536 lines
73 KiB
XML
536 lines
73 KiB
XML
<document id="6E4B3F1C0BD48E38BE66186BE7DF358C" ID-CLB-Dataset="188710" ID-DOI="10.11646/zootaxa.5209.2.2" ID-GBIF-Dataset="87405f1b-232d-4e0c-a7d8-4d6824a801c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7325861" ID-ZooBank="40F0B633-691B-404E-95B3-0C23F5133791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668583694219" checkinUser="plazi" docAuthor="Zumbado Echavarria, Marco A., Barrantes Barrantes, Edwin A., Helmick, Ericka E., Bartlett, Charles R. & Bahder, Brian W." docDate="2022" docId="154B032E356B4278F2F47B8EFA300AC9" docLanguage="en" docName="zootaxa.5209.2.2.pdf" docOrigin="Zootaxa 5209 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tico villablanca Bahder & Bartlett 2022, sp. n." docType="treatment" docVersion="8" lastPageNumber="195" masterDocId="E9727B5635684270F2637D0DFFAC0C24" masterDocTitle="A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms in Costa Rica" masterLastPageNumber="198" masterPageNumber="187" pageNumber="190" updateTime="1699411561995" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BA703FEE37FA78C542CE1F771C3E974C">
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<mods:title id="E498A594F8F544907B9A865938CB1FA7">A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms in Costa Rica</mods:title>
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<mods:name id="8F0B48FC7F649B1494F74E861750D601" type="personal">
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<mods:namePart id="7D186BB5D7345BDC7DBEA9DEB591B5D5">Zumbado Echavarria, Marco A.</mods:namePart>
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<mods:affiliation id="45CAEA183AC6C23E444A5B55338084EC">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="69C1613F6B6511C8132142BBD58E4135">Barrantes Barrantes, Edwin A.</mods:namePart>
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<mods:affiliation id="BAC617D78423D83F39BADC7B24EA30AC">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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</mods:name>
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<mods:namePart id="C5969865D730EDF0FE48EC0EA430A313">Helmick, Ericka E.</mods:namePart>
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<mods:affiliation id="55F7B208F0EE39E58C6A62DDE38116B6">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. E-mail: ehelmick @ ufl. edu</mods:affiliation>
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</mods:name>
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<mods:namePart id="A50DCEB7648B87071EC1DAD91EFE3DA5">Bartlett, Charles R.</mods:namePart>
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<mods:affiliation id="D77293C23188309992F24483E978B0AA">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
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<mods:namePart id="E66C89EF9671C6540A9E80D30E9964ED">Bahder, Brian W.</mods:namePart>
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<mods:nameIdentifier id="4E938947E849B3A509305B37F917CDA7" type="ORCID">0000-0002-1118-4832</mods:nameIdentifier>
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<mods:affiliation id="F82B0D9E84698B9E55DF0C97724C1C33">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. bbahder @ ufl. edu; https: // orcid. org / 0000 - 0002 - 1118 - 4832</mods:affiliation>
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<mods:nameIdentifier id="76797635E959C1BB18328EC988C2B543" type="email">bbahder@ufl.edu</mods:nameIdentifier>
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<mods:title id="36CD6FB4C6DB62BF4DE3FD6B18A0F383">Zootaxa</mods:title>
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<mods:part id="827F20EC4FE6D82BB62F7965A9FB0618">
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<mods:date id="72CD9B0A7E2A075F1B76B7379D2B68FA">2022</mods:date>
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<mods:number id="1AF20C378F343CD49BA8F8E88F1819DE">2022-11-16</mods:number>
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<mods:number id="B458C3B485A4E7A2B695E4C75FF3C694">5209</mods:number>
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<treatment id="154B032E356B4278F2F47B8EFA300AC9" ID-DOI="http://doi.org/10.5281/zenodo.7330825" ID-GBIF-Taxon="205079189" ID-Zenodo-Dep="7330825" LSID="urn:lsid:plazi:treatment:154B032E356B4278F2F47B8EFA300AC9" httpUri="http://treatment.plazi.org/id/154B032E356B4278F2F47B8EFA300AC9" lastPageId="8" lastPageNumber="195" pageId="3" pageNumber="190">
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<subSubSection id="D5F8E1B3356B4273F2F47B8EFD230AB9" box="[151,655,1666,1693]" pageId="3" pageNumber="190" type="nomenclature">
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<paragraph id="9D5DB238356B4273F2F47B8EFD230AB9" blockId="3.[151,655,1666,1729]" box="[151,655,1666,1693]" pageId="3" pageNumber="190">
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<heading id="C6150554356B4273F2F47B8EFD230AB9" bold="true" box="[151,655,1666,1693]" fontSize="11" level="1" pageId="3" pageNumber="190" reason="1">
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<emphasis id="AF966E2A356B4273F2F47B8EFD230AB9" bold="true" box="[151,655,1666,1693]" pageId="3" pageNumber="190">
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<taxonomicName id="5AE2C9BB356B4273F2F47B8EFDEA0AB9" authority="Bahder & Bartlett" authorityName="Bahder & Bartlett" authorityYear="2022" box="[151,582,1666,1693]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
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<emphasis id="AF966E2A356B4273F2F47B8EFEF50AB9" bold="true" box="[151,345,1666,1693]" italics="true" pageId="3" pageNumber="190">Tico villablanca</emphasis>
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Bahder & Bartlett
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</taxonomicName>
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<taxonomicNameLabel id="B4A5D351356B4273F02E7B8EFD230AB9" box="[589,655,1667,1693]" pageId="3" pageNumber="190" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D5F8E1B3356B4273F2F47BAAFE830AE5" box="[151,303,1703,1729]" pageId="3" pageNumber="190" type="description">
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<paragraph id="9D5DB238356B4273F2F47BAAFE830AE5" blockId="3.[151,655,1666,1729]" box="[151,303,1703,1729]" pageId="3" pageNumber="190">
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(
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<figureCitation id="05D9AEBD356B4273F2FC7BAAFE8B0AE5" box="[159,295,1703,1729]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[152,255,1373,1397]" captionStartId-1="5.[152,255,926,950]" captionStartId-2="5.[152,255,1943,1967]" captionStartId-3="7.[152,255,1310,1334]" captionStartId-4="7.[152,255,1845,1869]" captionTargetBox-0="[151,1436,273,1350]" captionTargetBox-1="[156,1430,390,895]" captionTargetBox-2="[160,1428,1012,1914]" captionTargetBox-3="[151,1428,182,1279]" captionTargetBox-4="[151,1436,1358,1821]" captionTargetId-0="figure-65@4.[151,1436,273,1350]" captionTargetId-1="figure-122@5.[152,1436,381,903]" captionTargetId-2="figure-162@5.[151,1436,1011,1920]" captionTargetId-3="figure-18@7.[151,1436,181,1285]" captionTargetId-4="figure-43@7.[151,1436,1358,1821]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="7" captionText-0="FIGURE 2. Adult male habitus of Tico villablanca sp. n.; A) lateral view, and B) dorsal view; scale = 1 mm." captionText-1="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." captionText-2="FIGURE 4. Adult male Tico villablanca sp. n. forewing venation." captionText-3="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." captionText-4="FIGURE 6. Male Tico villablanca sp. n. gonostylus; A) lateral view and B) inner-ventral view displaying sclerotized ridge." figureDoi-0="http://doi.org/10.5281/zenodo.7325867" figureDoi-1="http://doi.org/10.5281/zenodo.7325869" figureDoi-2="http://doi.org/10.5281/zenodo.7325871" figureDoi-3="http://doi.org/10.5281/zenodo.7325873" figureDoi-4="http://doi.org/10.5281/zenodo.7325877" httpUri-0="https://zenodo.org/record/7325867/files/figure.png" httpUri-1="https://zenodo.org/record/7325869/files/figure.png" httpUri-2="https://zenodo.org/record/7325871/files/figure.png" httpUri-3="https://zenodo.org/record/7325873/files/figure.png" httpUri-4="https://zenodo.org/record/7325877/files/figure.png" pageId="3" pageNumber="190">Figures 2–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D5F8E1B3356B4273F2F47BE2FA870B52" pageId="3" pageNumber="190" type="diagnosis">
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<paragraph id="9D5DB238356B4273F2F47BE2FA870B52" blockId="3.[151,1437,1775,2018]" pageId="3" pageNumber="190">
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<emphasis id="AF966E2A356B4273F2F47BE2FEBE0B2D" bold="true" box="[151,274,1775,1801]" pageId="3" pageNumber="190">Diagnosis.</emphasis>
