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<document id="9EAE26F2C219D605383816B1F5823C57" ID-CLB-Dataset="24105" ID-DOI="10.5852/ejt.2019.585" ID-GBIF-Dataset="dca608e8-fccf-4c1c-b8df-ef0c28e1d518" ID-Zenodo-Dep="3660259" ID-ZooBank="181B6FBA-ED75-4BB4-84C4-FB512B794749" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581251975682" checkinUser="plazi" docAuthor="Ebersole, Jun A., Cicimurri, David J. &amp; Stringer, Gary L." docDate="2019" docId="18174D41FF42FF68FDFC9E8C4C8609BD" docLanguage="en" docName="ejt-585_ebersole_cicimurri_stinger.pdf.imf" docOrigin="European Journal of Taxonomy 585" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Signata nicoli Dante &amp; Frizzell in Frizzell &amp; Dante 1965" docType="treatment" docVersion="18" lastPageNumber="200" masterDocId="E42E3539FF85FFAFFFC69F344D4B0F37" masterDocTitle="Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths" masterLastPageNumber="274" masterPageNumber="1" pageNumber="200" updateTime="1698783570407" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DC060E92B6DE8A948D5B672B97B184B1">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<mods:namePart id="9C473B54BABFF6D64BB13AC6494ED549">Ebersole, Jun A.</mods:namePart>
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<mods:namePart id="D5595E6323C85CF50CA0D9AC158FC575">Cicimurri, David J.</mods:namePart>
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<mods:title id="A96755628A2352C71520345EBAF083D9">European Journal of Taxonomy</mods:title>
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<treatment id="18174D41FF42FF68FDFC9E8C4C8609BD" ID-DOI="http://doi.org/10.5281/zenodo.3664406" ID-GBIF-Taxon="162072901" ID-Zenodo-Dep="3664406" LSID="urn:lsid:plazi:treatment:18174D41FF42FF68FDFC9E8C4C8609BD" httpUri="http://treatment.plazi.org/id/18174D41FF42FF68FDFC9E8C4C8609BD" lastPageNumber="200" pageId="199" pageNumber="200">
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<heading id="CB494B3BFF42FF68FDFC9E8C4EB20EE4" box="[570,1017,439,467]" centered="true" fontSize="11" level="2" pageId="199" pageNumber="200" reason="2">
<taxonomicName id="57BE87D4FF42FF68FDFC9E8C4EB20EE4" authority="Frizzell &amp; Dante, 1965" authorityName="Dante &amp; Frizzell in Frizzell &amp; Dante" authorityYear="1965" box="[570,1017,439,467]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FDFC9E8C4FA80EE5" bold="true" box="[570,739,439,466]" italics="true" pageId="199" pageNumber="200">Signata nicoli</emphasis>
<bibRefCitation id="F42F81A6FF42FF68FD2C9E8C4EB20EE4" author="Frizzell D. L. &amp; Dante J. H." box="[746,1017,440,467]" pageId="199" pageNumber="200" pagination="368 - 718" refId="ref142032" refString="Frizzell D. L. &amp; Dante J. H. 1965. Otoliths of some early Cenozoic fishes of the Gulf Coast. Journal of Paleontology 39 (4): 368 - 718." type="journal article" year="1965">Frizzell &amp; Dante, 1965</bibRefCitation>
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</heading>
</paragraph>
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<paragraph id="9001FC57FF42FF68FD189EE84E1E0EC1" blockId="199.[570,1017,439,502]" box="[734,853,476,502]" pageId="199" pageNumber="200">
<figureCitation id="0885E0D2FF42FF68FD189EE84E7A0EC1" box="[734,817,476,502]" captionStart="Fig" captionStartId="195.[189,232,1645,1671]" captionTargetBox="[205,1383,287,1593]" captionTargetId="figure@195.[189,1398,276,1606]" captionTargetPageId="195" captionText="Fig. 70. Claiborne Group, “upper” Lisbon Formation, otoliths. A–B. Diretmus? cf. D. serratus (Müller, 1999), MSC 42212. A. Inner face (left sagitta). B. Anterior view. — C–D. Preophidion meyeri (Koken, 1888), MSC 39055. C. Inner face (right sagitta). D. Dorsal view. — E–F. Preophidion cf. P. petropolis Frizzell &amp; Dante, 1965, MSC 42193. E. Inner face (right sagitta). F. Dorsal view. — G–H. Signata stenzeli Frizzell &amp; Dante, 1965, MSC 39051. G. Inner face (left sagitta). H. Dorsal view. — I–J. Signata nicoli Frizzell &amp; Dante, 1965, MSC 39046. I. Inner face (left sagitta). J. Dorsal view. — K–L. Hoplobrotula melrosensis (Frizzell &amp; Dante, 1965), MSC 37059.3. K. Inner face (right sagitta). L. Dorsal view. — M–N. Lepophidium? mucronata (Koken, 1888), MSC 39054. M. Inner face (left sagitta). N. Dorsal view. — O–P. Gobiidae Cuvier, 1816, indet. MSC 39045. O. Inner face (left sagitta). P. Dorsal view. — Q–R. Bothidae Smitt, 1892, indet. MSC 42235. Q. Inner face (right sagitta). R. Dorsal view. Scale bars: A–P= 2 mm; Q–R =1 mm." figureDoi="http://doi.org/10.5281/zenodo.3660395" httpUri="https://zenodo.org/record/3660395/files/figure.png" pageId="199" pageNumber="200">Fig. 70</figureCitation>
