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<mods:title id="CEA25C3E6C5553DBB4F5060C72A30A2A">A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms in Costa Rica</mods:title>
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<mods:namePart id="8C58EB22FE9379578180B03F28DA930A">Zumbado Echavarria, Marco A.</mods:namePart>
<mods:affiliation id="B3DD3C1F47893EDE7BBEAC6DF828589B">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="3907BA553B483A0DD74D5D891D472CAE">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="FAAB67785FD772724C735307D5804178">Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica.</mods:affiliation>
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<mods:namePart id="162993AC3908CEAD79649E9F181F4B95">Helmick, Ericka E.</mods:namePart>
<mods:affiliation id="079E51E4E9652422E2D2635178D312CA">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. E-mail: ehelmick @ ufl. edu</mods:affiliation>
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<mods:namePart id="FD2A902FB3075BF655594A45676D3A3F">Bartlett, Charles R.</mods:namePart>
<mods:affiliation id="7131CE5A8FA82EC2F6255C1868AC7E4F">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
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<mods:namePart id="AD70B0252F23E3FCF3E550738772CBD3">Bahder, Brian W.</mods:namePart>
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<mods:affiliation id="48AA60A5184C7CEAF161154984280E14">University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. bbahder @ ufl. edu; https: // orcid. org / 0000 - 0002 - 1118 - 4832</mods:affiliation>
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<emphasis id="AF966E2A356B4273F2F47CBAFDFE0DF5" bold="true" box="[151,594,439,465]" pageId="3" pageNumber="190">
Genus
<taxonomicName id="5AE2C9BB356B4273F28D7CBAFDFE0DF5" ID-CoL="9DNFB" authority="Bahder &amp; Bartlett 2021" authorityName="Bahder &amp; Bartlett" authorityYear="2021" box="[238,594,439,465]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="AF966E2A356B4273F28D7CBAFE8F0DF5" bold="true" box="[238,291,439,465]" italics="true" pageId="3" pageNumber="190">Tico</emphasis>
<bibRefCitation id="F973CFC9356B4273F3497CBAFDFE0DF5" author="Bahder, B. W. &amp; Echavarria, M. A. Z. &amp; Barrantes, E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[298,594,439,465]" pageId="3" pageNumber="190" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">Bahder &amp; Bartlett 2021</bibRefCitation>
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<typeStatus id="42590C9A356B4273F2F47CF1FF650E30" box="[151,201,508,532]" pageId="3" pageNumber="190">Type</typeStatus>
species:
<taxonomicName id="5AE2C9BB356B4273F3457CF1FD670E30" authority="Bahder &amp; Bartlett 2021" authorityName="Bahder &amp; Bartlett" authorityYear="2021" box="[294,715,508,532]" class="Insecta" family="Derbidae" genus="Tico" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="emmettcarri">
<emphasis id="AF966E2A356B4273F3457CF1FE780E37" box="[294,468,508,531]" italics="true" pageId="3" pageNumber="190">Tico emmettcarri</emphasis>
<bibRefCitation id="F973CFC9356B4273F3B97CF1FD670E30" author="Bahder, B. W. &amp; Echavarria, M. A. Z. &amp; Barrantes, E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[474,715,508,532]" pageId="3" pageNumber="190" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">Bahder &amp; Bartlett 2021</bibRefCitation>
</taxonomicName>
, by original designation.
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<emphasis id="AF966E2A356B4273F2F47F4CFE280E7F" bold="true" box="[151,388,577,603]" pageId="3" pageNumber="190">Amended Diagnosis</emphasis>
. (Revised from
<bibRefCitation id="F973CFC9356B4273F0547F4FFCA50E78" author="Bahder, B. W. &amp; Echavarria, M. A. Z. &amp; Barrantes, E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[567,777,578,604]" pageId="3" pageNumber="190" pagination="283 - 294" refId="ref5934" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2022) A new species of planthopper in the genus Tico (Hemiptera: Auchenorrhyncha: Derbidae) on palms from lowland tropical rainforest in Costa Rica. Zootaxa, 5125 (3), 283 - 294. https: // doi. org / 10.11646 / zootaxa. 5125.3.2" type="journal article" year="2022">
Bahder
<emphasis id="AF966E2A356B4273F0EC7F4EFD130E78" box="[655,703,578,604]" italics="true" pageId="3" pageNumber="190">et al</emphasis>
. 2022
</bibRefCitation>
.) Small (~2.53.0 mm), cenchreine derbids. Vertex roughly triangular or trapezoidal, widest posteriorly, disc concave, lateral margins keeled, with transverse carinae near fastigium. Frons moderately compressed (length at midline about 4x dorsal width), median carina absent, lateral carinae keeled. Forewing with ScP branch preceding apex of clavus (resulting in long marginal cell). Pygofer opening with distinct lateral expansions, ventral margin of opening without medioventral lobe. Gonostyli long and spatulate, apex rounded, with a medially directed dorsal lobe near midlength (or beyond; medial lobes absent on ventral margin) and sclerotized process on inner margin. Aedeagus nearly bilaterally symmetrical, lacking projections on shaft, except apically, terminating in complex array of retrorse processes. Anal tube elongate, in lateral view ventral margin weakly concave, apex inflected downward, apex truncate; in dorsal view broadly spatulate.