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Body pale (generally yellowish) strongly contrasting with red-fuscous wings. Head pale with fuscous patches on posterior half of genae below eyes. Lateral carinae of vertex and frons foliate with sensorial pits obsolete. Dorsal apex of aedeagal shaft bearing large, heart-like flanges with serrations on anterior margin. Gonostyli with elongate sclerotized ridge on inner margin bearing low, irregular serrations and elongate process at apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="D5F8E1B3356B4276F2A47A72FCDA0AB0" lastPageId="6" lastPageNumber="193" pageId="3" pageNumber="190" type="description">
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<paragraph id="9D5DB238356B4274F2A47A72FEFB0CDD" blockId="3.[151,1437,1775,2018]" lastBlockId="4.[151,1436,151,249]" lastPageId="4" lastPageNumber="191" pageId="3" pageNumber="190">
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<emphasis id="AF966E2A356B4273F2A47A72FEF50BBD" bold="true" box="[199,345,1919,1945]" pageId="3" pageNumber="190">Description.</emphasis>
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<emphasis id="AF966E2A356B4273F3007A72FE080BBD" box="[355,420,1919,1945]" italics="true" pageId="3" pageNumber="190">Color</emphasis>
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. General body color yellow-orange (
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<figureCitation id="05D9AEBD356B4273F1317A72FC360BBD" box="[850,922,1919,1945]" captionStart="FIGURE 2" captionStartId="4.[152,255,1373,1397]" captionTargetBox="[151,1436,273,1350]" captionTargetId="figure-65@4.[151,1436,273,1350]" captionTargetPageId="4" captionText="FIGURE 2. Adult male habitus of Tico villablanca sp. n.; A) lateral view, and B) dorsal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325867" httpUri="https://zenodo.org/record/7325867/files/figure.png" pageId="3" pageNumber="190">Fig. 2</figureCitation>
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), head and venter paler, posterior portion of genae marked with fuscous patch from eyes to frontoclypeal suture (washed with red near lateral ocelli); clypeus washed with red (
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<figureCitation id="05D9AEBD356B4273F3017ACAFE100BC5" box="[354,444,1991,2017]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="3" pageNumber="190">Fig. 3C</figureCitation>
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). Foliate region of paranota of prothorax paler then dorsal region, legs pale. Forewings red-fuscous, veins red distally, orange basally (
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<figureCitation id="05D9AEBD356C4274F0C77D9AFD450C95" box="[676,745,151,177]" captionStart="FIGURE 2" captionStartId="4.[152,255,1373,1397]" captionTargetBox="[151,1436,273,1350]" captionTargetId="figure-65@4.[151,1436,273,1350]" captionTargetPageId="4" captionText="FIGURE 2. Adult male habitus of Tico villablanca sp. n.; A) lateral view, and B) dorsal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325867" httpUri="https://zenodo.org/record/7325867/files/figure.png" pageId="4" pageNumber="191">Fig. 2</figureCitation>
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). Tergites of abdominal segment washed with bright red, sternites yellow-orange. Gonostyli pale, translucent, and slightly tinted with red and orange on dorsal margin, anal tube light red (
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<figureCitation id="05D9AEBD356C4274F3667DD2FEE50CDD" box="[261,329,222,249]" captionStart="FIGURE 5" captionStartId="7.[152,255,1310,1334]" captionTargetBox="[151,1428,182,1279]" captionTargetId="figure-18@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325873" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="4" pageNumber="191">Fig. 5</figureCitation>
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).
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</paragraph>
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<caption id="C99DE2B0356C4274F2FB7850FB5C0951" ID-DOI="http://doi.org/10.5281/zenodo.7325867" ID-Zenodo-Dep="7325867" box="[152,1264,1373,1398]" httpUri="https://zenodo.org/record/7325867/files/figure.png" pageId="4" pageNumber="191" startId="4.[152,255,1373,1397]" targetBox="[151,1436,273,1350]" targetPageId="4">
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<paragraph id="9D5DB238356C4274F2FB7850FB5C0951" blockId="4.[152,1264,1373,1398]" box="[152,1264,1373,1398]" pageId="4" pageNumber="191">
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<emphasis id="AF966E2A356C4274F2FB7850FEB40951" bold="true" box="[152,280,1373,1397]" pageId="4" pageNumber="191">FIGURE 2.</emphasis>
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Adult male habitus of
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<taxonomicName id="5AE2C9BB356C4274F0627853FD090952" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[513,677,1374,1398]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
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<emphasis id="AF966E2A356C4274F0627853FD090952" box="[513,677,1374,1398]" italics="true" pageId="4" pageNumber="191">Tico villablanca</emphasis>
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</taxonomicName>
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<emphasis id="AF966E2A356C4274F0CF7853FD490952" bold="true" box="[684,741,1374,1398]" pageId="4" pageNumber="191">
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<taxonomicNameLabel id="B4A5D351356C4274F0CF7853FD490952" box="[684,741,1374,1398]" pageId="4" pageNumber="191" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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; A) lateral view, and B) dorsal view; scale = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="9D5DB238356C4274F2A478A9FD130A9E" blockId="4.[151,1437,1444,2010]" pageId="4" pageNumber="191">
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<emphasis id="AF966E2A356C4274F2A478A9FE9A099A" box="[199,310,1444,1470]" italics="true" pageId="4" pageNumber="191">Structure.</emphasis>
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Body length without wings
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<quantity id="5A1A1FDD356C4274F01478A9FCB6099A" box="[631,794,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" metricValueMax="1.78" metricValueMin="1.74" pageId="4" pageNumber="191" unit="mm" value="1.76" valueMax="1.78" valueMin="1.74">1.74–1.78 mm</quantity>
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(males),
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<quantity id="5A1A1FDD356C4274F1E778A9FB84099A" box="[900,1064,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" metricValueMax="1.83" metricValueMin="1.81" pageId="4" pageNumber="191" unit="mm" value="1.82" valueMax="1.83" valueMin="1.81">1.81–1.83 mm</quantity>
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(females); with wings
|
||
<quantity id="5A1A1FDD356C4274F74F78A9FF6F09C6" metricMagnitude="-3" metricUnit="m" metricValue="2.695" metricValueMax="2.71" metricValueMin="2.68" pageId="4" pageNumber="191" unit="mm" value="2.695" valueMax="2.71" valueMin="2.68">2.68–2.71 mm</quantity>
|
||
(males),
|
||
<quantity id="5A1A1FDD356C4274F34D78C5FE3809C6" box="[302,404,1480,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="4" pageNumber="191" unit="mm" value="2.75">2.75 mm</quantity>
|
||
(females;
|
||
<tableCitation id="D0608783356C4274F06F78C5FDCE09C6" box="[524,610,1480,1506]" captionStart="TABLE 3" captionStartId="6.[152,244,151,175]" captionTargetPageId="6" captionText="TABLE 3. Biometric data for Tico villablanca sp. n." httpUri="http://table.plazi.org/id/C99DE2B0356E4276F2FB7D9AFD630C94" pageId="4" pageNumber="191" tableUuid="C99DE2B0356E4276F2FB7D9AFD630C94">Table 3</tableCitation>
|
||
). Head. In lateral view, rounded with slight angle representing the transverse carina at the fastigium (
|
||
<figureCitation id="05D9AEBD356C4274F38078E1FD970A22" box="[483,571,1516,1542]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="4" pageNumber="191">Fig. 3B</figureCitation>
|
||
). In dorsal view, vertex trapezoidal, truncate at anterior margin (at transverse carina), concave between eyes at posterior margin, lateral carinae foliate smooth (lacking sensorial pits) (
|
||
<figureCitation id="05D9AEBD356C4274F7567B1DFA230A0E" box="[1333,1423,1552,1578]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="4" pageNumber="191">Fig. 3A</figureCitation>
|
||
), disc concave, median carina absent. In frontal view, frons concave with lateral margins foliate and irregularly sinuate (sensorial pits lacking), narrowest between eyes, slightly wider at ventral margin then dorsal margin. Lateral ocelli conspicuous below eyes, antennae short, scale ringlike, pedicle spheroid bearing inconspicuous sensory plaques, flagellum bristle-like with bulbous base.