I–J
</paragraph>
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<subSubSection id="D8A4AFDCFF42FF68FF7B9D2B4EC30D4B" pageId="199" pageNumber="200" type="reference_group">
<paragraph id="9001FC57FF42FF68FF7B9D2B49F10D0E" blockId="199.[189,1210,542,569]" box="[189,1210,542,569]" pageId="199" pageNumber="200">
<treatmentCitationGroup id="B0AEDB79FF42FF68FF7B9D2B49F10D0E" box="[189,1210,542,569]" pageId="199" pageNumber="200">
<taxonomicName id="57BE87D4FF42FF68FF7B9D2B4EEB0D0E" authority="Dante &amp; Frizzell in Frizzell &amp; Dante, 1965: 709" authorityName="Dante &amp; Frizzell in Frizzell &amp; Dante" authorityPageNumber="709" authorityYear="1965" box="[189,928,542,569]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FF7B9D2B4C2A0D0E" box="[189,353,543,569]" italics="true" pageId="199" pageNumber="200">Signata nicoli</emphasis>
Dante &amp; Frizzell in
<treatmentCitation id="111FDA46FF42FF68FD959D2B4EEB0D0E" author="Frizzell D. L. &amp; Dante J. H." box="[595,928,542,569]" page="709" pageId="199" pageNumber="200" year="1965">
<bibRefCitation id="F42F81A6FF42FF68FD959D2B4EEB0D0E" author="Frizzell D. L. &amp; Dante J. H." box="[595,928,542,569]" pageId="199" pageNumber="200" pagination="368 - 718" refId="ref142032" refString="Frizzell D. L. &amp; Dante J. H. 1965. Otoliths of some early Cenozoic fishes of the Gulf Coast. Journal of Paleontology 39 (4): 368 - 718." type="journal article" year="1965">Frizzell &amp; Dante, 1965: 709</bibRefCitation>
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, pl. 88, figs 3–4, 21, 35.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B9D554EC30D4B" blockId="199.[189,904,609,672]" box="[189,904,609,636]" pageId="199" pageNumber="200">
“genus
<taxonomicName id="57BE87D4FF42FF68FED09D554CF70D4C" baseAuthorityName="Koken" baseAuthorityYear="1885" box="[278,444,609,635]" class="Actinopterygii" family="Ophidiidae" genus="Ophidiidarum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF42FF68FED09D554CF70D4C" box="[278,444,609,635]" italics="true" pageId="199" pageNumber="200">Ophidiidarum</emphasis>
</taxonomicName>
”
<taxonomicName id="57BE87D4FF42FF68FE169D554FAD0D4C" authority="- Nolf 1980: 138" authorityName="Nolf" authorityPageNumber="138" authorityYear="1980" box="[464,742,609,636]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FE169D554F580D4C" box="[464,531,609,635]" italics="true" pageId="199" pageNumber="200">nicoli</emphasis>
–
<bibRefCitation id="F42F81A6FF42FF68FDF69D554FAD0D4C" author="Nolf D." box="[560,742,609,635]" pageId="199" pageNumber="200" pagination="71 - 195" refId="ref147547" refString="Nolf D. 1980. Etude monographique des otolithes des Ophidiiformes actuels et revision des especes fossiles (Pisces, Teleostei). Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 17 (2): 71 - 195." type="journal article" year="1980">Nolf 1980: 138</bibRefCitation>
</taxonomicName>
, pl. 20, fig. 7.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF42FF68FF7B9DB14F890DA8" box="[189,706,645,672]" pageId="199" pageNumber="200" type="description">
<paragraph id="9001FC57FF42FF68FF7B9DB14F890DA8" blockId="199.[189,904,609,672]" box="[189,706,645,672]" pageId="199" pageNumber="200">
“
<emphasis id="A2CA2045FF42FF68FF0C9DB14C0A0DA8" box="[202,321,645,671]" italics="true" pageId="199" pageNumber="200">Ophidiida</emphasis>
”
<taxonomicName id="57BE87D4FF42FF68FE939DB14F160D97" authority="- Nolf 2013: 65" authorityName="- Nolf" authorityPageNumber="65" authorityYear="2013" box="[341,605,645,672]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FE939DB14CD30DA8" box="[341,408,645,671]" italics="true" pageId="199" pageNumber="200">nicoli</emphasis>
–