</paragraph>
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<emphasis id="AF966E2A356B4273F2A47E88FEF50FBB" bold="true" box="[199,345,901,927]" pageId="3" pageNumber="190">Distribution</emphasis>
.
<collectingCountry id="E5F5F2A8356B4273F3047E8BFE480F84" box="[359,484,902,928]" name="Costa Rica" pageId="3" pageNumber="190">Costa Rica</collectingCountry>
,
<collectingCountry id="E5F5F2A8356B4273F38C7E8BFDFD0F84" box="[495,593,902,928]" name="Trinidad and Tobago" pageId="3" pageNumber="190">Trinidad</collectingCountry>
,
<collectingCountry id="E5F5F2A8356B4273F03F7E8BFD630F84" box="[604,719,902,928]" name="Venezuela" pageId="3" pageNumber="190">Venezuela</collectingCountry>
(
<bibRefCitation id="F973CFC9356B4273F0BE7E8BFCD80F85" author="Fennah, R. G." box="[733,884,902,929]" pageId="3" pageNumber="190" pagination="109 - 170" refId="ref6296" refString="Fennah, R. G. (1952) On the generic classification of Derbidae (Homoptera: Fulgoroidea) with descriptions of new Neotropical species. Transactions of the Royal Entomological Society of London, 103 (4), 109 - 170. https: // doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01063. x" type="journal article" year="1952">Fennah 1952</bibRefCitation>
,
<bibRefCitation id="F973CFC9356B4273F11C7E8BFBFE0F84" author="Bahder, B. W. &amp; Echavarria, M. A. Z. &amp; Barrantes, E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[895,1106,902,928]" pageId="3" pageNumber="190" pagination="369 - 392" refId="ref5842" refString="Bahder, B. W., Echavarria, M. A. Z., Barrantes, E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2021) A new genus and two new species of planthopper in the tribe Cenchreini (Hemiptera: Auchenorrhyncha: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 4908 (3), 369 - 392. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
Bahder
<emphasis id="AF966E2A356B4273F1B47E8AFBA40F84" box="[983,1032,902,928]" italics="true" pageId="3" pageNumber="190">et al</emphasis>
. 2021
</bibRefCitation>
).
</paragraph>
<paragraph id="9D5DB238356B4273F2A47EA7FA270FE0" blockId="3.[151,1437,577,964]" box="[199,1419,937,964]" pageId="3" pageNumber="190">
<emphasis id="AF966E2A356B4273F2A47EA7FE300FE7" bold="true" box="[199,412,937,964]" pageId="3" pageNumber="190">Plant associations</emphasis>
.
<taxonomicName id="5AE2C9BB356B4273F3C97EA7FCAE0FE0" box="[426,770,938,964]" class="Liliopsida" family="Cyclanthaceae" genus="Asplundia" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="190" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="AF966E2A356B4273F3C97EA7FDB00FE0" box="[426,540,938,964]" italics="true" pageId="3" pageNumber="190">Asplundia</emphasis>
sp. (Cyclanthaceae)
</taxonomicName>
,
<taxonomicName id="5AE2C9BB356B4273F16E7EA7FBF60FE0" box="[781,1114,938,964]" class="Insecta" family="Nymphalidae" genus="Heliconia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="AF966E2A356B4273F16E7EA7FCD70FE0" box="[781,891,938,964]" italics="true" pageId="3" pageNumber="190">Heliconia</emphasis>
sp. (Heliconiaceae)
</taxonomicName>
<taxonomicName id="5AE2C9BB356B4273F6027EA7FA2B0FE0" box="[1121,1415,938,964]" class="Liliopsida" family="Arecaceae" genus="Geonoma" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="190" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="AF966E2A356B4273F6027EA7FB620FE0" box="[1121,1230,938,964]" italics="true" pageId="3" pageNumber="190">Geonoma</emphasis>
sp. (Arecaceae)
</taxonomicName>
.
</paragraph>
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