|
||
</paragraph>
|
||
<paragraph id="9D5DB238356C4275F2A47BC9FE200D41" blockId="4.[151,1437,1444,2010]" lastBlockId="5.[151,1436,150,357]" lastPageId="5" lastPageNumber="192" pageId="4" pageNumber="191">
|
||
Thorax. Pronotum relatively elongate, at midline approximately equal length to vertex (
|
||
<figureCitation id="05D9AEBD356C4274F6FF7BC9FB5A0AFA" box="[1180,1270,1732,1758]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="4" pageNumber="191">Fig. 3A</figureCitation>
|
||
), anterior margin truncate, posterior margin moderately concave; distinctly tricarinate (lateral carinae laterally arched, reaching hind margin), paradiscal region distinctly declinate at lateral carinae (from dorsal view); in lateral view (
|
||
<figureCitation id="05D9AEBD356C4274F7557A01FA220B02" box="[1334,1422,1804,1830]" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="4" pageNumber="191">Fig. 3B</figureCitation>
|
||
), posterior margin raised, declining anteriorly; paradiscal regions broadly foliate, forming cup-like fossa surrounding antennae posteriorly; in frontal view, foliations broadly exceeding antennae, appearing rounded laterally (Fig, 3C). Mesonotum approximately as long at midline as wide (longer at midline than vertex and pronotum combined,
|
||
<figureCitation id="05D9AEBD356C4274F7137A75FF110B92" captionStart="FIGURE 3" captionStartId="5.[152,255,926,950]" captionTargetBox="[156,1430,390,895]" captionTargetId="figure-122@5.[152,1436,381,903]" captionTargetPageId="5" captionText="FIGURE 3. Adult Tico villablanca sp. n.; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="4" pageNumber="191">Fig. 3A</figureCitation>
|
||
); carinate at midline, lateral carinae obscure. Forewing (
|
||
<figureCitation id="05D9AEBD356C4274F1237A91FC290B92" box="[832,901,1948,1974]" captionStart="FIGURE 4" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[160,1428,1012,1914]" captionTargetId="figure-162@5.[151,1436,1011,1920]" captionTargetPageId="5" captionText="FIGURE 4. Adult male Tico villablanca sp. n. forewing venation." figureDoi="http://doi.org/10.5281/zenodo.7325871" httpUri="https://zenodo.org/record/7325871/files/figure.png" pageId="4" pageNumber="191">Fig. 4</figureCitation>
|
||
) with apex of clavus near forewing midlength, fused vein Pcu+A1 reaching wing margin distantly proximad to CuP at claval apex. Fork of R (forming C1 cell) just preceding claval apex, fork of CuA proximad of R fork (cell C5 longer than C1); composite vein M+R+ScP forming elongate stem from basal cell before branching of MP from ScP+R. Branching pattern: RA 1-branched, RP 2-branched (RP
|
||
<subScript id="0166B07D356D4275F32A7DFCFECE0CDB" attach="left" box="[329,354,241,255]" fontSize="6" pageId="5" pageNumber="192">1+2</subScript>
|
||
and RP
|
||
<subScript id="0166B07D356D4275F3DE7DFCFE7B0CDB" attach="left" box="[445,471,241,255]" fontSize="6" pageId="5" pageNumber="192">3+4</subScript>
|
||
), MP 4-branched (a forked MP
|
||
<subScript id="0166B07D356D4275F15F7DFCFCE80CDB" attach="left" box="[828,836,241,255]" fontSize="6" pageId="5" pageNumber="192">1</subScript>
|
||
plus MP
|
||
<subScript id="0166B07D356D4275F1C87DFCFC1F0CDB" attach="left" box="[939,947,241,255]" fontSize="6" pageId="5" pageNumber="192">2</subScript>
|
||
and MP
|
||
<subScript id="0166B07D356D4275F6777DFCFB820CDB" attach="left" box="[1044,1070,241,255]" fontSize="6" pageId="5" pageNumber="192">3+4</subScript>
|
||
; CuA 2 branched (forming cells C5 and C5’ by icua crossvein and icu forming marginal C6 cell). Large pustules along subcostal vein, Pcu and costal vein between ScP and Ra, very small pustules present along all veins, branches and crossveins from basal region to distal margin (
|
||
<figureCitation id="05D9AEBD356D4275F35A7C46FED20D41" box="[313,382,331,357]" captionStart="FIGURE 4" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[160,1428,1012,1914]" captionTargetId="figure-162@5.[151,1436,1011,1920]" captionTargetPageId="5" captionText="FIGURE 4. Adult male Tico villablanca sp. n. forewing venation." figureDoi="http://doi.org/10.5281/zenodo.7325871" httpUri="https://zenodo.org/record/7325871/files/figure.png" pageId="5" pageNumber="192">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C99DE2B0356D4275F2FB7E93FCD00FFE" ID-DOI="http://doi.org/10.5281/zenodo.7325869" ID-Zenodo-Dep="7325869" httpUri="https://zenodo.org/record/7325869/files/figure.png" pageId="5" pageNumber="192" startId="5.[152,255,926,950]" targetBox="[156,1430,390,895]" targetPageId="5">
|
||
<paragraph id="9D5DB238356D4275F2FB7E93FCD00FFE" blockId="5.[151,1436,926,987]" pageId="5" pageNumber="192">
|
||
<emphasis id="AF966E2A356D4275F2FB7E93FEB40F93" bold="true" box="[152,280,926,951]" pageId="5" pageNumber="192">FIGURE 3.</emphasis>
|
||
Adult
|
||
<taxonomicName id="5AE2C9BB356D4275F33E7E92FDAC0F93" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[349,512,927,951]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="192" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356D4275F33E7E92FDAC0F93" box="[349,512,927,951]" italics="true" pageId="5" pageNumber="192">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356D4275F0657E92FD920F93" bold="true" box="[518,574,927,951]" pageId="5" pageNumber="192">
|
||
<taxonomicNameLabel id="B4A5D351356D4275F0657E92FD920F93" box="[518,574,927,951]" pageId="5" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; A) head, pronotum and mesonotum dorsal view, B) head, pronotum, and mesonotum right lateral view, and C) head and pronotum frontal view; scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C99DE2B0356D4275F2FB7A9AFCEC0B8B" ID-DOI="http://doi.org/10.5281/zenodo.7325871" ID-Zenodo-Dep="7325871" box="[152,832,1943,1967]" httpUri="https://zenodo.org/record/7325871/files/figure.png" pageId="5" pageNumber="192" startId="5.[152,255,1943,1967]" targetBox="[160,1428,1012,1914]" targetPageId="5">
|
||
<paragraph id="9D5DB238356D4275F2FB7A9AFCEC0B8B" blockId="5.[152,832,1943,1967]" box="[152,832,1943,1967]" pageId="5" pageNumber="192">
|
||
<emphasis id="AF966E2A356D4275F2FB7A9AFEB40B8B" bold="true" box="[152,280,1943,1967]" pageId="5" pageNumber="192">FIGURE 4.</emphasis>
|
||
Adult male
|
||
<taxonomicName id="5AE2C9BB356D4275F3F57A95FD960B8B" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[406,570,1943,1967]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="192" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356D4275F3F57A95FD960B8B" box="[406,570,1943,1967]" italics="true" pageId="5" pageNumber="192">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356D4275F0227A9AFDD60B8B" bold="true" box="[577,634,1943,1967]" pageId="5" pageNumber="192">
|
||
<taxonomicNameLabel id="B4A5D351356D4275F0227A9AFDD60B8B" box="[577,634,1943,1967]" pageId="5" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
forewing venation.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C99DE2B0356E4276F2FB7D9AFD630C94" ID-Table-UUID="C99DE2B0356E4276F2FB7D9AFD630C94" box="[152,719,151,176]" httpUri="http://table.plazi.org/id/C99DE2B0356E4276F2FB7D9AFD630C94" pageId="6" pageNumber="193" startId="6.[152,244,151,175]" targetBox="[166,1350,192,860]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="9D5DB238356E4276F2FB7D9AFD630C94" blockId="6.[152,719,151,176]" box="[152,719,151,176]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2FB7D9AFEA30C8B" bold="true" box="[152,271,151,175]" pageId="6" pageNumber="193">TABLE 3.</emphasis>
