DF952FD0DDCB6" box="[429,661,1732,1756]" italics="true" pageId="18" pageNumber="222">Anobothrus laubieri</emphasis>
(Desbruyères, 1978)
</taxonomicName>
and
<taxonomicName id="6FBC919CEF0DBF4AFC4CF952FBC8DCB6" box="[972,1104,1732,1756]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="wilhelmi" status="sp. nov.">
<emphasis id="9AC8360DEF0DBF4AFC4CF952FBC8DCB6" box="[972,1104,1732,1756]" italics="true" pageId="18" pageNumber="222">A. wilhelmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0DBF4AFBDDF952FB0ADCB1" box="[1117,1170,1732,1755]" pageId="18" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
The most outstanding character of
<taxonomicName id="6FBC919CEF0DBF4AFEA2F971FE47DC95" box="[290,479,1767,1791]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0DBF4AFEA2F971FE47DC95" box="[290,479,1767,1791]" italics="true" pageId="18" pageNumber="222">A. rubropaleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0DBF4AFE66F97EFD83DC95" box="[486,539,1768,1791]" pageId="18" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
however is the unique reddish colouration of the paleae. In shape they resemble those of
<taxonomicName id="6FBC919CEF0DBF4AFF79F89AFD6DDD49" authority="Schuller, 2008" authorityName="Schuller" authorityYear="2008" box="[249,757,1803,1828]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="pseudoampharete">
<emphasis id="9AC8360DEF0DBF4AFF79F89AFDD2DD4E" box="[249,586,1804,1828]" italics="true" pageId="18" pageNumber="222">Anobothrus pseudoampharete</emphasis>
Schüller, 2008
</taxonomicName>
with their medium size, stout base and an abruptly tapering distal quarter with a fine but pronounced tip. Under SEM, a simple accessory chaetae dorsal to the paleae was observed (
<figureCitation id="3087F69AEF0DBF4AFE8BF8C5FEC2DD01" box="[267,346,1875,1899]" captionStart="FIGURE 11" captionStartId="19.[151,250,1715,1738]" captionTargetBox="[151,1436,385,1667]" captionTargetId="figure@19.[151,1436,385,1668]" captionTargetPageId="19" captionText="FIGURE 11. Anobothrus rubropaleatus n. sp. (paratype, Knipovich 258, ZMH- 26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU- 9, D—uncini of neuropodium of TU- 9." httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="18" pageNumber="222">Fig. 11</figureCitation>
B). The surface of this chaeta was covered by regularly scattered holes, that were also present (but less numerous) on the paleal surface. This
<typeStatus id="770754BDEF0DBF4AFDFEF8EEFD36DDFA" box="[638,686,1912,1936]" pageId="18" pageNumber="222">type</typeStatus>
of chaeta has never been described for the genus before and only further observation of material under SEM will help to resolve its significance as a taxonomic character.
<taxonomicName id="6FBC919CEF0DBF4AFF17F829FE26DDBD" box="[151,446,1983,2007]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0DBF4AFF17F829FE26DDBD" box="[151,446,1983,2007]" italics="true" pageId="18" pageNumber="222">Anobothrus rubropaleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0DBF4AFE44F856FE61DDBD" box="[452,505,1984,2007]" pageId="18" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
together with
<taxonomicName id="6FBC919CEF0DBF4AFD1AF829FBB4DDBD" authority="Fauchald, 1972" authorityName="Fauchald" authorityYear="1972" box="[666,1068,1983,2008]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="18" pageNumber="222" phylum="Annelida" rank="species" species="mancus">
<emphasis id="9AC8360DEF0DBF4AFD1AF829FCE1DDB2" box="[666,889,1983,2008]" italics="true" pageId="18" pageNumber="222">Anobothrus mancus</emphasis>
Fauchald, 1972
</taxonomicName>
are the only species of the genus with only 14TCs. Additionally, among the examined material, a few specimens bear only 11 neuropodia on one side, starting in TC-4 (TS-7). While uncini were not observed in AU-1 and AU-
<quantity id="6F4447FAEF0CBF4BFBB8FF2AFBFDDABE" box="[1080,1125,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="19" pageNumber="223" unit="in" value="2.0">2 in</quantity>
the species, under SEM, a surface structure that might resemble uncini in development was observed in midline of the neuropodia of AU-1, however, the true identity of this structure is unknown.
</paragraph>
<caption id="FCC3BA97EF0CBF4BFF17F925FEE5DD6D" httpUri="https://zenodo.org/record/247916/files/figure.png" pageId="19" pageNumber="223" targetBox="[151,1436,385,1667]" targetPageId="19">
<paragraph id="A803EA1FEF0CBF4BFF17F925FEE5DD6D" blockId="19.[151,1436,1715,1799]" pageId="19" pageNumber="223">
<emphasis id="9AC8360DEF0CBF4BFF17F925FEB9DCA3" bold="true" box="[151,289,1715,1738]" pageId="19" pageNumber="223">FIGURE 11.</emphasis>
<taxonomicName id="6FBC919CEF0CBF4BFEAAF922FDA0DCA0" box="[298,568,1716,1738]" class="Polychaeta" family="Ampharetidae" genus="Anobothrus" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="223" phylum="Annelida" rank="species" species="rubropaleatus" status="sp. nov.">
<emphasis id="9AC8360DEF0CBF4BFEAAF922FDA0DCA0" box="[298,568,1716,1738]" italics="true" pageId="19" pageNumber="223">Anobothrus rubropaleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81FB8B76EF0CBF4BFDC1F923FDEADCA0" box="[577,626,1717,1738]" pageId="19" pageNumber="223" rank="species">n.sp.</taxonomicNameLabel>
(paratype, Knipovich 258, ZMH-26066) SEM A—prostomium, dorso-frontal view, B—paleae, arrow—chaeta of unknown origin associated with paleae, C—notochaetae of TU-9, D—uncini of neuropodium of TU-9.
</paragraph>
</caption>
<paragraph id="A803EA1FEF0CBF4BFF47F8B8FE1FDD00" blockId="19.[151,1436,1838,1898]" pageId="19" pageNumber="223">Etymology: The name refers to the bright reddish colour of the paleae that remains present even in ethanolpreserved specimens.</paragraph>
</subSubSection>
</treatment>
</document>