<bibRefCitation id="F42F81A6FF42FF68FE739DB14F160D97" author="Nolf D." box="[437,605,645,672]" pageId="199" pageNumber="200" refId="ref147755" refString="Nolf D. 2013. The Diversity of Fish Otoliths, Past and Present. Operational directorate &quot; Earth and History of Life &quot; of the Royal Belgian Institute of Natural Sciences, Brussels." type="book" year="2013">Nolf 2013: 65</bibRefCitation>
</taxonomicName>
, pl. 124.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF42FF68FF7B9DFC494A0C26" pageId="199" pageNumber="200" type="materials_examined">
<paragraph id="9001FC57FF42FF68FF7B9DFC4CE10DD5" blockId="199.[189,426,712,738]" box="[189,426,712,738]" pageId="199" pageNumber="200">
<heading id="CB494B3BFF42FF68FF7B9DFC4CE10DD5" bold="true" box="[189,426,712,738]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
<emphasis id="A2CA2045FF42FF68FF7B9DFC4CE10DD5" bold="true" box="[189,426,712,738]" pageId="199" pageNumber="200">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B9DCC494A0C26" blockId="199.[189,1025,759,786]" box="[189,1025,759,786]" pageId="199" pageNumber="200">
<materialsCitation id="20D6F60AFF42FF68FF7B9DCC4EB70C26" ID-GBIF-Occurrence="2564277978" box="[189,1020,759,786]" collectionCode="MSC" country="United States of America" location="Alabama" pageId="199" pageNumber="200" specimenCount="1" stateProvince="Alabama">
<collectingCountry id="E8A9BCC7FF42FF68FF7B9DCC4F1B0C25" box="[189,592,760,786]" name="United States of America" pageId="199" pageNumber="200">UNITED STATES OF AMERICA</collectingCountry>
–
<emphasis id="A2CA2045FF42FF68FDAB9DC34F940C26" bold="true" box="[621,735,759,785]" pageId="199" pageNumber="200">
<collectingRegion id="527A32B5FF42FF68FDAB9DC34F940C26" box="[621,735,759,785]" country="United States of America" name="Alabama" pageId="199" pageNumber="200">Alabama</collectingRegion>
</emphasis>
• 1 otolith;
<collectionCode id="F6AF6492FF42FF68FCAC9DCC4EE30C25" box="[874,936,760,786]" pageId="199" pageNumber="200">MSC</collectionCode>
39046
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF42FF68FF7B9C0E49BF0A24" pageId="199" pageNumber="200" type="description">
<paragraph id="9001FC57FF42FF68FF7B9C0E4C050C63" blockId="199.[189,334,826,852]" box="[189,334,826,852]" pageId="199" pageNumber="200">
<heading id="CB494B3BFF42FF68FF7B9C0E4C050C63" bold="true" box="[189,334,826,852]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
<emphasis id="A2CA2045FF42FF68FF7B9C0E4C050C63" bold="true" box="[189,334,826,852]" pageId="199" pageNumber="200">Description</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B9C5D49BF0A24" blockId="199.[189,1399,873,1299]" pageId="199" pageNumber="200">Sagittae primarily of ovate shape; height/length ratio around 70%. Smooth margins. Anterior margin broadly rounded. Anterodorsal slope shorter and noticeably steeper than posterodorsal slope. Dorsal margin quite short, almost straight. Posterodorsal slope incurved, longer than anterodorsal slope.Posterior sharply rounded. Ventral margin broadly rounded, but posteroventral slope is slightly steeper. Smooth inner face moderately convex or inflated, slopes toward the margins. Extremely wide, undivided, long (approximately 98% of the otolith length) sulcus. Very fine incised line marks sulcus. Sulcus extending essentially from anterior margin to very near posterior margin. Very small ostial channel opens on anterior margin (visible on well-preserved specimens). Dorsal margin of sulcus closer to dorsal margin than ventral margin of sulcus is to ventral margin. Dorsal margin of sulcus noticeably flexed upward anterior to the vertical midline of the otolith. Ventral margin of sulcus slightly curved or sinuous. Cristae not developed; no depressed area observed. Ventral furrow absent. Outer face slightly and irregularly concave, commonly sculptured with smooth bosses and rugosities, especially in dorsal area.</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF42FF68FF7B9A0F4C8609BD" pageId="199" pageNumber="200" type="discussion">
<paragraph id="9001FC57FF42FF68FF7B9A0F4C640A62" blockId="199.[189,303,1339,1365]" box="[189,303,1339,1365]" pageId="199" pageNumber="200">