|
||
Biometric data for
|
||
<taxonomicName id="5AE2C9BB356E4276F3837D95FD200C8B" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[480,652,151,175]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356E4276F3837D95FD200C8B" box="[480,652,151,175]" italics="true" pageId="6" pageNumber="193">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4A5D351356E4276F0F07D95FD630C94" box="[659,719,152,176]" pageId="6" pageNumber="193" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9D5DB238356E4276F0377DCDFA8C0F78" pageId="6" pageNumber="193">
|
||
<table id="EFE24098356EBD8FF2C57DCDFAEA0F78" box="[166,1350,192,860]" gridcols="5" gridrows="17" pageId="6" pageNumber="193">
|
||
<tr id="23D2B07A356EBD8FF2C57DCDFAEA0CFC" box="[166,1350,192,216]" gridrow="0" pageId="6" pageNumber="193" rowspan-0="1" rowspan-2="1" rowspan-4="1">
|
||
<th id="6003D906356EBD8FF0377DCDFD7F0CFC" box="[596,723,192,216]" gridcol="1" gridrow="0" pageId="6" pageNumber="193">
|
||
Male (
|
||
<emphasis id="AF966E2A356E4276F0F47DCCFD080CFC" box="[663,676,193,216]" italics="true" pageId="6" pageNumber="193">n</emphasis>
|
||
=11)
|
||
</th>
|
||
<th id="6003D906356EBD8FF19C7DCDFB240CFC" box="[1023,1160,192,216]" gridcol="3" gridrow="0" pageId="6" pageNumber="193">
|
||
Female (
|
||
<emphasis id="AF966E2A356E4276F63A7DCCFBCA0CFC" box="[1113,1126,193,216]" italics="true" pageId="6" pageNumber="193">n</emphasis>
|
||
=4)
|
||
</th>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57DE5FAEA0D25" box="[166,1350,232,257]" gridrow="1" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57DE5FE120D25" box="[166,446,232,257]" gridcol="0" gridrow="1" pageId="6" pageNumber="193">Character</th>
|
||
<td id="6003D906356EBD8FF0377DE5FD7F0D25" box="[596,723,232,257]" gridcol="1" gridrow="1" pageId="6" pageNumber="193">Range</td>
|
||
<td id="6003D906356EBD8FF16D7DE5FC360D25" box="[782,922,232,257]" gridcol="2" gridrow="1" pageId="6" pageNumber="193">Average ± SE</td>
|
||
<td id="6003D906356EBD8FF19C7DE5FB240D25" box="[1023,1160,232,257]" gridcol="3" gridrow="1" pageId="6" pageNumber="193">Range</td>
|
||
<td id="6003D906356EBD8FF6DA7DE5FAEA0D25" box="[1209,1350,232,257]" gridcol="4" gridrow="1" pageId="6" pageNumber="193">Average ± SE</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57C1DFAEA0D0C" box="[166,1350,272,296]" gridrow="2" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57C1DFE120D0C" box="[166,446,272,296]" gridcol="0" gridrow="2" pageId="6" pageNumber="193">Body length, with wings</th>
|
||
<td id="6003D906356EBD8FF0377C1DFD7F0D0C" box="[596,723,272,296]" gridcol="1" gridrow="2" pageId="6" pageNumber="193">2.68–2.71</td>
|
||
<td id="6003D906356EBD8FF16D7C1DFC360D0C" box="[782,922,272,296]" gridcol="2" gridrow="2" pageId="6" pageNumber="193">2.69±0.01</td>
|
||
<td id="6003D906356EBD8FF19C7C1DFB240D0C" box="[1023,1160,272,296]" gridcol="3" gridrow="2" pageId="6" pageNumber="193">2.75–2.75</td>
|
||
<td id="6003D906356EBD8FF6DA7C1DFAEA0D0C" box="[1209,1350,272,296]" gridcol="4" gridrow="2" pageId="6" pageNumber="193">2.75±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57C35FAEA0D75" box="[166,1350,312,337]" gridrow="3" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57C35FE120D75" box="[166,446,312,337]" gridcol="0" gridrow="3" pageId="6" pageNumber="193">Body length, no wings</th>
|
||
<td id="6003D906356EBD8FF0377C35FD7F0D75" box="[596,723,312,337]" gridcol="1" gridrow="3" pageId="6" pageNumber="193">1.69–1.71</td>
|
||
<td id="6003D906356EBD8FF16D7C35FC360D75" box="[782,922,312,337]" gridcol="2" gridrow="3" pageId="6" pageNumber="193">1.7±0.01</td>
|
||
<td id="6003D906356EBD8FF19C7C35FB240D75" box="[1023,1160,312,337]" gridcol="3" gridrow="3" pageId="6" pageNumber="193">1.81–1.81</td>
|
||
<td id="6003D906356EBD8FF6DA7C35FAEA0D75" box="[1209,1350,312,337]" gridcol="4" gridrow="3" pageId="6" pageNumber="193">1.81±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57C6CFAEA0D5D" box="[166,1350,353,377]" gridrow="4" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57C6CFE120D5D" box="[166,446,353,377]" gridcol="0" gridrow="4" pageId="6" pageNumber="193">Forewing length</th>
|
||
<td id="6003D906356EBD8FF0377C6CFD7F0D5D" box="[596,723,353,377]" gridcol="1" gridrow="4" pageId="6" pageNumber="193">2.19–2.21</td>
|
||
<td id="6003D906356EBD8FF16D7C6CFC360D5D" box="[782,922,353,377]" gridcol="2" gridrow="4" pageId="6" pageNumber="193">2.21±0.01</td>
|
||
<td id="6003D906356EBD8FF19C7C6CFB240D5D" box="[1023,1160,353,377]" gridcol="3" gridrow="4" pageId="6" pageNumber="193">2.24–2.24</td>
|
||
<td id="6003D906356EBD8FF6DA7C6CFAEA0D5D" box="[1209,1350,353,377]" gridcol="4" gridrow="4" pageId="6" pageNumber="193">2.24±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57C84FAEA0D85" box="[166,1350,393,417]" gridrow="5" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57C84FE120D85" box="[166,446,393,417]" gridcol="0" gridrow="5" pageId="6" pageNumber="193">Vertex length</th>
|
||
<td id="6003D906356EBD8FF0377C84FD7F0D85" box="[596,723,393,417]" gridcol="1" gridrow="5" pageId="6" pageNumber="193">0.14–0.14</td>
|
||
<td id="6003D906356EBD8FF16D7C84FC360D85" box="[782,922,393,417]" gridcol="2" gridrow="5" pageId="6" pageNumber="193">0.14±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7C84FB240D85" box="[1023,1160,393,417]" gridcol="3" gridrow="5" pageId="6" pageNumber="193">0.15–0.15</td>
|
||
<td id="6003D906356EBD8FF6DA7C84FAEA0D85" box="[1209,1350,393,417]" gridcol="4" gridrow="5" pageId="6" pageNumber="193">0.15±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57CBCFAEA0DED" box="[166,1350,433,457]" gridrow="6" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57CBCFE120DED" box="[166,446,433,457]" gridcol="0" gridrow="6" pageId="6" pageNumber="193">Vertex width, basal margin</th>
|
||
<td id="6003D906356EBD8FF0377CBCFD7F0DED" box="[596,723,433,457]" gridcol="1" gridrow="6" pageId="6" pageNumber="193">0.16–0.16</td>
|
||
<td id="6003D906356EBD8FF16D7CBCFC360DED" box="[782,922,433,457]" gridcol="2" gridrow="6" pageId="6" pageNumber="193">0.16±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7CBCFB240DED" box="[1023,1160,433,457]" gridcol="3" gridrow="6" pageId="6" pageNumber="193">0.17–0.17</td>
|
||
<td id="6003D906356EBD8FF6DA7CBCFAEA0DED" box="[1209,1350,433,457]" gridcol="4" gridrow="6" pageId="6" pageNumber="193">0.17±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57CD4FAEA0DD6" box="[166,1350,473,498]" gridrow="7" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57CD4FE120DD6" box="[166,446,473,498]" gridcol="0" gridrow="7" pageId="6" pageNumber="193">Vertex width, distal margin</th>
|
||
<td id="6003D906356EBD8FF0377CD4FD7F0DD6" box="[596,723,473,498]" gridcol="1" gridrow="7" pageId="6" pageNumber="193">0.08–0.08</td>
|
||
<td id="6003D906356EBD8FF16D7CD4FC360DD6" box="[782,922,473,498]" gridcol="2" gridrow="7" pageId="6" pageNumber="193">0.08±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7CD4FB240DD6" box="[1023,1160,473,498]" gridcol="3" gridrow="7" pageId="6" pageNumber="193">0.08–0.08</td>
|
||
<td id="6003D906356EBD8FF6DA7CD4FAEA0DD6" box="[1209,1350,473,498]" gridcol="4" gridrow="7" pageId="6" pageNumber="193">0.08±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57F0FFAEA0E3E" box="[166,1350,514,538]" gridrow="8" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57F0FFE120E3E" box="[166,446,514,538]" gridcol="0" gridrow="8" pageId="6" pageNumber="193">Pronotum length, midline</th>
|
||
<td id="6003D906356EBD8FF0377F0FFD7F0E3E" box="[596,723,514,538]" gridcol="1" gridrow="8" pageId="6" pageNumber="193">0.14–0.14</td>
|
||
<td id="6003D906356EBD8FF16D7F0FFC360E3E" box="[782,922,514,538]" gridcol="2" gridrow="8" pageId="6" pageNumber="193">0.14±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7F0FFB240E3E" box="[1023,1160,514,538]" gridcol="3" gridrow="8" pageId="6" pageNumber="193">0.16–0.16</td>
|
||
<td id="6003D906356EBD8FF6DA7F0FFAEA0E3E" box="[1209,1350,514,538]" gridcol="4" gridrow="8" pageId="6" pageNumber="193">0.16±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57F27FAEA0E66" box="[166,1350,554,578]" gridrow="9" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57F27FE120E66" box="[166,446,554,578]" gridcol="0" gridrow="9" pageId="6" pageNumber="193">Mesonotum length, midline</th>
|
||
<td id="6003D906356EBD8FF0377F27FD7F0E66" box="[596,723,554,578]" gridcol="1" gridrow="9" pageId="6" pageNumber="193">0.44–0.44</td>
|
||
<td id="6003D906356EBD8FF16D7F27FC360E66" box="[782,922,554,578]" gridcol="2" gridrow="9" pageId="6" pageNumber="193">0.44±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7F27FB240E66" box="[1023,1160,554,578]" gridcol="3" gridrow="9" pageId="6" pageNumber="193">0.46–0.46</td>
|
||
<td id="6003D906356EBD8FF6DA7F27FAEA0E66" box="[1209,1350,554,578]" gridcol="4" gridrow="9" pageId="6" pageNumber="193">0.46±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57F5FFAEA0E4E" box="[166,1350,594,618]" gridrow="10" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57F5FFE120E4E" box="[166,446,594,618]" gridcol="0" gridrow="10" pageId="6" pageNumber="193">Mesonotum width</th>
|
||
<td id="6003D906356EBD8FF0377F5FFD7F0E4E" box="[596,723,594,618]" gridcol="1" gridrow="10" pageId="6" pageNumber="193">0.55–0.56</td>
|
||
<td id="6003D906356EBD8FF16D7F5FFC360E4E" box="[782,922,594,618]" gridcol="2" gridrow="10" pageId="6" pageNumber="193">0.55±0.01</td>
|
||
<td id="6003D906356EBD8FF19C7F5FFB240E4E" box="[1023,1160,594,618]" gridcol="3" gridrow="10" pageId="6" pageNumber="193">0.58–0.58</td>
|
||
<td id="6003D906356EBD8FF6DA7F5FFAEA0E4E" box="[1209,1350,594,618]" gridcol="4" gridrow="10" pageId="6" pageNumber="193">0.58±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57F77FAEA0EB7" box="[166,1350,634,659]" gridrow="11" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57F77FE120EB7" box="[166,446,634,659]" gridcol="0" gridrow="11" pageId="6" pageNumber="193">Frons width, dorsal margin</th>
|
||
<td id="6003D906356EBD8FF0377F77FD7F0EB7" box="[596,723,634,659]" gridcol="1" gridrow="11" pageId="6" pageNumber="193">0.08–0.08</td>
|
||
<td id="6003D906356EBD8FF16D7F77FC360EB7" box="[782,922,634,659]" gridcol="2" gridrow="11" pageId="6" pageNumber="193">0.08±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7F77FB240EB7" box="[1023,1160,634,659]" gridcol="3" gridrow="11" pageId="6" pageNumber="193">0.08–0.08</td>
|
||
<td id="6003D906356EBD8FF6DA7F77FAEA0EB7" box="[1209,1350,634,659]" gridcol="4" gridrow="11" pageId="6" pageNumber="193">0.08±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57FAEFAEA0E9F" box="[166,1350,675,699]" gridrow="12" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57FAEFE120E9F" box="[166,446,675,699]" gridcol="0" gridrow="12" pageId="6" pageNumber="193">Frons width, clypeal suture</th>
|
||
<td id="6003D906356EBD8FF0377FAEFD7F0E9F" box="[596,723,675,699]" gridcol="1" gridrow="12" pageId="6" pageNumber="193">0.09–0.09</td>
|
||
<td id="6003D906356EBD8FF16D7FAEFC360E9F" box="[782,922,675,699]" gridcol="2" gridrow="12" pageId="6" pageNumber="193">0.09±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7FAEFB240E9F" box="[1023,1160,675,699]" gridcol="3" gridrow="12" pageId="6" pageNumber="193">0.09–0.09</td>
|
||
<td id="6003D906356EBD8FF6DA7FAEFAEA0E9F" box="[1209,1350,675,699]" gridcol="4" gridrow="12" pageId="6" pageNumber="193">0.09±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57FC6FAEA0EC7" box="[166,1350,715,739]" gridrow="13" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57FC6FE120EC7" box="[166,446,715,739]" gridcol="0" gridrow="13" pageId="6" pageNumber="193">Frons width, widest</th>
|
||
<td id="6003D906356EBD8FF0377FC6FD7F0EC7" box="[596,723,715,739]" gridcol="1" gridrow="13" pageId="6" pageNumber="193">0.09–0.09</td>
|
||
<td id="6003D906356EBD8FF16D7FC6FC360EC7" box="[782,922,715,739]" gridcol="2" gridrow="13" pageId="6" pageNumber="193">0.09±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7FC6FB240EC7" box="[1023,1160,715,739]" gridcol="3" gridrow="13" pageId="6" pageNumber="193">0.09–0.09</td>
|
||
<td id="6003D906356EBD8FF6DA7FC6FAEA0EC7" box="[1209,1350,715,739]" gridcol="4" gridrow="13" pageId="6" pageNumber="193">0.09±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57FFEFAEA0F2F" box="[166,1350,755,779]" gridrow="14" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57FFEFE120F2F" box="[166,446,755,779]" gridcol="0" gridrow="14" pageId="6" pageNumber="193">Frons width, narrowest</th>
|
||
<td id="6003D906356EBD8FF0377FFEFD7F0F2F" box="[596,723,755,779]" gridcol="1" gridrow="14" pageId="6" pageNumber="193">0.05–0.05</td>
|
||
<td id="6003D906356EBD8FF16D7FFEFC360F2F" box="[782,922,755,779]" gridcol="2" gridrow="14" pageId="6" pageNumber="193">0.05±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7FFEFB240F2F" box="[1023,1160,755,779]" gridcol="3" gridrow="14" pageId="6" pageNumber="193">0.05–0.05</td>
|
||
<td id="6003D906356EBD8FF6DA7FFEFAEA0F2F" box="[1209,1350,755,779]" gridcol="4" gridrow="14" pageId="6" pageNumber="193">0.05±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57E16FAEA0F17" box="[166,1350,795,819]" gridrow="15" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57E16FE120F17" box="[166,446,795,819]" gridcol="0" gridrow="15" pageId="6" pageNumber="193">Frons length, midline</th>
|
||
<td id="6003D906356EBD8FF0377E16FD7F0F17" box="[596,723,795,819]" gridcol="1" gridrow="15" pageId="6" pageNumber="193">0.36–0.37</td>
|
||
<td id="6003D906356EBD8FF16D7E16FC360F17" box="[782,922,795,819]" gridcol="2" gridrow="15" pageId="6" pageNumber="193">0.36±0.01</td>
|
||
<td id="6003D906356EBD8FF19C7E16FB240F17" box="[1023,1160,795,819]" gridcol="3" gridrow="15" pageId="6" pageNumber="193">0.36–0.36</td>
|
||
<td id="6003D906356EBD8FF6DA7E16FAEA0F17" box="[1209,1350,795,819]" gridcol="4" gridrow="15" pageId="6" pageNumber="193">0.36±0.00</td>
|
||
</tr>
|
||
<tr id="23D2B07A356EBD8FF2C57E49FAEA0F78" box="[166,1350,836,860]" gridrow="16" pageId="6" pageNumber="193">
|
||
<th id="6003D906356EBD8FF2C57E49FE120F78" box="[166,446,836,860]" gridcol="0" gridrow="16" pageId="6" pageNumber="193">Clypeus length</th>
|
||
<td id="6003D906356EBD8FF0377E49FD7F0F78" box="[596,723,836,860]" gridcol="1" gridrow="16" pageId="6" pageNumber="193">0.19–0.19</td>
|
||
<td id="6003D906356EBD8FF16D7E49FC360F78" box="[782,922,836,860]" gridcol="2" gridrow="16" pageId="6" pageNumber="193">0.19±0.00</td>
|
||
<td id="6003D906356EBD8FF19C7E49FB240F78" box="[1023,1160,836,860]" gridcol="3" gridrow="16" pageId="6" pageNumber="193">0.19–0.19</td>
|
||
<td id="6003D906356EBD8FF6DA7E49FAEA0F78" box="[1209,1350,836,860]" gridcol="4" gridrow="16" pageId="6" pageNumber="193">0.19±0.00</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="9D5DB238356E4276F2A47E8BFBBB0A54" blockId="6.[151,1437,902,2045]" pageId="6" pageNumber="193">