<heading id="CB494B3BFF42FF68FF7B9A0F4C640A62" bold="true" box="[189,303,1339,1365]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
<emphasis id="A2CA2045FF42FF68FF7B9A0F4C640A62" bold="true" box="[189,303,1339,1365]" pageId="199" pageNumber="200">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B9A5E4E3A0AC5" blockId="199.[189,1399,1386,1522]" pageId="199" pageNumber="200">
The sagitta of
<taxonomicName id="57BE87D4FF42FF68FEAC9A5F4F5B0AB3" authorityName="- Nolf" authorityYear="2013" box="[362,528,1386,1413]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FEAC9A5F4F5B0AB3" box="[362,528,1386,1413]" italics="true" pageId="199" pageNumber="200">Signata nicoli</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="57BE87D4FF42FF68FD2A9A5F4E280AB3" authorityName="- Nolf" authorityYear="2013" box="[748,867,1386,1413]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="stenzeli">
<emphasis id="A2CA2045FF42FF68FD2A9A5F4E280AB3" box="[748,867,1386,1413]" italics="true" pageId="199" pageNumber="200">S. stenzeli</emphasis>
</taxonomicName>
, but there are some distinct differences. The flexure on the dorsal margin of the sulcus of
<taxonomicName id="57BE87D4FF42FF68FD089ABB4E650A9E" authorityName="- Nolf" authorityYear="2013" box="[718,814,1423,1449]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FD089ABB4E650A9E" box="[718,814,1423,1449]" italics="true" pageId="199" pageNumber="200">S. nicoli</emphasis>
</taxonomicName>
is not nearly as pronounced as on
<taxonomicName id="57BE87D4FF42FF68FB019ABB48760A9E" authorityName="- Nolf" authorityYear="2013" box="[1223,1341,1423,1449]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="stenzeli">
<emphasis id="A2CA2045FF42FF68FB019ABB48760A9E" box="[1223,1341,1423,1449]" italics="true" pageId="199" pageNumber="200">S. stenzeli</emphasis>
</taxonomicName>
, and the ventral margin of
<taxonomicName id="57BE87D4FF42FF68FE799A874F6A0AFA" authorityName="- Nolf" authorityYear="2013" box="[447,545,1459,1485]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FE799A874F6A0AFA" box="[447,545,1459,1485]" italics="true" pageId="199" pageNumber="200">S. nicoli</emphasis>
</taxonomicName>
has little or no flexure upward. Additionally, the posterodorsal slope on
<taxonomicName id="57BE87D4FF42FF68FF7B9AEC4C560AC5" authorityName="- Nolf" authorityYear="2013" box="[189,285,1496,1522]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="nicoli">
<emphasis id="A2CA2045FF42FF68FF7B9AEC4C560AC5" box="[189,285,1496,1522]" italics="true" pageId="199" pageNumber="200">S. nicoli</emphasis>
</taxonomicName>
is incurved rather than outcurved as on
<emphasis id="A2CA2045FF42FF68FD329AEC4E3A0AC5" box="[756,881,1496,1522]" italics="true" pageId="199" pageNumber="200">
<taxonomicName id="57BE87D4FF42FF68FD329AEC4E260AC5" authorityName="- Nolf" authorityYear="2013" box="[756,877,1496,1522]" class="Actinopterygii" family="Ophidiidae" genus="Signata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="199" pageNumber="200" phylum="Chordata" rank="species" species="stenzeli">S. stenzeli</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B992F4E5F0902" blockId="199.[189,788,1563,1589]" box="[189,788,1563,1589]" pageId="199" pageNumber="200">
<heading id="CB494B3BFF42FF68FF7B992F4E5F0902" bold="true" box="[189,788,1563,1589]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
<emphasis id="A2CA2045FF42FF68FF7B992F4E5F0902" bold="true" box="[189,788,1563,1589]" pageId="199" pageNumber="200">
Stratigraphic and geographic range in
<collectingRegion id="527A32B5FF42FF68FD64992F4E5F0902" box="[674,788,1563,1589]" country="United States of America" name="Alabama" pageId="199" pageNumber="200">Alabama</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF42FF68FF7B997F4C8609BD" blockId="199.[189,1398,1611,1674]" pageId="199" pageNumber="200">
The singular specimen was recovered from the “upper”
<collectingRegion id="527A32B5FF42FF68FCA1997F4EF30952" box="[871,952,1611,1637]" country="Portugal" name="Lisboa" pageId="199" pageNumber="200">Lisbon</collectingRegion>
Formation at site ACh-8. Bartonian, zones NP16 and NP17.
</paragraph>
</subSubSection>
</treatment>
</document>
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