|
||
Terminalia. Pygofer in lateral view irregular in shape, widest at dorsal margin, narrowing ventrally then expanding at ventral margin, irregularly sinuate on anterior and posterior margin (
|
||
<figureCitation id="05D9AEBD356E4276F6497EA7FB2A0FE1" box="[1066,1158,938,965]" captionStart="FIGURE 5" captionStartId="7.[152,255,1310,1334]" captionTargetBox="[151,1428,182,1279]" captionTargetId="figure-18@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325873" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="6" pageNumber="193">Fig. 5A</figureCitation>
|
||
); in ventral view, posterior margin lacking medioventral process (
|
||
<figureCitation id="05D9AEBD356E4276F01B7EC3FD7D0FCD" box="[632,721,974,1001]" captionStart="FIGURE 5" captionStartId="7.[152,255,1310,1334]" captionTargetBox="[151,1428,182,1279]" captionTargetId="figure-18@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325873" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="6" pageNumber="193">Fig. 5B</figureCitation>
|
||
). Gonostyli in lateral view broadly spatulate with large angular process at base, distally rounded on ventral margin, truncate at apex with slight apical invagination, dorsal margin irregularly sinuate, subapical lobe bearing sclerotized hook, curving distad (
|
||
<figureCitation id="05D9AEBD356E4276F1BC791BFB940814" box="[991,1080,1046,1072]" captionStart="FIGURE 6" captionStartId="7.[152,255,1845,1869]" captionTargetBox="[151,1436,1358,1821]" captionTargetId="figure-43@7.[151,1436,1358,1821]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico villablanca sp. n. gonostylus; A) lateral view and B) inner-ventral view displaying sclerotized ridge." figureDoi="http://doi.org/10.5281/zenodo.7325877" httpUri="https://zenodo.org/record/7325877/files/figure.png" pageId="6" pageNumber="193">Fig. 6A</figureCitation>
|
||
); inner lateral margin with elongate sclerotized ridge with slight serrations, curved mesad with subapical process (
|
||
<figureCitation id="05D9AEBD356E4276F64C7937FB2A0870" box="[1071,1158,1082,1108]" captionStart="FIGURE 6" captionStartId="7.[152,255,1845,1869]" captionTargetBox="[151,1436,1358,1821]" captionTargetId="figure-43@7.[151,1436,1358,1821]" captionTargetPageId="7" captionText="FIGURE 6. Male Tico villablanca sp. n. gonostylus; A) lateral view and B) inner-ventral view displaying sclerotized ridge." figureDoi="http://doi.org/10.5281/zenodo.7325877" httpUri="https://zenodo.org/record/7325877/files/figure.png" pageId="6" pageNumber="193">Fig. 6B</figureCitation>
|
||
); in ventral view, narrow basally, expanded at large angular process, then abruptly constricted, expanded again at distal 2/3 then subapically constricting, apices rounded (
|
||
<figureCitation id="05D9AEBD356E4276F380798FFD9008B9" box="[483,572,1154,1181]" captionStart="FIGURE 5" captionStartId="7.[152,255,1310,1334]" captionTargetBox="[151,1428,182,1279]" captionTargetId="figure-18@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325873" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="6" pageNumber="193">Fig. 5B</figureCitation>
|
||
). Aedeagal shaft upcurved, cylindrical, simple with complex array of mostly retrorse aedeagal and endosomal processes, approximately bilaterally symmetrical; aedeagal processes are a pair of subapical heart-shaped flanges on dorsal margin (A1 & A2) angled cephalad with anterior margin serrate (
|
||
<figureCitation id="05D9AEBD356E4276F73579C7FA3008C0" box="[1366,1436,1226,1252]" captionStart="FIGURE 7" captionStartId="8.[152,255,1424,1448]" captionTargetBox="[151,1437,181,1400]" captionTargetId="figure-24@8.[151,1437,181,1400]" captionTargetPageId="8" captionText="FIGURE 7. Aedeagus of Tico villablanca sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325879" httpUri="https://zenodo.org/record/7325879/files/figure.png" pageId="6" pageNumber="193">Figs 7</figureCitation>
|
||
&
|
||
<figureCitation id="05D9AEBD356E4276F2D779E3FF68092C" box="[180,196,1262,1288]" captionStart="FIGURE 8" captionStartId="9.[152,255,1135,1159]" captionTargetBox="[157,1430,187,1110]" captionTargetId="figure-18@9.[151,1436,181,1112]" captionTargetPageId="9" captionText="FIGURE 8. Line art of aedeagus of Tico villablanca sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view; A = aedeagus process, E = endosoma process, s = secondary, p = primary, d = dorsal and v = ventral." figureDoi="http://doi.org/10.5281/zenodo.7325883" httpUri="https://zenodo.org/record/7325883/files/figure.png" pageId="6" pageNumber="193">8</figureCitation>
|
||
). Aedeagal endosoma very complex with five pairs of processes; first pair (E1 & E2) dorsal most processes in lateral view, slender, moderate length, curving ventrad; second pair (E3 & E4) strongly bifid with secondary processes (E3s & E4s) arising on dorsal margin of primary processes (E3p & E4p), E3s and E4s short, about as thick as E1 and E2, terminating approximately at same point as E1 and E2, curved ventrad; E3p and E4p long, reaching base of aedeagus, laterally flattened (broad in lateral view), dorsal and ventral margins sinuate, apex curved dorsad; third pair of processes (E5 & E6) simple, slender, slightly sinuate, nearly reaching base of aedeagus; fourth pair (E7 & E8) simple, slender, reaching approximately midpoint of E3p and E4p, angled slightly dorsad; final pair (E9 and E10) strongly bifid, dorsal subprocess (E9d & E10d) slender, strongly sinuate, curving dorsad, ventral subprocesses (E9v & E10v) slender, strongly sinuate, curving laterally and dorsad then curving mesad at apex (
|
||
<figureCitation id="05D9AEBD356E4276F6BB7B03FAB30A0C" box="[1240,1311,1550,1576]" captionStart="FIGURE 7" captionStartId="8.[152,255,1424,1448]" captionTargetBox="[151,1437,181,1400]" captionTargetId="figure-24@8.[151,1437,181,1400]" captionTargetPageId="8" captionText="FIGURE 7. Aedeagus of Tico villablanca sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325879" httpUri="https://zenodo.org/record/7325879/files/figure.png" pageId="6" pageNumber="193">Figs 7</figureCitation>
|
||
&
|
||
<figureCitation id="05D9AEBD356E4276F7217B03FAFE0A0C" box="[1346,1362,1550,1576]" captionStart="FIGURE 8" captionStartId="9.[152,255,1135,1159]" captionTargetBox="[157,1430,187,1110]" captionTargetId="figure-18@9.[151,1436,181,1112]" captionTargetPageId="9" captionText="FIGURE 8. Line art of aedeagus of Tico villablanca sp. n.; A) right lateral view, B) left lateral view, and C) dorsal view; A = aedeagus process, E = endosoma process, s = secondary, p = primary, d = dorsal and v = ventral." figureDoi="http://doi.org/10.5281/zenodo.7325883" httpUri="https://zenodo.org/record/7325883/files/figure.png" pageId="6" pageNumber="193">8</figureCitation>
|
||
). Anal segment moderately long, not reaching apex of gonostyli, irregularly sinuate on dorsal and ventral margins, apex elongate, apex blunt, curved slightly ventrad (
|
||
<figureCitation id="05D9AEBD356E4276F0FA7B5BFD5F0A55" box="[665,755,1622,1649]" captionStart="FIGURE 5" captionStartId="7.[152,255,1310,1334]" captionTargetBox="[151,1428,182,1279]" captionTargetId="figure-18@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 5. Male Tico villablanca sp. n. terminalia; A) left lateral view, B) ventral view, and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7325873" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="6" pageNumber="193">Fig. 5A</figureCitation>
|
||
); paraproct short, conical.
|
||
</paragraph>
|
||
<paragraph id="9D5DB238356E4276F2A47B77FCDA0AB0" blockId="6.[151,1437,902,2045]" box="[199,886,1658,1684]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2A47B77FE0F0AB0" bold="true" box="[199,419,1658,1684]" pageId="6" pageNumber="193">Plant associations.</emphasis>
|
||
Undetermined palm species (
|
||
<taxonomicName id="5AE2C9BB356E4276F08C7B77FCC70AB0" box="[751,875,1658,1684]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="193" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5F8E1B3356E4276F2A47B93FDF60A9C" box="[199,602,1694,1720]" pageId="6" pageNumber="193" type="distribution">
|
||
<paragraph id="9D5DB238356E4276F2A47B93FDF60A9C" blockId="6.[151,1437,902,2045]" box="[199,602,1694,1720]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2A47B93FECD0A9C" bold="true" box="[199,353,1694,1720]" pageId="6" pageNumber="193">Distribution.</emphasis>
|
||
<collectingCountry id="E5F5F2A8356E4276F3047B93FE4D0A9C" box="[359,481,1694,1720]" name="Costa Rica" pageId="6" pageNumber="193">Costa Rica</collectingCountry>
|
||
(
|
||
<collectingRegion id="5F267CDA356E4276F3937B93FDFF0A9C" box="[496,595,1694,1720]" country="Costa Rica" name="Alajuela" pageId="6" pageNumber="193">Alajuela</collectingRegion>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5F8E1B3356E4276F2A47BCFFB330AF8" box="[199,1183,1730,1756]" pageId="6" pageNumber="193" type="etymology">
|
||
<paragraph id="9D5DB238356E4276F2A47BCFFB330AF8" blockId="6.[151,1437,902,2045]" box="[199,1183,1730,1756]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2A47BCFFEE20AF8" bold="true" box="[199,334,1730,1756]" pageId="6" pageNumber="193">Etymology.</emphasis>
|
||
The specific name is given in reference to
|
||
<typeStatus id="42590C9A356E4276F1537BCFFCCC0AF8" box="[816,864,1730,1756]" pageId="6" pageNumber="193">type</typeStatus>
|
||
locality, Hotel Villa Blanca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D5F8E1B3356E4276F2A47BEBFB690BD9" pageId="6" pageNumber="193" type="materials_examined">
|
||
<paragraph id="9D5DB238356E4276F2A47BEBFF420B6C" blockId="6.[151,1437,902,2045]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2A47BEBFE1D0B24" bold="true" box="[199,433,1766,1792]" pageId="6" pageNumber="193">Material examined.</emphasis>
|
||
<materialsCitation id="2D8AB865356E4276F3DB7BEBFCF10B00" ID-GBIF-Occurrence="3966323301" collectingDate="2018-05-15" collectionCode="FLREC" country="Costa Rica" location="Hotel Villa Blanca" pageId="6" pageNumber="193" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
|
||
<typeStatus id="42590C9A356E4276F3DB7BEBFD8C0B24" box="[440,544,1766,1792]" pageId="6" pageNumber="193" type="holotype">Holotype</typeStatus>
|
||
male, “
|
||
<collectingCountry id="E5F5F2A8356E4276F0157BEBFD5A0B24" box="[630,758,1766,1792]" name="Costa Rica" pageId="6" pageNumber="193">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingRegion id="5F267CDA356E4276F1627BEBFCF60B24" box="[769,858,1766,1792]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="193">Heredia</collectingRegion>
|
||
/
|
||
<location id="983DE4E3356E4276F1137BEBFBED0B24" LSID="urn:lsid:plazi:treatment:154B032E356B4278F2F47B8EFA300AC9:983DE4E3356E4276F1137BEBFBED0B24" box="[880,1089,1766,1792]" country="Costa Rica" name="Hotel Villa Blanca" pageId="6" pageNumber="193" stateProvince="Heredia">Hotel Villa Blanca</location>
|
||
/
|
||
<date id="E95C94F8356E4276F6347BEBFB650B25" box="[1111,1225,1766,1793]" pageId="6" pageNumber="193" value="2018-05-15">
|
||
<collectingDate id="F9186D10356E4276F6347BEBFB650B25" box="[1111,1225,1766,1793]" pageId="6" pageNumber="193" value="2018-05-15">15.V.2018</collectingDate>
|
||
</date>
|
||
/ Coll.: B.W. Bahder, sweeping palms / Holotype
|
||
<taxonomicName id="5AE2C9BB356E4276F0647A06FD110B00" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[519,701,1802,1828]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="villablanca">
|
||
<emphasis id="AF966E2A356E4276F0647A06FD110B00" box="[519,701,1802,1828]" italics="true" pageId="6" pageNumber="193">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="8BE479B1356E4276F0A67A07FD770B00" box="[709,731,1802,1828]" pageId="6" pageNumber="193" type="male">♂</specimenCount>
|
||
” (
|
||
<collectionCode id="FBF32AFD356E4276F0947A07FCF90B00" box="[759,853,1802,1828]" pageId="6" pageNumber="193">FLREC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2D8AB865356E4276F10A7A07FF460B6C" ID-GBIF-Occurrence="3966323302" collectionCode="FSCA" country="Costa Rica" pageId="6" pageNumber="193" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Heredia" typeStatus="paratype">
|
||
<typeStatus id="42590C9A356E4276F10A7A07FC7A0B00" box="[873,982,1802,1828]" pageId="6" pageNumber="193" type="paratype">Paratypes</typeStatus>
|
||
same as holotype (
|
||
<specimenCount id="8BE479B1356E4276F6D17A07FAB70B00" box="[1202,1307,1802,1828]" pageId="6" pageNumber="193" type="male">10 males</specimenCount>
|
||
,
|
||
<specimenCount id="8BE479B1356E4276F74B7A07FA340B00" box="[1320,1432,1802,1828]" pageId="6" pageNumber="193" type="female">4 females</specimenCount>
|
||
,
|
||
<collectionCode id="FBF32AFD356E4276F2F47A23FF4E0B6C" box="[151,226,1838,1864]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="193" type="Museum">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9D5DB238356E4276F2A47A5FFB690BD9" blockId="6.[151,1437,902,2045]" pageId="6" pageNumber="193">
|
||
<emphasis id="AF966E2A356E4276F2A47A5FFDB00B48" bold="true" box="[199,540,1874,1900]" pageId="6" pageNumber="193">Sequence Data and Analysis.</emphasis>
|
||
For COI, a 681 bp product was generated (GenBank Accession No.
|
||
<accessionNumber id="82B12FDB356E4276F7757A5FFA390B49" box="[1302,1429,1874,1901]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/ON797450" pageId="6" pageNumber="193">ON797450</accessionNumber>
|
||
) and for 18S, a 1,601 bp product was generated (GenBank Accession No.
|
||
<accessionNumber id="82B12FDB356E4276F1AC7A7BFBFD0BB4" box="[975,1105,1910,1936]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/ON783969" pageId="6" pageNumber="193">ON783969</accessionNumber>
|
||
). While support for all clades based on COI is weak, bootstrap support for placement of
|
||
<taxonomicName id="5AE2C9BB356E4276F17C7A96FC780B90" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[799,980,1946,1972]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356E4276F17C7A96FC780B90" box="[799,980,1946,1972]" italics="true" pageId="6" pageNumber="193">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356E4276F1B87A97FBB50B90" bold="true" box="[987,1049,1946,1972]" pageId="6" pageNumber="193">
|
||
<taxonomicNameLabel id="B4A5D351356E4276F1B87A97FBB50B90" box="[987,1049,1946,1972]" pageId="6" pageNumber="193" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
within
|
||
<taxonomicName id="5AE2C9BB356E4276F60E7A96FB310B90" authorityName="Bahder & Bartlett" authorityYear="2021" box="[1133,1181,1947,1972]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF966E2A356E4276F60E7A96FB310B90" box="[1133,1181,1947,1972]" italics="true" pageId="6" pageNumber="193">Tico</emphasis>
|
||
</taxonomicName>
|
||
based on the 18S gene is strong (99) and based on the consensus analysis, there is also strong bootstrap support (100) for the genus
|
||
<taxonomicName id="5AE2C9BB356E4276F7067AB2FA390BFC" authorityName="Bahder & Bartlett" authorityYear="2021" box="[1381,1429,1983,2008]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF966E2A356E4276F7067AB2FA390BFC" box="[1381,1429,1983,2008]" italics="true" pageId="6" pageNumber="193">Tico</emphasis>
|
||
</taxonomicName>
|
||
, where
|
||
<taxonomicName id="5AE2C9BB356E4276F2807AEEFE340BD8" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[227,408,2018,2044]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356E4276F2807AEEFE340BD8" box="[227,408,2018,2044]" italics="true" pageId="6" pageNumber="193">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356E4276F3FC7AEFFE720BD8" bold="true" box="[415,478,2018,2044]" pageId="6" pageNumber="193">
|
||
<taxonomicNameLabel id="B4A5D351356E4276F3FC7AEFFE720BD8" box="[415,478,2018,2044]" pageId="6" pageNumber="193" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resolves adjacent to
|
||
<taxonomicName id="5AE2C9BB356E4276F0AA7AEEFCEF0BD8" box="[713,835,2019,2044]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="sierra">
|
||
<emphasis id="AF966E2A356E4276F0AA7AEEFCEF0BD8" box="[713,835,2019,2044]" italics="true" pageId="6" pageNumber="193">Tico sierra</emphasis>
|
||
</taxonomicName>
|
||
with weak bootstrap support (56).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C99DE2B0356F4277F2FB7813FB430912" ID-DOI="http://doi.org/10.5281/zenodo.7325873" ID-Zenodo-Dep="7325873" box="[152,1263,1310,1334]" httpUri="https://zenodo.org/record/7325873/files/figure.png" pageId="7" pageNumber="194" startId="7.[152,255,1310,1334]" targetBox="[151,1428,182,1279]" targetPageId="7">
|
||
<paragraph id="9D5DB238356F4277F2FB7813FB430912" blockId="7.[152,1263,1310,1334]" box="[152,1263,1310,1334]" pageId="7" pageNumber="194">
|
||
<emphasis id="AF966E2A356F4277F2FB7813FEB40912" bold="true" box="[152,280,1310,1334]" pageId="7" pageNumber="194">FIGURE 5.</emphasis>
|
||
Male
|
||
<taxonomicName id="5AE2C9BB356F4277F33A7812FE510912" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[345,509,1310,1334]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="194" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356F4277F33A7812FE510912" box="[345,509,1310,1334]" italics="true" pageId="7" pageNumber="194">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356F4277F0607813FD900912" bold="true" box="[515,572,1310,1334]" pageId="7" pageNumber="194">
|
||
<taxonomicNameLabel id="B4A5D351356F4277F0607813FD900912" box="[515,572,1310,1334]" pageId="7" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
terminalia; A) left lateral view, B) ventral view, and C) dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C99DE2B0356F4277F2FB7A38FA290B69" ID-DOI="http://doi.org/10.5281/zenodo.7325877" ID-Zenodo-Dep="7325877" box="[152,1413,1845,1869]" httpUri="https://zenodo.org/record/7325877/files/figure.png" pageId="7" pageNumber="194" startId="7.[152,255,1845,1869]" targetBox="[151,1436,1358,1821]" targetPageId="7">
|
||
<paragraph id="9D5DB238356F4277F2FB7A38FA290B69" blockId="7.[152,1413,1845,1869]" box="[152,1413,1845,1869]" pageId="7" pageNumber="194">
|
||
<emphasis id="AF966E2A356F4277F2FB7A38FEB40B69" bold="true" box="[152,280,1845,1869]" pageId="7" pageNumber="194">FIGURE 6.</emphasis>
|
||
Male
|
||
<taxonomicName id="5AE2C9BB356F4277F33A7A3BFE510B69" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[345,509,1845,1869]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="194" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A356F4277F33A7A3BFE510B69" box="[345,509,1845,1869]" italics="true" pageId="7" pageNumber="194">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A356F4277F0607A38FD900B69" bold="true" box="[515,572,1845,1869]" pageId="7" pageNumber="194">
|
||
<taxonomicNameLabel id="B4A5D351356F4277F0607A38FD900B69" box="[515,572,1845,1869]" pageId="7" pageNumber="194" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
gonostylus; A) lateral view and B) inner-ventral view displaying sclerotized ridge.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C99DE2B035604278F2FB789DFB57098C" ID-DOI="http://doi.org/10.5281/zenodo.7325879" ID-Zenodo-Dep="7325879" box="[152,1275,1424,1449]" httpUri="https://zenodo.org/record/7325879/files/figure.png" pageId="8" pageNumber="195" startId="8.[152,255,1424,1448]" targetBox="[151,1437,181,1400]" targetPageId="8">
|
||
<paragraph id="9D5DB23835604278F2FB789DFB57098C" blockId="8.[152,1275,1424,1449]" box="[152,1275,1424,1449]" pageId="8" pageNumber="195">
|
||
<emphasis id="AF966E2A35604278F2FB789DFEB4098D" bold="true" box="[152,280,1424,1449]" pageId="8" pageNumber="195">FIGURE 7.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="5AE2C9BB35604278F3C0789CFDEB098C" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[419,583,1424,1448]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A35604278F3C0789CFDEB098C" box="[419,583,1424,1448]" italics="true" pageId="8" pageNumber="195">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A35604278F02E789CFD2A098D" bold="true" box="[589,646,1425,1449]" pageId="8" pageNumber="195">
|
||
<taxonomicNameLabel id="B4A5D35135604278F02E789CFD2A098D" box="[589,646,1425,1449]" pageId="8" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; A) right lateral view, B) left lateral view, and C) dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D5F8E1B335604278F2A478DAFA300AC9" pageId="8" pageNumber="195" type="discussion">
|
||
<paragraph id="9D5DB23835604278F2A478DAFC8A0AA5" blockId="8.[151,1437,1495,1773]" pageId="8" pageNumber="195">
|
||
<emphasis id="AF966E2A35604278F2A478DAFE9709D5" bold="true" box="[199,315,1495,1521]" pageId="8" pageNumber="195">Remarks.</emphasis>
|
||
Features observed in
|
||
<taxonomicName id="5AE2C9BB35604278F05A78D5FD7809D5" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[569,724,1495,1521]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A35604278F05A78D5FD7809D5" box="[569,724,1495,1521]" italics="true" pageId="8" pageNumber="195">T. villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A35604278F0BE78DAFCB309D5" bold="true" box="[733,799,1495,1521]" pageId="8" pageNumber="195">
|
||
<taxonomicNameLabel id="B4A5D35135604278F0BE78DAFCB309D5" box="[733,799,1495,1521]" pageId="8" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(angular projection at base of gonostyli and sclerotized structure on inner margin of the gonostyli) are consistent with other members of the genus
|
||
<taxonomicName id="5AE2C9BB35604278F6DB78F1FB440A31" authorityName="Bahder & Bartlett" authorityYear="2021" box="[1208,1256,1532,1557]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF966E2A35604278F6DB78F1FB440A31" box="[1208,1256,1532,1557]" italics="true" pageId="8" pageNumber="195">Tico</emphasis>
|
||
</taxonomicName>
|
||
and support its placement therein. Furthermore, molecular support (while weak) places
|
||
<taxonomicName id="5AE2C9BB35604278F1DA7B2DFBFE0A1D" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[953,1106,1567,1593]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A35604278F1DA7B2DFBFE0A1D" box="[953,1106,1567,1593]" italics="true" pageId="8" pageNumber="195">T. villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A35604278F63B7B12FB3B0A1D" bold="true" box="[1112,1175,1567,1593]" pageId="8" pageNumber="195">
|
||
<taxonomicNameLabel id="B4A5D35135604278F63B7B12FB3B0A1D" box="[1112,1175,1567,1593]" pageId="8" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
adjacent to
|
||
<taxonomicName id="5AE2C9BB35604278F77F7B2DFAD50A1D" box="[1308,1401,1568,1593]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="species" species="sierra">
|
||
<emphasis id="AF966E2A35604278F77F7B2DFAD50A1D" box="[1308,1401,1568,1593]" italics="true" pageId="8" pageNumber="195">T. sierra</emphasis>
|
||
</taxonomicName>
|
||
, to which it bears the closest resemblance, superficially based on color. However, there is strong bootstrap support for the monophyly of
|
||
<taxonomicName id="5AE2C9BB35604278F3067B65FE390AA5" authorityName="Bahder & Bartlett" authorityYear="2021" box="[357,405,1640,1665]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AF966E2A35604278F3067B65FE390AA5" box="[357,405,1640,1665]" italics="true" pageId="8" pageNumber="195">Tico</emphasis>
|
||
</taxonomicName>
|
||
based on the loci analyzed thus far.
|
||
</paragraph>
|
||
<paragraph id="9D5DB23835604278F2A47B81FA300AC9" blockId="8.[151,1437,1495,1773]" pageId="8" pageNumber="195">
|
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<taxonomicName id="5AE2C9BB35604278F2A47B81FED70A81" authority="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder, 2022" authorityName="Zumbado Echavarria & Barrantes Barrantes & Helmick & Bartlett & Bahder" authorityYear="2022" box="[199,379,1675,1701]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="species" species="villablanca" status="sp. nov.">
|
||
<emphasis id="AF966E2A35604278F2A47B81FED70A81" box="[199,379,1675,1701]" italics="true" pageId="8" pageNumber="195">Tico villablanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AF966E2A35604278F3E27B86FE130A81" bold="true" box="[385,447,1675,1701]" pageId="8" pageNumber="195">
|
||
<taxonomicNameLabel id="B4A5D35135604278F3E27B86FE130A81" box="[385,447,1675,1701]" pageId="8" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unusual in lacking the sensorial pits on the lateral carinae of the vertex and frons. Since the presence of these pits is usually taken to be a tribal feature, our presumption is that the absence of these pits is a derived feature, an assertion that is supported by the placement of this species as derived in the
|
||
<taxonomicName id="5AE2C9BB35604278F6CC7BDEFA860AC9" authorityName="Muir" authorityYear="1913" box="[1199,1322,1747,1773]" class="Insecta" family="Derbidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="197" phylum="Arthropoda" rank="tribe" tribe="Cenchreini">Cenchreini</taxonomicName>
|
||
(
|
||
<figureCitation id="05D9AEBD35604278F75B7BDEFA230AC9" box="[1336,1423,1747,1773]" captionStart="FIGURE 9" captionStartId="9.[152,255,1707,1731]" captionTargetBox="[154,1428,1223,1682]" captionTargetId="figure-68@9.[151,1436,1220,1684]" captionTargetPageId="9" captionText="FIGURE 9. Maximum Likelihood phylogenetic trees based on 1,000 replicates; A) COI gene, B) 18S gene, and C) consensus of concatenated COI and 18S sequences." figureDoi="http://doi.org/10.5281/zenodo.7325885" httpUri="https://zenodo.org/record/7325885/files/figure.png" pageId="8" pageNumber="195">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |