treatments-xml/data/05/76/94/0576944CFFCCFF992F094CB80C9EF6AF.xml

<document id="F2623D0E61F04BB098F10E9E4A16C707" ID-DOI="10.5281/zenodo.174943" ID-GBIF-Dataset="29840302-24dc-4f5a-8879-63b60cb0ed59" ID-ISSN="1175-5326" ID-Zenodo-Dep="174943" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459791486740" checkinUser="plazi" docAuthor="Sendra, Alberto, Ortuño, Vicente M., Moreno, Agustín, Montagud, Sergio &amp; Teruel, Santiago" docDate="2006" docId="0576944CFFCCFF992F094CB80C9EF6AF" docLanguage="en" docName="zt01372p052.pdf" docOrigin="Zootaxa 1372" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Gollumjapyx smeagol Sendra &amp; Ortuno, sp. nov." docType="treatment" docVersion="10" lastPageNumber="49" masterDocId="F94FEC34FFCBFF972E014A590F65F001" masterDocTitle="Gollumjapyx smeagol gen. n., sp. n., an enigmatic hypogean japygid (Diplura: Japygidae) from the eastern Iberian Peninsula" masterLastPageNumber="52" masterPageNumber="35" pageNumber="42" updateTime="1698219304337" updateUser="plazi">
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<mods:title id="5F66754F575BF94C2F61A703BF3B95BF">Gollumjapyx smeagol gen. n., sp. n., an enigmatic hypogean japygid (Diplura: Japygidae) from the eastern Iberian Peninsula</mods:title>
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<mods:namePart id="97522F7DA0812FB052E14A7523DB528D">Sendra, Alberto</mods:namePart>
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<mods:namePart id="80DC0ED4DBFC008AF856E20FD5316E3F">Ortuño, Vicente M.</mods:namePart>
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<mods:namePart id="5B705135FF4F8CE8EF19E57883360674">Moreno, Agustín</mods:namePart>
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<mods:namePart id="AF93BF5F7B40023DCF235A6BDB53F364">Montagud, Sergio</mods:namePart>
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<mods:namePart id="DF40FF3DF4C79204F504BA30A5E657A5">Teruel, Santiago</mods:namePart>
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<mods:date id="D7DBE624C037716C688D8999414CF40B">2006</mods:date>
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<subSubSection id="C5C576D1FFCCFF902F094CB80C3DF6FB" box="[264,856,1760,1787]" pageId="7" pageNumber="42" type="nomenclature">
<paragraph id="8D60255AFFCCFF902F094CB80C3DF6FB" blockId="7.[264,856,1760,1787]" box="[264,856,1760,1787]" pageId="7" pageNumber="42">
<heading id="D6289236FFCCFF902F094CB80C3DF6FB" bold="true" box="[264,856,1760,1787]" fontSize="11" level="1" pageId="7" pageNumber="42" reason="1">
<emphasis id="BFABF948FFCCFF902F094CB80C3DF6FB" bold="true" box="[264,856,1760,1787]" pageId="7" pageNumber="42">
<taxonomicName id="4ADF5ED9FFCCFF902F094CB80D91F6FB" authority="Sendra &amp; Ortuno" authorityName="Sendra &amp; Ortuno" box="[264,756,1760,1787]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="smeagol" status="sp. nov.">
<emphasis id="BFABF948FFCCFF902F094CB80D6AF6FA" bold="true" box="[264,527,1761,1787]" italics="true" pageId="7" pageNumber="42">Gollumjapyx smeagol</emphasis>
Sendra &amp; Ortuño
</taxonomicName>
<taxonomicNameLabel id="A4984433FFCCFF902CFA4CB90C3DF6FB" box="[763,856,1760,1786]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5C576D1FFCCFF9F2F094D720CF1F2EF" lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42" type="materials_examined">
<paragraph id="8D60255AFFCCFF9F2F094D720B5CF137" blockId="7.[264,1323,1835,1901]" lastBlockId="8.[264,1323,284,910]" lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42">
<emphasis id="BFABF948FFCCFF902F094D720E1FF744" bold="true" box="[264,378,1835,1861]" pageId="7" pageNumber="42">
<typeStatus id="52649BF8FFCCFF902F094D720E1FF744" box="[264,378,1835,1861]" pageId="7" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
: Ψ 
<quantity id="4A2788BFFFCCFF902FA84D720D47F744" box="[425,546,1835,1861]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="7" pageNumber="42" unit="mm" value="23.4">23.40 mm</quantity>
(cerci inclusive) nº 0 815, Castellón, Serra d’en Galceran, “Cova del Mas de Gaspar” cave; 
<date id="F961039AFFCCFF902C3D4D0A0DDBF76C" box="[572,702,1875,1901]" pageId="7" pageNumber="42" value="1980-05-26">26-V-1980</date>
; J. Comas &amp; O. Escolà leg.; preserved in ethanol 70º + glycerine and deposited at the Barcelona Zoology Museum (
<collectingCountry id="F5C865CAFFC3FF9F2DE54B450B48F137" box="[996,1069,284,310]" name="Spain" pageId="8" pageNumber="43">Spain</collectingCountry>
).
</paragraph>
<paragraph id="8D60255AFFC3FF9F2F394B1A0B57F2C7" blockId="8.[264,1323,284,910]" pageId="8" pageNumber="43">
<emphasis id="BFABF948FFC3FF9F2F394B1A0EDBF15C" bold="true" box="[312,446,323,349]" pageId="8" pageNumber="43">
<typeStatus id="52649BF8FFC3FF9F2F394B1A0EDFF15C" box="[312,442,323,349]" pageId="8" pageNumber="43" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 Ψ, Tarragona, Ulldecona, “Avenc Canals” pit; 
<date id="F961039AFFC3FF9F2A0A4B1D0BF3F15F" box="[1035,1174,324,350]" pageId="8" pageNumber="43" value="2000-03-11">11-III-2000</date>
; F. Fadrique leg. 1 Ψ, Castellón, Cabanes, “Avenc d’en Serenge” pit; 
<date id="F961039AFFC3FF9F2D994B350B41F187" box="[920,1060,364,390]" pageId="8" pageNumber="43" value="1982-03-14">14-III-1982</date>
; J. Comas leg. 1 sex?, idem; 
<date id="F961039AFFC3FF9F2F554BCD0E87F1AF" box="[340,482,404,430]" pageId="8" pageNumber="43" value="1995-11-12">12-XI-1995</date>
; E. Carabajal leg. 1 ɗ, idem; 
<date id="F961039AFFC3FF9F2D4F4BCD0C80F1AF" box="[846,997,404,430]" pageId="8" pageNumber="43" value="2003-07-19">19-VII-2003</date>
; S. Montagud leg. 2 sex?, idem; 
<date id="F961039AFFC3FF9F2F534BE50EB1F1D7" box="[338,468,444,470]" pageId="8" pageNumber="43" value="2006-02-26">26-II-2006</date>
; S. Montagud &amp; A. Sendra leg. 1 ɗ, Castellón, Cabanes, “Avenc Mas de la Cova” pit; 
<date id="F961039AFFC3FF9F2FAC4BBD0D20F1FF" box="[429,581,484,510]" pageId="8" pageNumber="43" value="1995-07-22">22-VII-1995</date>
; E. Carabajal leg. 1 ɗ, Castellón, Coves de Vinromá, “Avenc Mas Nou” pit; 
<date id="F961039AFFC3FF9F2FC748550D28F227" box="[454,589,524,550]" pageId="8" pageNumber="43" value="2005-03-12">12.III.2005</date>
; S. Teruel leg. 1 sex?, Castellón, Sant Mateu, “Cova dels Encenalls” cave; 
<date id="F961039AFFC3FF9F2FD7486D0D31F24F" box="[470,596,564,590]" pageId="8" pageNumber="43" value="2006-03-09">9-III-2006</date>
; J. Comas leg. Preserved in ethanol 70º or 96º and deposited at: Muséum d’Histoire Naturelle Genève, 
<collectingCountry id="F5C865CAFFC3FF9F2D1C48050CC8F277" box="[797,941,604,630]" name="Switzerland" pageId="8" pageNumber="43">Switzerland</collectingCountry>
; National Museum of Natural Sciences, Madrid, 
<collectingCountry id="F5C865CAFFC3FF9F2FF248DD0D5CF29F" box="[499,569,644,670]" name="Spain" pageId="8" pageNumber="43">Spain</collectingCountry>
; Barcelona Zoology Museum (
<collectingCountry id="F5C865CAFFC3FF9F2DC748DD0B69F29F" box="[966,1036,644,670]" name="Spain" pageId="8" pageNumber="43">Spain</collectingCountry>
) and Valencia Natural History Museum (
<collectingCountry id="F5C865CAFFC3FF9F2FDC48F50D46F2C7" box="[477,547,684,710]" name="Spain" pageId="8" pageNumber="43">Spain</collectingCountry>
) — Fundación Entomolólogica Torres Sala-.
</paragraph>
<paragraph id="8D60255AFFC3FF9F2F09488D0CF1F2EF" blockId="8.[264,1323,284,910]" box="[264,916,723,750]" pageId="8" pageNumber="43">
Total 
<typeStatus id="52649BF8FFC3FF9F2F4A488D0EA5F2EF" box="[331,448,724,750]" pageId="8" pageNumber="43" type="paratype">paratypes</typeStatus>
: 
<specimenCount id="9BD9EED3FFC3FF9F2FCB488D0D3AF2EF" box="[458,607,724,750]" pageId="8" pageNumber="43" type="generic">9 specimens</specimenCount>
; 3 ɗ ɗ, 2 Ψ Ψ and 4 sex?.
</paragraph>
</subSubSection>
<subSubSection id="C5C576D1FFC3FF9F2F3948A50D3FF6CC" pageId="8" pageNumber="43" type="etymology">
<paragraph id="8D60255AFFC3FF9F2F3948A50DD9F33F" blockId="8.[264,1323,284,910]" pageId="8" pageNumber="43">
<emphasis id="BFABF948FFC3FF9F2F3948A50EA7F317" bold="true" box="[312,450,764,790]" pageId="8" pageNumber="43">
<emphasis id="BFABF948FFC3FF9F2F3948A50EDDF317" bold="true" box="[312,440,764,790]" italics="true" pageId="8" pageNumber="43">Etymology</emphasis>
:
</emphasis>
the specific epithet refers to Gollum’s original name, who, in his epigean origin, was a hobbit named 
<taxonomicName id="4ADF5ED9FFC3FF9F2C4C497D0DD2F33F" box="[589,695,804,830]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="smeagol">Smeagol</taxonomicName>
.
</paragraph>
<paragraph id="8D60255AFFC3FF9F2F3949150BB9F38F" blockId="8.[264,1323,284,910]" pageId="8" pageNumber="43">
DNA barcode: A nucleotide sequence of the 5' half of the cytochrome c oxydase subunit I (cox1) has been deposited in Genbank with accession number 
<accessionNumber id="928CB8B9FFC3FF9F2A53492D0BBDF38F" box="[1106,1240,884,910]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ993154" pageId="8" pageNumber="43">DQ993154</accessionNumber>
.
</paragraph>
<caption id="D9A075D2FFC3FF9F2F09499A0B42F3FF" ID-Table-UUID="D9A075D2FFC3FF9F2F09499A0B42F3FF" httpUri="http://table.plazi.org/id/D9A075D2FFC3FF9F2F09499A0B42F3FF" pageId="8" pageNumber="43" targetBox="[272,1305,1097,1283]" targetIsTable="true" targetPageId="8">
<paragraph id="8D60255AFFC3FF9F2F09499A0B42F3FF" blockId="8.[264,1323,963,1022]" pageId="8" pageNumber="43">
<emphasis id="BFABF948FFC3FF9F2F09499A0E1AF3DA" bold="true" box="[264,383,963,987]" pageId="8" pageNumber="43">TABLE 1.</emphasis>
Length (in mm) of the body, appendixes and number of antennomeres found in the holotype and paratypes of 
<taxonomicName id="4ADF5ED9FFC3FF9F2C2249BE0CAFF3FC" authority="Sendra &amp; Ortuno" authorityName="Sendra &amp; Ortuno" box="[547,970,998,1022]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="smeagol" status="sp. nov.">
<emphasis id="BFABF948FFC3FF9F2C2249BE0C69F3FF" box="[547,780,999,1022]" italics="true" pageId="8" pageNumber="43">Gollumjapyx smeagol</emphasis>
Sendra &amp; Ortuño
</taxonomicName>
<taxonomicNameLabel id="A4984433FFC3FF9F2DD049BF0B42F3FF" box="[977,1063,998,1022]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8D60255AFFC3FF9F2C3C4E470BD6F5AC" blockId="8.[573,1305,1054,1077]" lastBlockId="8.[272,1241,1430,1573]" pageId="8" pageNumber="43">Body* Cerci** Antennae** Antennomeres** Legs III** sex?, Avenc d’en Serenge broken and - 8.0 5 55 -</paragraph>
<paragraph id="8D60255AFFC3FF9F2F114E100BBDF502" pageId="8" pageNumber="43">
<table id="FFDFD7FAFFC300682F114E100A7CF502" box="[272,1305,1097,1283]" gridcols="6" gridrows="4" pageId="8" pageNumber="43">
<tr id="33EF2718FFC300682F114E100A7CF461" box="[272,1305,1097,1120]" gridrow="0" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114E100D6DF461" box="[272,520,1097,1120]" gridcol="0" gridrow="0" pageId="8" pageNumber="43">Ψ, Avenc Canals</th>
<th id="703E4E64FFC300682C3C4E100D1CF461" box="[573,633,1097,1120]" gridcol="1" gridrow="0" pageId="8" pageNumber="43">21.20</th>
<th id="703E4E64FFC300682CDC4E100C68F461" box="[733,781,1097,1120]" gridcol="2" gridrow="0" pageId="8" pageNumber="43">2.90</th>
<th id="703E4E64FFC300682D434E100CE9F461" box="[834,908,1097,1120]" gridcol="3" gridrow="0" pageId="8" pageNumber="43">8.20</th>
<th id="703E4E64FFC300682DE34E100C98F461" box="[994,1021,1097,1120]" gridcol="4" gridrow="0" pageId="8" pageNumber="43">53</th>
<th id="703E4E64FFC300682AAB4E100A7CF461" box="[1194,1305,1097,1120]" gridcol="5" gridrow="0" pageId="8" pageNumber="43">9.75</th>
</tr>
<tr id="33EF2718FFC300682F114E2A0A7CF48A" box="[272,1305,1139,1163]" gridrow="1" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114E2A0D6DF48A" box="[272,520,1139,1163]" gridcol="0" gridrow="1" pageId="8" pageNumber="43">Ψ, holotype</th>
<td id="703E4E64FFC300682C3C4E2A0D1CF48A" box="[573,633,1139,1163]" gridcol="1" gridrow="1" pageId="8" pageNumber="43">20.15</td>
<td id="703E4E64FFC300682CDC4E2A0C68F48A" box="[733,781,1139,1163]" gridcol="2" gridrow="1" pageId="8" pageNumber="43">3.25</td>
<td id="703E4E64FFC300682D434E2A0CE9F48A" box="[834,908,1139,1163]" gridcol="3" gridrow="1" pageId="8" pageNumber="43">9.00</td>
<td id="703E4E64FFC300682DE34E2A0C98F48A" box="[994,1021,1139,1163]" gridcol="4" gridrow="1" pageId="8" pageNumber="43">54</td>
<td id="703E4E64FFC300682AAB4E2A0A7CF48A" box="[1194,1305,1139,1163]" gridcol="5" gridrow="1" pageId="8" pageNumber="43">9.79</td>
</tr>
<tr id="33EF2718FFC300682F114EC70A7CF4D9" box="[272,1305,1182,1240]" gridrow="2" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114EC70D6DF4D9" box="[272,520,1182,1240]" gridcol="0" gridrow="2" pageId="8" pageNumber="43">♂, Avenc del Mas de la Cova</th>
<td id="703E4E64FFC300682C3C4EC70D1CF4D9" box="[573,633,1182,1240]" gridcol="1" gridrow="2" pageId="8" pageNumber="43">19.40</td>
<td id="703E4E64FFC300682CDC4EC70C68F4D9" box="[733,781,1182,1240]" gridcol="2" gridrow="2" pageId="8" pageNumber="43">2.20</td>
<td id="703E4E64FFC300682D434EC70CE9F4D9" box="[834,908,1182,1240]" gridcol="3" gridrow="2" pageId="8" pageNumber="43">broken</td>
<td id="703E4E64FFC300682DE34EC70C98F4D9" box="[994,1021,1182,1240]" gridcol="4" gridrow="2" pageId="8" pageNumber="43">-</td>
<td id="703E4E64FFC300682AAB4EC70A7CF4D9" box="[1194,1305,1182,1240]" gridcol="5" gridrow="2" pageId="8" pageNumber="43">broken</td>
</tr>
<tr id="33EF2718FFC300682F114EB20A7CF502" box="[272,1305,1259,1283]" gridrow="3" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114EB20D6DF502" box="[272,520,1259,1283]" gridcol="0" gridrow="3" pageId="8" pageNumber="43">Ψ, Avenc d’en Serenge</th>
<td id="703E4E64FFC300682C3C4EB20D1CF502" box="[573,633,1259,1283]" gridcol="1" gridrow="3" pageId="8" pageNumber="43">17.00</td>
<td id="703E4E64FFC300682CDC4EB20C68F502" box="[733,781,1259,1283]" gridcol="2" gridrow="3" pageId="8" pageNumber="43">2.40</td>
<td id="703E4E64FFC300682D434EB20CE9F502" box="[834,908,1259,1283]" gridcol="3" gridrow="3" pageId="8" pageNumber="43">8.8</td>
<td id="703E4E64FFC300682DE34EB20C98F502" box="[994,1021,1259,1283]" gridcol="4" gridrow="3" pageId="8" pageNumber="43">55</td>
<td id="703E4E64FFC300682AAB4EB20A7CF502" box="[1194,1305,1259,1283]" gridcol="5" gridrow="3" pageId="8" pageNumber="43">8.75</td>
</tr>
</table>
</paragraph>
<paragraph id="8D60255AFFC3FF9F2F114F4F0BBCF582" pageId="8" pageNumber="43">
<table id="FFDFD7FAFFC300682F114F4F0A7CF582" box="[272,1305,1302,1411]" gridcols="4" gridrows="3" pageId="8" pageNumber="43">
<tr id="33EF2718FFC300682F114F4F0A7CF52C" box="[272,1305,1302,1325]" gridrow="0" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114F4F0D1CF52C" box="[272,633,1302,1325]" gridcol="0" gridrow="0" pageId="8" pageNumber="43">sex?, Avenc d’en Serenge 16.40</th>
<th id="703E4E64FFC300682CDF4F4F0C14F52C" box="[734,881,1302,1325]" gridcol="1" gridrow="0" pageId="8" pageNumber="43">2.20 8.50</th>
<th id="703E4E64FFC300682DE34F4F0C98F52C" box="[994,1021,1302,1325]" gridcol="2" gridrow="0" pageId="8" pageNumber="43">54</th>
<th id="703E4E64FFC300682AAB4F4F0A7CF52C" box="[1194,1305,1302,1325]" gridcol="3" gridrow="0" pageId="8" pageNumber="43">8.66</th>
</tr>
<tr id="33EF2718FFC300682F114F180A7CF559" box="[272,1305,1345,1368]" gridrow="1" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114F180D1CF559" box="[272,633,1345,1368]" gridcol="0" gridrow="1" pageId="8" pageNumber="43">sex?, Cova dels Encenalls 16.00</th>
<td id="703E4E64FFC300682CDF4F180C14F559" box="[734,881,1345,1368]" gridcol="1" gridrow="1" pageId="8" pageNumber="43">1.95 7.20</td>
<td id="703E4E64FFC300682DE34F180C98F559" box="[994,1021,1345,1368]" gridcol="2" gridrow="1" pageId="8" pageNumber="43">53</td>
<td id="703E4E64FFC300682AAB4F180A7CF559" box="[1194,1305,1345,1368]" gridcol="3" gridrow="1" pageId="8" pageNumber="43">7.30</td>
</tr>
<tr id="33EF2718FFC300682F114F350A7CF582" box="[272,1305,1388,1411]" gridrow="2" pageId="8" pageNumber="43">
<th id="703E4E64FFC300682F114F350D1CF582" box="[272,633,1388,1411]" gridcol="0" gridrow="2" pageId="8" pageNumber="43">sex?, Avenc d’en Serenge 14.80</th>
<td id="703E4E64FFC300682CDF4F350C14F582" box="[734,881,1388,1411]" gridcol="1" gridrow="2" pageId="8" pageNumber="43">2.10 8.30</td>
<td id="703E4E64FFC300682DE34F350C98F582" box="[994,1021,1388,1411]" gridcol="2" gridrow="2" pageId="8" pageNumber="43">54</td>
<td id="703E4E64FFC300682AAB4F350A7CF582" box="[1194,1305,1388,1411]" gridcol="3" gridrow="2" pageId="8" pageNumber="43">7.62</td>
</tr>
</table>
</paragraph>
<paragraph id="8D60255AFFC3FF9F2C3C4FE00DADF5D1" blockId="8.[272,1241,1430,1573]" box="[573,712,1465,1488]" pageId="8" pageNumber="43">not complete</paragraph>
<paragraph id="8D60255AFFC3FF9F2F114FBA0BBCF5FA" blockId="8.[272,1241,1430,1573]" box="[272,1241,1507,1531]" pageId="8" pageNumber="43">♂, Avenc Mas Nou 13.26 1.50 7.40 53 4.81</paragraph>
<paragraph id="8D60255AFFC3FF9F2F114C570BBDF624" blockId="8.[272,1241,1430,1573]" box="[272,1240,1550,1573]" pageId="8" pageNumber="43">♂, Avenc d’en Serenge 12.00 1.17 6.75 54 4.76</paragraph>
<paragraph id="8D60255AFFC3FF9F2F094C170D26F664" blockId="8.[264,1323,1614,1741]" box="[264,579,1614,1637]" pageId="8" pageNumber="43">* Length body without cerci.</paragraph>
<paragraph id="8D60255AFFC3FF9F2F094C280D3FF6CC" blockId="8.[264,1323,1614,1741]" pageId="8" pageNumber="43">** Measurements taken of the two appendixes from the same specimen, although only the longest is recorded. In the case of two intact antennae from the same specimen, only the antennae with the most antennomeres is recorded.</paragraph>
</subSubSection>
<subSubSection id="C5C576D1FFC3FF9B2F394CA50EADF70B" lastPageId="12" lastPageNumber="47" pageId="8" pageNumber="43" type="description">
<paragraph id="8D60255AFFC3FF9F2F394CA50A4EF767" blockId="8.[264,1323,1788,1894]" pageId="8" pageNumber="43">
<emphasis id="BFABF948FFC3FF9F2F394CA50EB7F717" bold="true" box="[312,466,1788,1814]" pageId="8" pageNumber="43">Description:</emphasis>
Long body, maximum and minimum body length (excluding cerci) from 21.20 to 12.00 mm. Other lengths described in 
<tableCitation id="C05D10E1FFC3FF9F2D2C4D7D0CE0F73F" box="[813,901,1828,1854]" captionStart="TABLE 1" captionStartId="8.[264,352,963,987]" captionTargetBox="[272,1305,1097,1283]" captionTargetPageId="8" captionText="TABLE 1. Length (in mm) of the body, appendixes and number of antennomeres found in the holotype and paratypes of Gollumjapyx smeagol Sendra &amp; Ortuño sp. nov." httpUri="http://table.plazi.org/id/D9A075D2FFC3FF9F2F09499A0B42F3FF" pageId="8" pageNumber="43" tableUuid="D9A075D2FFC3FF9F2F09499A0B42F3FF">Table 1</tableCitation>
. All three thoracic segments show a remarkable lengthening more obvious in mesothoracic and metathoracic prescutum.
</paragraph>
<paragraph id="8D60255AFFC2FF9E2F094B450D19F227" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
Almost totally unpigmented cuticle. More or less apparent sclerotization of buccal pieces, anterior margin of head, claws and a large portion of urite X, becoming particularly evident in cerci. Abdominal segments clearly show a lateral expansion in the tergi, increasingly apparent from segment I to VII (
<figureCitation id="15E439DFFFC2FF9E2D1F4BCD0C06F1AF" box="[798,867,404,430]" captionStart="FIGURE 1" captionStartId="4.[264,367,1867,1891]" captionTargetBox="[468,1103,375,1750]" captionTargetId="figure@4.[351,1233,319,1845]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n. (scale: 3 mm)." httpUri="https://zenodo.org/record/174944/files/figure.png" pageId="9" pageNumber="44">Fig. 1</figureCitation>
and 
<figureCitation id="15E439DFFFC2FF9E2D9C4BCD0CA1F1AF" box="[925,964,404,430]" captionStart="FIGURE 2. a" captionStartId="5.[264,367,1746,1770]" captionTargetBox="[264,1323,313,1734]" captionTargetId="figure@5.[264,1323,313,1734]" captionTargetPageId="5" captionText="FIGURE 2. a. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., Avenc d’en Serenge cave from Cabanes, Castellón (Spain); b. detail of the contents from the digestive tract belonging to a specimen from Avenc d’en Serenge; all the fragments belong to a male of Speleotyphlus aurouxi (Carabidae, Anillini). Terms: a, antennomeres; rgs, ring of the genital segment; ml, median lobe; p, paramere; t, tarsomeres. (scales, a: 3 mm, b: 0.3 mm)." httpUri="https://zenodo.org/record/174945/files/figure.png" pageId="9" pageNumber="44">2 a</figureCitation>
). Ordinary setae noticeable on head, pronotum and mesonotum, more abundant in larger specimens. These ordinary setae are shorter in the mesonotum, becoming even shorter and sparse in the metanotum and in the abdominal sclerites (
<figureCitation id="15E439DFFFC2FF9E2C2748550D0AF227" box="[550,623,524,550]" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8D60255AFFC2FF9E2F39486D0D85F42F" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
Head. Antennae with 53 to 55 antennomeres, measuring around half of body length (
<tableCitation id="C05D10E1FFC2FF9E2F1148050E03F277" box="[272,358,604,630]" captionStart="TABLE 1" captionStartId="8.[264,352,963,987]" captionTargetBox="[272,1305,1097,1283]" captionTargetPageId="8" captionText="TABLE 1. Length (in mm) of the body, appendixes and number of antennomeres found in the holotype and paratypes of Gollumjapyx smeagol Sendra &amp; Ortuño sp. nov." httpUri="http://table.plazi.org/id/D9A075D2FFC3FF9F2F09499A0B42F3FF" pageId="9" pageNumber="44" tableUuid="D9A075D2FFC3FF9F2F09499A0B42F3FF">Table 1</tableCitation>
); 13 typical trichobothria on antennomeres IV–VI (1 ventral and 2 dorsal in IV), 
<emphasis id="BFABF948FFC2FF9E2B1D48050A4EF274" box="[1308,1323,604,629]" italics="true" pageId="9" pageNumber="44">a</emphasis>
clearly distal; all antennomeres generally pilose (
<figureCitation id="15E439DFFFC2FF9E2D6A48DD0CD2F29F" box="[875,951,644,670]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="9" pageNumber="44">Fig. 3</figureCitation>
b); antennomeres II to IV are largest and from antennomere V tapering to penultimate antennomere; pilose patches dense with long, robust setae, arranged in two whorls in proximal antennomeres, tapering to one whorl in distal antennomeres. These long and robust setae are accompanied by many thin setae, clearly shorter, covering almost all of each antennomere. Joining the pilose patch are many long, fine and hooked setae-shaped sensilla and also a few fine and short sensilla (
<figureCitation id="15E439DFFFC2FF9E2FB2492D0E9FF38F" box="[435,506,884,910]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="9" pageNumber="44">Fig. 3</figureCitation>
a). Placoid sensilla of apical antennomere 
<quantity id="4A2788BFFFC2FF9E2DEF492D0B0AF38F" box="[1006,1135,884,910]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="9" pageNumber="44" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14 to 16 in</quantity>
number, in two random groups (
<figureCitation id="15E439DFFFC2FF9E2FCC49C50D73F3B7" box="[461,534,924,950]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="9" pageNumber="44">Fig. 3</figureCitation>
a). One or two placoid sensilla in the penultimate antennomere in some of the studied specimens. In three of the specimens from “Avenc d’en Serenge” placoid sensilla appear from antennomere XI or XII, ventrally and the next antennomeres show 0, 1 or 2 placoid sensilla (
<figureCitation id="15E439DFFFC2FF9E2C7A4E4D0DA4F42F" box="[635,705,1044,1070]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="9" pageNumber="44">Fig. 3</figureCitation>
b).
</paragraph>
<paragraph id="8D60255AFFC2FF9E2F394E650EB8F547" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
Simple vertex setae increase noticeably the larger the body size of the specimen (
<figureCitation id="15E439DFFFC2FF9E2AFA4E650E7DF47F" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
a, b). However, the number and pattern of the large setae does not vary in the specimens under study. A comparison with the pattern of large setae proposed by 
<bibRefCitation id="E94E58ABFFC2FF9E2A474ED50B85F4A7" author="Pages" box="[1094,1248,1164,1190]" pageId="9" pageNumber="52" refString="Pages, J. (1984) Dicellurata Genavensia XIII. Japygides du Sud-Est asiatique Nº 4. Revue suisse Zoologie, 91, 329 - 368." type="journal article" year="1984">Pagés (1984)</bibRefCitation>
in the case of 
<taxonomicName id="4ADF5ED9FFC2FF9E2F644EED0DB2F4CF" authority="Pages, 1984" authorityName="Pages" authorityYear="1984" box="[357,727,1204,1230]" class="Entognatha" family="Japygidae" genus="Indjapyx" kingdom="Animalia" order="Diplura" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="uvaianus">
<emphasis id="BFABF948FFC2FF9E2F644EED0D5AF4CC" box="[357,575,1204,1229]" italics="true" pageId="9" pageNumber="44">Indjapyx uvaianus</emphasis>
<bibRefCitation id="E94E58ABFFC2FF9E2C494EED0DB2F4CF" author="Pages" box="[584,727,1204,1230]" pageId="9" pageNumber="52" refString="Pages, J. (1984) Dicellurata Genavensia XIII. Japygides du Sud-Est asiatique Nº 4. Revue suisse Zoologie, 91, 329 - 368." type="journal article" year="1984">Pagés, 1984</bibRefCitation>
</taxonomicName>
proved complex; nonetheless the setae have been numbered, comparing their pattern of distribution. Out of 40 large setae proposed for 
<taxonomicName id="4ADF5ED9FFC2FF9E2B184E850E17F51C" class="Entognatha" family="Japygidae" genus="Indjapyx" kingdom="Animalia" order="Diplura" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="uvaianus">
<emphasis id="BFABF948FFC2FF9E2B184E850E17F51C" italics="true" pageId="9" pageNumber="44">I. uvaianus</emphasis>
</taxonomicName>
distributed in seven groups (A, S, V, M, I, L y P), only 14 seem to appear in 
<taxonomicName id="4ADF5ED9FFC2FF9E2B134F5D0E0CF544" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC2FF9E2B134F5D0E0CF544" italics="true" pageId="9" pageNumber="44">G. smeagol</emphasis>
</taxonomicName>
(
<figureCitation id="15E439DFFFC2FF9E2F7B4F750EA7F547" box="[378,450,1324,1350]" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
a).
</paragraph>
<paragraph id="8D60255AFFC2FF9E2F394F0D0C6FF65F" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
Buccal pieces are consistent with the family: five laminae of the internal lobe of the maxilae with all five laminae pectinate; labial palpi 3.70 times longer than wide at base (423 μm 
<typeStatus id="52649BF8FFC2FF9E2F7C4FFD0E80F5BF" box="[381,485,1444,1470]" pageId="9" pageNumber="44" type="holotype">holotype</typeStatus>
) covered with long setae; submentum with 1+1 large setae (almost equivalent to real macrosetae), postmentum with 1+1 large setae in the internal margin of the labial palpus; admentum with some large setae of which 2+2 are more noticeable; prementum (external lobe) with many setae among which a dozen small short setae. All large setae are longer than the labial palpus.
</paragraph>
<paragraph id="8D60255AFFC2FF9D2F394C350B5BF15F" blockId="9.[264,1323,284,1870]" lastBlockId="10.[264,1323,284,1036]" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44">
Thorax. Extremely elongate (
<figureCitation id="15E439DFFFC2FF9E2C964C350D85F687" box="[663,736,1644,1670]" captionStart="FIGURE 1" captionStartId="4.[264,367,1867,1891]" captionTargetBox="[468,1103,375,1750]" captionTargetId="figure@4.[351,1233,319,1845]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n. (scale: 3 mm)." httpUri="https://zenodo.org/record/174944/files/figure.png" pageId="9" pageNumber="44">Fig. 1</figureCitation>
and 
<figureCitation id="15E439DFFFC2FF9E2D1F4C350C58F687" box="[798,829,1644,1670]" captionStart="FIGURE 2. a" captionStartId="5.[264,367,1746,1770]" captionTargetBox="[264,1323,313,1734]" captionTargetId="figure@5.[264,1323,313,1734]" captionTargetPageId="5" captionText="FIGURE 2. a. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., Avenc d’en Serenge cave from Cabanes, Castellón (Spain); b. detail of the contents from the digestive tract belonging to a specimen from Avenc d’en Serenge; all the fragments belong to a male of Speleotyphlus aurouxi (Carabidae, Anillini). Terms: a, antennomeres; rgs, ring of the genital segment; ml, median lobe; p, paramere; t, tarsomeres. (scales, a: 3 mm, b: 0.3 mm)." httpUri="https://zenodo.org/record/174945/files/figure.png" pageId="9" pageNumber="44">2a</figureCitation>
). Pronotum (
<figureCitation id="15E439DFFFC2FF9E2DDB4C350B40F687" box="[986,1061,1644,1670]" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
c): typical 5+5 M and 1+1 M posterior lateral. Ordinary setae abundant, increasing relative to body length. Mesonotum (
<figureCitation id="15E439DFFFC2FF9E2FAB4CE50E9DF6D7" box="[426,504,1724,1750]" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
d): prescutum with few setae longer than ordinary (0+1 M in the 
<typeStatus id="52649BF8FFC2FF9E2F094CBD0E08F6FF" box="[264,365,1764,1790]" pageId="9" pageNumber="44" type="paratype">paratype</typeStatus>
from “Avenc Canals”); scutum with 5+5 M (M1 and M 
<quantity id="4A2788BFFFC2FF9E2A384CBD0B0EF6FF" box="[1081,1131,1764,1790]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="44" unit="in" value="2.0">2 in</quantity>
more posterior position, M1 even shorter than half of those remaining); with ordinary setae but less abundant than in the pronotum. Metanotum (
<figureCitation id="15E439DFFFC2FF9E2D244D6D0C15F74F" box="[805,880,1844,1870]" captionStart="FIGURE 4" captionStartId="7.[264,367,1542,1566]" captionTargetBox="[450,1157,474,1526]" captionTargetId="figure@7.[396,1190,464,1533]" captionTargetPageId="7" captionText="FIGURE 4. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. Vertex, young specimen from Avenc Mas Nou; b. Vertex, holotype; c. Pronotum, holotype; d. Mesonotum, holotype; e. Metanotum, holotype. Terms: A, S, V, M, I, P groups of large setae in vertex; M, macrochaeta thoracic. (scales: 1 mm)." httpUri="https://zenodo.org/record/174947/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
e): Prescutum with 1+1 M or 0+1 M (male from “Avenc Mas de la Cova” lacking M); scutum with 4+4 M (M2 absent; M1 shorter than those remaining); ordinary setae very short and abundant.
</paragraph>
<paragraph id="8D60255AFFC1FF9D2F394B350E89F29F" blockId="10.[264,1323,284,1036]" pageId="10" pageNumber="45">
Exceptionally long legs. The end of metathoracic pretarsus exceeds half of urite X. Length (units in mm) of the metathoracic leg of the 
<typeStatus id="52649BF8FFC1FF9D2D6D4BCD0CB7F1AF" box="[876,978,404,430]" pageId="10" pageNumber="45" type="holotype">holotype</typeStatus>
(total length 
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): coxa (0.95), trochanter (0.60), femur (2.85), tibia (3.06), tarsus (1.92) and pretarsus (0.41). The tibia is the longest segment in the legs. In all specimens examined legs length is either equal or inferior to half the total length of the body (
<tableCitation id="C05D10E1FFC1FF9D2D7C48550CB3F227" box="[893,982,524,550]" captionStart="TABLE 1" captionStartId="8.[264,352,963,987]" captionTargetBox="[272,1305,1097,1283]" captionTargetPageId="8" captionText="TABLE 1. Length (in mm) of the body, appendixes and number of antennomeres found in the holotype and paratypes of Gollumjapyx smeagol Sendra &amp; Ortuño sp. nov." httpUri="http://table.plazi.org/id/D9A075D2FFC3FF9F2F09499A0B42F3FF" pageId="10" pageNumber="45" tableUuid="D9A075D2FFC3FF9F2F09499A0B42F3FF">Table 1</tableCitation>
). Femur with short ordinary setae; femur and tibia with dozens of large setae; tarsus with 16–18 spiniform setae scattered in two ventral rows; well developed claws, posterior 1.2 times longer than anterior (
<typeStatus id="52649BF8FFC1FF9D2F7348DD0E85F29F" box="[370,480,644,670]" pageId="10" pageNumber="45" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8D60255AFFC1FF9D2F3948F50C37F36D" blockId="10.[264,1323,284,1036]" pageId="10" pageNumber="45">
Abdomen. Tergite I. Prescutum and scutum: 0+0 M. Tergite II: 1+1 M (ma=M). Tergites III–VII (
<figureCitation id="15E439DFFFC1FF9D2FD4488D0D7BF2EF" box="[469,542,724,750]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="10" pageNumber="45">Fig. 5</figureCitation>
a): 5+5 M and 1+ 
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3 (ma=M, M1 absent). Tergite VIII (
<figureCitation id="15E439DFFFC1FF9D2ACD488D0A76F2EF" box="[1228,1299,724,750]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="10" pageNumber="45">Fig. 5</figureCitation>
a) nearly 2.30 times wider than long, 5+5 M (2+2 anterolateral, 2+2 posterolateral and 1+1 consistent with M5). Tergite IX (
<figureCitation id="15E439DFFFC1FF9D2C87497E0DA9F340" box="[646,716,807,833]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="10" pageNumber="45">Fig. 5</figureCitation>
a) around 3.30 times wider than long, M absent but with 4+4 M in the 
<typeStatus id="52649BF8FFC1FF9D2FE2490B0D2DF36D" box="[483,584,850,876]" pageId="10" pageNumber="45" type="paratype">paratype</typeStatus>
from “Avenc Canals”.
</paragraph>
<paragraph id="8D60255AFFC1FF9D2F3949230C01F40D" blockId="10.[264,1323,284,1036]" pageId="10" pageNumber="45">
Urite X (
<figureCitation id="15E439DFFFC1FF9D2FA349230E8DF395" box="[418,488,890,916]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="10" pageNumber="45">Fig. 3</figureCitation>
c and 5b) nearly 1.60 longer than wide, distinctly marked carinae with subparallel margins, 4+4 M intracarinal (D) (3+3 M lateral and 1+1 M posterior) in both small and large specimens, although larger specimens show 3+3 large setae on both sides preceding posterior macrosetae; 5+5 M carinae (L).
</paragraph>
<caption id="D9A075D2FFC1FF9D2F094D710B83F763" httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="10" pageNumber="45" targetBox="[416,1165,1091,1777]" targetPageId="10">
<paragraph id="8D60255AFFC1FF9D2F094D710B83F763" blockId="10.[264,1322,1832,1890]" pageId="10" pageNumber="45">
<emphasis id="BFABF948FFC1FF9D2F094D710EECF741" bold="true" box="[264,393,1832,1856]" pageId="10" pageNumber="45">FIGURE 5.</emphasis>
<taxonomicName id="4ADF5ED9FFC1FF9D2F914D710C50F73E" authority="Sendra &amp; Ortuno" authorityName="Sendra &amp; Ortuno" box="[400,821,1832,1855]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="smeagol" status="sp. nov.">
<emphasis id="BFABF948FFC1FF9D2F914D710D1CF73E" box="[400,633,1832,1855]" italics="true" pageId="10" pageNumber="45">Gollumjapyx smeagol</emphasis>
Sendra &amp; Ortuño
</taxonomicName>
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<taxonomicNameLabel id="A4984433FFC1FF9D2D3D4D710C12F741" box="[828,887,1832,1856]" pageId="10" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm).
</paragraph>
</caption>
<paragraph id="8D60255AFFC0FF9C2F394B450D0CF15F" blockId="11.[264,1323,284,950]" pageId="11" pageNumber="46">Very short ordinary setae, practically absent from the first tergites and progressively more abundant from tergite V.</paragraph>
<paragraph id="8D60255AFFC0FF9C2F394B350B00F187" blockId="11.[264,1323,284,950]" box="[312,1125,364,390]" pageId="11" pageNumber="46">
Acropygium triangular, slightly protruding, rounded (
<figureCitation id="15E439DFFFC0FF9C2DAD4B350C94F187" box="[940,1009,364,390]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="11" pageNumber="46">Fig. 3</figureCitation>
c and 5b).
</paragraph>
<paragraph id="8D60255AFFC0FF9C2F394BCD0E97F277" blockId="11.[264,1323,284,950]" pageId="11" pageNumber="46">
Tergites I–IV showing posterolateral angles either blunt or slightly rounded; angles found in tergite V are more conspicuous due to a small lobiform projection (
<figureCitation id="15E439DFFFC0FF9C2AC04BE50A69F1D7" box="[1217,1292,444,470]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="11" pageNumber="46">Fig. 5</figureCitation>
a); tergites VI and VII show this projection more manifestly, although they are not visibly sclerotized (
<figureCitation id="15E439DFFFC0FF9C2F9D48550E82F227" box="[412,487,524,550]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="11" pageNumber="46">Fig. 5</figureCitation>
a); tergite VIII maintains this angular lobe but shorter than the two preceding ones; finally, the posterior angle of tergite IX lacks the aforementioned projection (
<figureCitation id="15E439DFFFC0FF9C2F8E48050EB1F277" box="[399,468,604,630]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="11" pageNumber="46">Fig. 5</figureCitation>
a).
</paragraph>
<paragraph id="8D60255AFFC0FF9C2F3948DD0B40F29F" blockId="11.[264,1323,284,950]" box="[312,1061,644,670]" pageId="11" pageNumber="46">
Relative lengths of segments VI–X (
<figureCitation id="15E439DFFFC0FF9C2CE248DD0C4EF29F" box="[739,811,644,670]" captionStart="FIGURE 5" captionStartId="10.[264,367,1832,1856]" captionTargetBox="[416,1165,1091,1777]" captionTargetId="figure@10.[396,1190,1060,1793]" captionTargetPageId="10" captionText="FIGURE 5. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., holotype. a. Tergites V to IX; b. Urite X and cerci, dorsal view. Term: M, macrochaeta; m, microchaeta; ls, large setae. (scale: 1 mm)." httpUri="https://zenodo.org/record/174948/files/figure.png" pageId="11" pageNumber="46">Fig. 5</figureCitation>
): 28–28–33–20–100.
</paragraph>
<paragraph id="8D60255AFFC0FF9C2F3948F50C96F3B7" blockId="11.[264,1323,284,950]" pageId="11" pageNumber="46">
Sternite I (
<figureCitation id="15E439DFFFC0FF9C2FBA48F50D63F2C7" box="[443,518,684,710]" captionStart="FIGURE 6" captionStartId="11.[264,367,1736,1760]" captionTargetBox="[449,1144,991,1655]" captionTargetId="figure@11.[396,1190,964,1682]" captionTargetPageId="11" captionText="FIGURE 6. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., paratype female from Avenc Canals pit. a. urosternite I; b. Details of sensory and glandular setae of lateral subcoxal organ. Terms: st, styli; GS, glandular setae; SS, sensory setae. (scales, a: 0.5 mm; b: 0.05 mm)." httpUri="https://zenodo.org/record/174949/files/figure.png" pageId="11" pageNumber="46">Fig. 6</figureCitation>
a): Prescutum 5+5 M; scutum: 22+22 M and other 7–9+7–9 large setae consistent with true macrosetae; sternite totally covered with short ordinary setae, increasing in posteriorly and with longer and more robust setae scattered in two or three rows at the front of the lateral subcoxal organs (between 
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the male specimen from “Avenc Mas de la Cova” and 
<quantity id="4A2788BFFFC0FF9C2C6949150DC0F367" box="[616,677,844,870]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="46" unit="in" value="60.0">60 in</quantity>
the female specimen from “Avenc Canals”). Subcoxal organs appear either invaginated below the posterior margin of sternite I or evaginated outwards, protruding over the margin of the urosternite (
<figureCitation id="15E439DFFFC0FF9C2D9C49C50C80F3B7" box="[925,997,924,950]" captionStart="FIGURE 6" captionStartId="11.[264,367,1736,1760]" captionTargetBox="[449,1144,991,1655]" captionTargetId="figure@11.[396,1190,964,1682]" captionTargetPageId="11" captionText="FIGURE 6. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., paratype female from Avenc Canals pit. a. urosternite I; b. Details of sensory and glandular setae of lateral subcoxal organ. Terms: st, styli; GS, glandular setae; SS, sensory setae. (scales, a: 0.5 mm; b: 0.05 mm)." httpUri="https://zenodo.org/record/174949/files/figure.png" pageId="11" pageNumber="46">Fig. 6</figureCitation>
).
</paragraph>
<caption id="D9A075D2FFC0FF9C2F094C910B60F721" httpUri="https://zenodo.org/record/174949/files/figure.png" pageId="11" pageNumber="46" targetBox="[449,1144,991,1655]" targetPageId="11">
<paragraph id="8D60255AFFC0FF9C2F094C910B60F721" blockId="11.[264,1323,1736,1824]" pageId="11" pageNumber="46">
<emphasis id="BFABF948FFC0FF9C2F094C910EEFF6E1" bold="true" box="[264,394,1736,1760]" pageId="11" pageNumber="46">FIGURE 6.</emphasis>
<taxonomicName id="4ADF5ED9FFC0FF9C2F914C900C50F6E1" authority="Sendra &amp; Ortuno" authorityName="Sendra &amp; Ortuno" box="[400,821,1737,1760]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="smeagol" status="sp. nov.">
<emphasis id="BFABF948FFC0FF9C2F914C900D1CF6E1" box="[400,633,1737,1760]" italics="true" pageId="11" pageNumber="46">Gollumjapyx smeagol</emphasis>
Sendra &amp; Ortuño
</taxonomicName>
<emphasis id="BFABF948FFC0FF9C2D3D4C910C12F6E1" bold="true" box="[828,887,1736,1760]" pageId="11" pageNumber="46">
<taxonomicNameLabel id="A4984433FFC0FF9C2D3D4C910C12F6E1" box="[828,887,1736,1760]" pageId="11" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype female from Avenc Canals pit. a. urosternite I; b. Details of sensory and glandular setae of lateral subcoxal organ. Terms: st, styli; GS, glandular setae; SS, sensory setae. (scales, a: 0.5 mm; b: 0.05 mm).
</paragraph>
</caption>
<paragraph id="8D60255AFFC7FF9B2F394B450E3AF227" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">
The rows of setae on the subcoxal organs (
<figureCitation id="15E439DFFFC7FF9B2D464B450CEFF137" box="[839,906,284,310]" captionStart="FIGURE 6" captionStartId="11.[264,367,1736,1760]" captionTargetBox="[449,1144,991,1655]" captionTargetId="figure@11.[396,1190,964,1682]" captionTargetPageId="11" captionText="FIGURE 6. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., paratype female from Avenc Canals pit. a. urosternite I; b. Details of sensory and glandular setae of lateral subcoxal organ. Terms: st, styli; GS, glandular setae; SS, sensory setae. (scales, a: 0.5 mm; b: 0.05 mm)." httpUri="https://zenodo.org/record/174949/files/figure.png" pageId="12" pageNumber="47">Fig 6</figureCitation>
b) represent between 1/5 to 1/3 of interstyle width (
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females and 
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males and 
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a specimen of unknown sex from “Avenc d’en Serenge”). About 65 short glandular setae in the male from “Avenc del Mas de la Cova”, set in 1 or 2 rows, GS/st1= 0.14; nearly 80 glandular setae, equally short, in the female from “Avenc Canals”, in 2 rows, GS/st1= 0.13; 25–28 sensory setae, SS/st1= 
<quantity id="4A2788BFFFC7FF9B2C254BBD0D1FF1FF" box="[548,634,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="47" unit="in" value="0.1">0.10 in</quantity>
the male and 35–38 sensory setae, SS/st1= 
<quantity id="4A2788BFFFC7FF9B2AA54BBD0B9FF1FF" box="[1188,1274,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="47" unit="in" value="0.1">0.10 in</quantity>
the female.
</paragraph>
<paragraph id="8D60255AFFC7FF9B2F39486D0BC3F277" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">
Both visible median glandular organ and central setae-shaped sensilla missing. These sensilla appear in 4–8+ 
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the internal side of the subcoxal organs (
<figureCitation id="15E439DFFFC7FF9B2A4348050BECF277" box="[1090,1161,604,630]" captionStart="FIGURE 6" captionStartId="11.[264,367,1736,1760]" captionTargetBox="[449,1144,991,1655]" captionTargetId="figure@11.[396,1190,964,1682]" captionTargetPageId="11" captionText="FIGURE 6. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., paratype female from Avenc Canals pit. a. urosternite I; b. Details of sensory and glandular setae of lateral subcoxal organ. Terms: st, styli; GS, glandular setae; SS, sensory setae. (scales, a: 0.5 mm; b: 0.05 mm)." httpUri="https://zenodo.org/record/174949/files/figure.png" pageId="12" pageNumber="47">Fig. 6</figureCitation>
a).
</paragraph>
<paragraph id="8D60255AFFC7FF9B2F3948DD0DFBF2EF" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Urosternites II–VII: 28+28 M, although missing some typical macrosetae (C2, C8, C10). Urosternite VIII: 12+12 M, in the largest specimens and 7+7 M in the smallest. Paratergite VIII: 1 M subposterior.</paragraph>
<paragraph id="8D60255AFFC7FF9B2F3948A50D3CF340" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Elongate styli with sharp sclerotized points: s1/s7= 0.44–0.94; st1/st7= 0.59–0.76; s1/st7= 0.24–0.46; s1/st7= 0.26–0.59.</paragraph>
<paragraph id="8D60255AFFC7FF9B2F39490B0CE0F3BD" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Male genital papillae could not be examined in the largest specimen from “Avenc Mas de la Cova”; however, genital appendices are wholly membranous and cylindrical-conic. No ventral glandular fossae have been found in males.</paragraph>
<paragraph id="8D60255AFFC7FF9B2F3949930BE5F4FD" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">
Cerci. (
<figureCitation id="15E439DFFFC7FF9B2F9549930E85F3E5" box="[404,480,970,996]" captionStart="FIGURE 3" captionStartId="6.[264,367,1398,1422]" captionTargetBox="[269,1317,326,1373]" captionTargetId="figure@6.[269,1317,326,1374]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron microscope photography of Gollumjapyx smeagol Sendra &amp; Ortuño sp. n.. a. apical antennomere; b. XXXV antennomere; c. urite X and cerci, dorsal view; d. urite X and cerci, ventral view. Terms: p, placoid sensilla; ss, short sensilla. (scales, a and b: 25 µm; c and d: 500 µm)." httpUri="https://zenodo.org/record/174946/files/figure.png" pageId="12" pageNumber="47">Fig. 3</figureCitation>
c, d and 5b). Large, at first slightly curved, becoming more curved towards the extreme, pointed. Length ranges from 
<quantity id="4A2788BFFFC7FF9B2D5D49AB0CA0F40D" box="[860,965,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="12" pageNumber="47" unit="mm" value="3.25">3.25 mm</quantity>
in 
<typeStatus id="52649BF8FFC7FF9B2DEA49AB0B37F40D" box="[1003,1106,1010,1036]" pageId="12" pageNumber="47" type="holotype">holotype</typeStatus>
to 
<quantity id="4A2788BFFFC7FF9B2A7649AB0B84F40D" box="[1143,1249,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="12" pageNumber="47" unit="mm" value="1.17">1.17 mm</quantity>
in the smallest specimen; the relative size of the cercus increases larger in larger specimens (
<tableCitation id="C05D10E1FFC7FF9B2F114E1B0E0FF45D" box="[272,362,1090,1116]" captionStart="TABLE 1" captionStartId="8.[264,352,963,987]" captionTargetBox="[272,1305,1097,1283]" captionTargetPageId="8" captionText="TABLE 1. Length (in mm) of the body, appendixes and number of antennomeres found in the holotype and paratypes of Gollumjapyx smeagol Sendra &amp; Ortuño sp. nov." httpUri="http://table.plazi.org/id/D9A075D2FFC3FF9F2F09499A0B42F3FF" pageId="12" pageNumber="47" tableUuid="D9A075D2FFC3FF9F2F09499A0B42F3FF">Table 1</tableCitation>
). In all specimens the length of the right cercus is slightly inferior to that of the left (0.95 to 0.97). Cerci slightly longer than the normally uncovered part of urite X in smaller specimens, but shorter in the larger specimens. Both cerci heavily sclerotized, in particular both external and internal margins and also in the dorsal carinae projecting from the acetabula, running the length of the right cercus and halfway on the left.
</paragraph>
<paragraph id="8D60255AFFC7FF9B2F394F530D21F5EE" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Right cercus. Triangular tooth, very strong, distinctly proximal, r d= 0.40–0.65 (ΨΨ), 0.57–0.58 (ɗɗ); width of cercus at tooth level equals that of base; predental margin with 2 to 4 strong tubercles and very prominent in larger specimens, wherein 1 or 2 appear to be bitubercles; in smaller specimens tubercles are simple and smaller. Postdental arista is slightly concave, surrounded by tiny rounded denticles, tapering in size extremely even to the point of disappearance.</paragraph>
<paragraph id="8D60255AFFC7FF9B2F394FA40BE1F693" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Left cercus. Sharp postmedian tooth, r g= 1.14–1.60 (Ψ), 1.33–1.51 (ɗ); width of cercus at tooth level slightly larger or equal to half the base of the appendix; concave predental arista with 8–12/11–15 sharp/rounded tubercles; postdental arista slightly concave with very small denticles or showing a smooth surface in the smallest specimens.</paragraph>
<paragraph id="8D60255AFFC7FF9B2F094CF90EADF70B" blockId="12.[264,1323,284,1882]" pageId="12" pageNumber="47">Chaetotaxy: in dorsal view, the inside margin of both cerci has anterolateral M and a large setae, at tooth level, in the right cercus. The outside margins of both cerci support a dozen large setae each.</paragraph>
</subSubSection>
<subSubSection id="C5C576D1FFC7FF9A2F394D4E0DFCF317" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47" type="discussion">
<paragraph id="8D60255AFFC7FF9A2F394D4E0A4EF1AF" blockId="12.[264,1323,284,1882]" lastBlockId="13.[264,1323,284,1870]" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47">
<emphasis id="BFABF948FFC7FF9B2F394D4E0EDAF730" bold="true" box="[312,447,1815,1841]" pageId="12" pageNumber="47">Comments</emphasis>
. During this study, our colleague at the University of Murcia, José Luis Lencina, gave us a large japygid specimen (body length 
<quantity id="4A2788BFFFC7FF9B2DDC4D190B35F75B" box="[989,1104,1856,1882]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="12" pageNumber="47" unit="mm" value="22.6">22.6 mm</quantity>
, including cerci), displaying troglobiomorphic features. Both the cercal armature and the lengthening of thorax and legs in this specimen show a strong resemblance to 
<taxonomicName id="4ADF5ED9FFC6FF9A2DFD4B1D0B9BF15C" box="[1020,1278,324,349]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC6FF9A2DFD4B1D0B9BF15C" box="[1020,1278,324,349]" italics="true" pageId="13" pageNumber="48">Gollumjapyx smeagol</emphasis>
</taxonomicName>
; its poor state of conservation, however, made it impossible to conduct a thorough study that could have confirmed or denied its inclusion in the species described in the present article.
</paragraph>
<paragraph id="8D60255AFFC6FF9A2F394BE50DFCF317" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="48">
Here follow the collection data: 1 Ψ, Comunidad de Murcia, Murcia, Jumilla, “Cueva del Pozo”; 
<date id="F961039AFFC6FF9A2F984BBD0D47F1FF" box="[409,546,484,510]" pageId="13" pageNumber="48" value="1992-12-08">8-XII-1992</date>
; Grupo espeleológico Hinneni leg. (deposited at the Valencia Natural History Museum, 
<collectingCountry id="F5C865CAFFC6FF9A2C3B48550DE7F227" box="[570,642,524,550]" name="Spain" pageId="13" pageNumber="48">Spain</collectingCountry>
). Pozo cave (Jumilla, Murcia, 
<collectingCountry id="F5C865CAFFC6FF9A2DE248550B42F227" box="[995,1063,524,550]" name="Spain" pageId="13" pageNumber="48">Spain</collectingCountry>
) is a fossil cave, more than 
<quantity id="4A2788BFFFC6FF9A2F43486D0E18F24F" box="[322,381,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="48" unit="km" value="1.0">1 km</quantity>
in length and some 50 metres deep, located at an altitude of 498 metres, open in limestone and dolomites from the upper cretaceous at the Rajica de En Medio range, in the southern Prebetic mountain range. The specimen was collected 200 metres into the entrance, on damp clayey soil (despite low humidity), at a mean temperature of 26ºC, higher than expected according to the annual average for this locality (unpublished data, forwarded by the Hinneni Group).
</paragraph>
</subSubSection>
<subSubSection id="C5C576D1FFC6FF992F39497D0C9EF6AF" lastPageId="14" lastPageNumber="49" pageId="13" pageNumber="48" type="distribution">
<paragraph id="8D60255AFFC6FF9A2F39497D0D64F457" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="48">
<emphasis id="BFABF948FFC6FF9A2F39497D0DBAF33F" bold="true" box="[312,735,804,830]" pageId="13" pageNumber="48">Distribution, habitat and biology.</emphasis>
To date 
<taxonomicName id="4ADF5ED9FFC6FF9A2D4A497D0B2BF33C" box="[843,1102,804,829]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="smeagol" status="gen. nov.">
<emphasis id="BFABF948FFC6FF9A2D4A497D0B2BF33C" box="[843,1102,804,829]" italics="true" pageId="13" pageNumber="48">Gollumjapyx smeagol</emphasis>
</taxonomicName>
<emphasis id="BFABF948FFC6FF9A2A56497D0B9FF33F" bold="true" box="[1111,1274,804,830]" pageId="13" pageNumber="48">
<taxonomicNameLabel id="A4984433FFC6FF9A2A56497D0BC9F33F" box="[1111,1196,804,830]" pageId="13" pageNumber="48" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
has been found in six caves scattered along the limestone relief of “The Coastal Ranges”. This geomorphological region defined by 
<bibRefCitation id="E94E58ABFFC6FF9A2CD1492D0C13F38F" author="Garay" box="[720,886,884,910]" pageId="13" pageNumber="51" refString="Garay, P. (1995) Marco geologico estructural y geotectonica. In: Universitat de Valencia &amp; Asociacion Espanola para Estudio del Cuaternario (Ed.). El Cuaternario del Pais Valenciano. Valencia, 31 - 42." type="book chapter" year="1995">Garay (1995)</bibRefCitation>
, comprises the low altitude coastal ranges (400–600 metres average height) from the Oropesa range (Castellón, 
<collectingCountry id="F5C865CAFFC6FF9A2A9349C50BB2F3B7" box="[1170,1239,924,950]" name="Spain" pageId="13" pageNumber="48">Spain</collectingCountry>
) to the Montsià range (Tarragona, 
<collectingCountry id="F5C865CAFFC6FF9A2C4A499D0DF6F3DF" box="[587,659,964,990]" name="Spain" pageId="13" pageNumber="48">Spain</collectingCountry>
). It displays a series of particular palaeogeographic and stratigraphic traits which have seemingly allowed it to remain a refugium area throughout the Cenozoic era for a host of species, presently recognised to be relics or paleo-endemics (
<bibRefCitation id="E94E58ABFFC6FF9A2F114E650E90F457" author="Ortuno" box="[272,501,1084,1110]" pageId="13" pageNumber="52" refString="Ortuno, V. M.; Sendra, A.; Montagud, S. &amp; Teruel, S. (2005) Systematique et biologie d'une espece paleoendemique hypogee de la peninsule Iberique: Ildobates neboti Espanol, 1966 (Coleoptera: Carabidae: Dryptinae). Annales de la Societe Entomologique de France (Nouvelle Serie), 40, 459 - 475." type="journal article" year="2005">
Ortuño 
<emphasis id="BFABF948FFC6FF9A2F6D4E650EC5F454" box="[364,416,1084,1109]" italics="true" pageId="13" pageNumber="48">et al</emphasis>
., 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8D60255AFFC6FF9A2F394E3D0EF7F4F7" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="48">
The northernmost locality is the “Avenc Canals” in the Montsià range, a small cave with stalagmite formations. The specimen was retrieved at the base of the entry shaft, in a formation 
<quantity id="4A2788BFFFC6FF9A2F834EED0E92F4CF" box="[386,503,1204,1230]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="13" pageNumber="48" unit="cm" value="30.0" valueMax="40.0" valueMin="20.0">20–40 cm</quantity>
from a debris base made up of dirt and vegetable detritus (F. Fadrique com. pers.).
</paragraph>
<paragraph id="8D60255AFFC6FF9A2F394F5D0C2EF65F" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="48">
In the “la Vall d’Àngel” range, towards the south, there is “Cova dels Encenalls” cave; it hosts a considerable catalogue of troglobite elements, including 
<taxonomicName id="4ADF5ED9FFC6FF9A2A5E4F750BA1F547" box="[1119,1220,1324,1350]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="tribe" tribe="Zuphiini">Zuphiini</taxonomicName>
carabid 
<taxonomicName id="4ADF5ED9FFC6FF9A2F094F0D0DECF56F" authority="Espanol, 1965" authorityName="Espanol" authorityYear="1965" box="[264,649,1364,1390]" class="Insecta" family="Carabidae" genus="Ildobates" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="neboti">
<emphasis id="BFABF948FFC6FF9A2F094F0D0EA9F56C" box="[264,460,1364,1389]" italics="true" pageId="13" pageNumber="48">Ildobates neboti</emphasis>
Español, 1965
</taxonomicName>
and Anillini carabids 
<taxonomicName id="4ADF5ED9FFC6FF9A2DA94F0D0E28F597" authority="Espanol, 1971" authorityName="Espanol" authorityYear="1971" class="Insecta" family="Carabidae" genus="Speleotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="jusmeti">
<emphasis id="BFABF948FFC6FF9A2DA94F0D0BC8F56C" box="[936,1197,1364,1389]" italics="true" pageId="13" pageNumber="48">Speleotyphlus jusmeti</emphasis>
(Español, 1971)
</taxonomicName>
and 
<taxonomicName id="4ADF5ED9FFC6FF9A2F9B4F250C20F597" authority="Espanol, 1971" authorityName="Espanol" authorityYear="1971" box="[410,837,1404,1430]" class="Insecta" family="Carabidae" genus="Iberanillus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="vinyasi">
<emphasis id="BFABF948FFC6FF9A2F9B4F250D1AF594" box="[410,639,1404,1429]" italics="true" pageId="13" pageNumber="48">Iberanillus vinyasi</emphasis>
Español, 1971
</taxonomicName>
; 
<taxonomicName id="4ADF5ED9FFC6FF9A2D5D4F250C84F597" box="[860,993,1404,1430]" class="Arachnida" family="Dysderidae" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="48" phylum="Arthropoda" rank="family">Dysderidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC6FF9A2DF24F250B30F597" box="[1011,1109,1404,1430]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="48" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC6FF9A2A644F250D46F5BF" authority="Ribera, 1982" authorityName="Ribera" authorityYear="1982" class="Arachnida" family="Dysderidae" genus="Speleoharpactea" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="levantina">
<emphasis id="BFABF948FFC6FF9A2A644F250E10F5BC" italics="true" pageId="13" pageNumber="48">Speleoharpactea levantina</emphasis>
Ribera, 1982
</taxonomicName>
; Neobisidae 
<taxonomicName id="4ADF5ED9FFC6FF9A2CC64FFD0B3FF5BC" authority="Acantocreagris" authorityName="Acantocreagris" box="[711,1114,1444,1470]" class="Arachnida" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="48" phylum="Arthropoda" rank="order">
Pseudoscorpiones 
<emphasis id="BFABF948FFC6FF9A2DA74FFD0B3FF5BC" box="[934,1114,1444,1469]" italics="true" pageId="13" pageNumber="48">Acantocreagris</emphasis>
</taxonomicName>
relicta Mahnert, 1977; and 
<taxonomicName id="4ADF5ED9FFC6FF9A2F844F950D43F5E7" box="[389,550,1484,1510]" class="Entognatha" family="Campodeidae" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="family">Campodeidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC6FF9A2C2F4F950DEDF5E7" box="[558,648,1484,1510]" class="Entognatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="order">Diplura</taxonomicName>
<taxonomicName id="4ADF5ED9FFC6FF9A2C904F950B8FF5E7" authority="Bareth &amp; Conde, 1981" authorityName="Bareth &amp; Conde" authorityYear="1981" box="[657,1258,1484,1510]" class="Entognatha" family="Campodeidae" genus="Paratachycampa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="BFABF948FFC6FF9A2C904F950CB6F5E4" box="[657,979,1484,1509]" italics="true" pageId="13" pageNumber="48">Paratachycampa hispanica</emphasis>
Bareth &amp; Condé, 1981
</taxonomicName>
. This cave is, for the greater part of its 
<quantity id="4A2788BFFFC6FF9A2C974FAD0DB6F60F" box="[662,723,1524,1550]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="48" unit="m" value="75.0">75 m</quantity>
width and only 
<quantity id="4A2788BFFFC6FF9A2D964FAD0CB6F60F" box="[919,979,1524,1550]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="48" unit="m" value="10.0">10 m</quantity>
depth, covered in stalagmite floors, with small gours, generally flooded. The specimen was found wandering on the stony soil of a side gallery (J. Comas com. pers.).
</paragraph>
<paragraph id="8D60255AFFC6FF992F394C350EB4F227" blockId="13.[264,1323,284,1870]" lastBlockId="14.[264,1323,284,1710]" lastPageId="14" lastPageNumber="49" pageId="13" pageNumber="48">
“Avenc del Mas Nou” pit lies on the southern foothills of the “Serra d’en Galceran” range; access to its 
<quantity id="4A2788BFFFC6FF9A2FEA4CCD0D43F6AF" box="[491,550,1684,1710]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="48" unit="m" value="28.0">28 m</quantity>
drop is gained through a narrow gap. The cave extends for 
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and reaches a depth of 
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<quantity id="4A2788BFFFC6FF9A2C2A4CE50D0BF6D7" box="[555,622,1724,1750]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="48" unit="m" value="40.0">40 m</quantity>
.
</specimenCount>
It is covered with stalagmite formations as well as clay deposits, all in an extremely humid environment. The specimen was collected crawling on the damp formations at over 
<quantity id="4A2788BFFFC6FF9A2C774D550DD2F727" box="[630,695,1804,1830]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="48" unit="m" value="30.0">30 m</quantity>
depth. Also spotted in that same area were many specimens of two detrivorous species of 
<taxonomicName id="4ADF5ED9FFC6FF9A2D374D6D0CB2F74F" box="[822,983,1844,1870]" class="Entognatha" family="Campodeidae" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="family">Campodeidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC6FF9A2DED4D6D0B2FF74F" box="[1004,1098,1844,1870]" class="Entognatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="order">Diplura</taxonomicName>
: 
<taxonomicName id="4ADF5ED9FFC6FF992A634D6D0E1FF134" class="Entognatha" family="Campodeidae" genus="Paratachycampa" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="14" lastPageNumber="49" order="Diplura" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="BFABF948FFC6FF992A634D6D0E1FF134" italics="true" lastPageId="14" lastPageNumber="49" pageId="13" pageNumber="48">Paratachycampa hispanica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4ADF5ED9FFC5FF992FB54B450B7CF137" authority="Sendra &amp; Escola, 2004" authorityName="Sendra &amp; Escola" authorityYear="2004" box="[436,1049,284,310]" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="maestrazgoensis">
<emphasis id="BFABF948FFC5FF992FB54B450D9BF134" box="[436,766,284,309]" italics="true" pageId="14" pageNumber="49">Campodea maestrazgoensis</emphasis>
Sendra &amp; Escolà, 2004
</taxonomicName>
. In this same mountain range there is a small cave, less than 
<quantity id="4A2788BFFFC5FF992CCB4B1D0C6DF15F" box="[714,776,324,350]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="49" unit="m" value="70.0">70 m</quantity>
of practically horizontal extension, known as “Cova del Mas de Gaspar” or “Cova Santa”, where the first specimen of this new taxon was discovered — which has been used as 
<typeStatus id="52649BF8FFC5FF992D334BCD0CFCF1AF" box="[818,921,404,430]" pageId="14" pageNumber="49" type="holotype">holotype</typeStatus>
in the description. 
<taxonomicName id="4ADF5ED9FFC5FF992A924BCD0E0CF1D4" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC5FF992A924BCD0E0CF1D4" italics="true" pageId="14" pageNumber="49">Gollumjapyx smeagol</emphasis>
</taxonomicName>
shares this cave with other troglobite elements such as 
<taxonomicName id="4ADF5ED9FFC5FF992A294BE50BF8F1D7" box="[1064,1181,444,470]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992AAB4BE50A4EF1D7" box="[1194,1323,444,470]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992F094BBD0DD5F1FF" authority="Lagar, 1978" authorityName="Lagar" authorityYear="1978" box="[264,688,484,510]" class="Insecta" family="Leiodidae" genus="Anillochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="cullelli">
<emphasis id="BFABF948FFC5FF992F094BBD0D6BF1FC" box="[264,526,484,509]" italics="true" pageId="14" pageNumber="49">Anillochlamys cullelli</emphasis>
Lagar, 1978
</taxonomicName>
and 
<taxonomicName id="4ADF5ED9FFC5FF992CF44BBD0CF3F1FF" box="[757,918,484,510]" class="Entognatha" family="Campodeidae" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="family">Campodeidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992DA54BBD0C9BF1FF" box="[932,1022,484,510]" class="Entognatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="order">Diplura</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992A0A4BBD0EAFF224" class="Entognatha" family="Campodeidae" genus="Campodea" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="maestrazgoensis" subGenus="Campodea">
<emphasis id="BFABF948FFC5FF992A0A4BBD0EAFF224" italics="true" pageId="14" pageNumber="49">Campodea (Campodea) maestrazgoensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8D60255AFFC5FF992F39486D0DADF47F" blockId="14.[264,1323,284,1710]" pageId="14" pageNumber="49">
The two southernmost localities are two pits in close proximity, the “Avenc d’En Serenge” and the “Avenc Mas de la Cova”. Both are located in limestone elevations near the town of Cabanes, in Ferradura mountain. The “Avenc d’en Serenge”, as well as being the typical locality of 
<taxonomicName id="4ADF5ED9FFC5FF992C0D48F50DAEF2C4" box="[524,715,684,709]" class="Insecta" family="Carabidae" genus="Ildobates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="neboti">
<emphasis id="BFABF948FFC5FF992C0D48F50DAEF2C4" box="[524,715,684,709]" italics="true" pageId="14" pageNumber="49">Ildobates neboti</emphasis>
</taxonomicName>
, is home to a great variety of troglobite forms with great interest from a palaeo- and biogeographical point of view: Neobisidae 
<taxonomicName id="4ADF5ED9FFC5FF992F0948A50EBFF317" box="[264,474,764,790]" class="Arachnida" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="49" phylum="Arthropoda" rank="order">Pseudoscorpiones</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992FEF48A50B66F317" authority="Ellingsen, 1912" authorityName="Ellingsen" authorityYear="1912" box="[494,1027,764,790]" class="Arachnida" family="Bochicidae" genus="Troglobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="BFABF948FFC5FF992FEF48A50C76F314" box="[494,787,764,789]" italics="true" pageId="14" pageNumber="49">Troglobisium racovitzai</emphasis>
(Ellingsen, 1912)
</taxonomicName>
; 
<taxonomicName id="4ADF5ED9FFC5FF992A1D48A50BD8F317" box="[1052,1213,764,790]" class="Entognatha" family="Campodeidae" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="family">Campodeidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992AD048A50A4EF317" box="[1233,1323,764,790]" class="Entognatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="order">Diplura</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992F09497D0D28F33C" box="[264,589,804,829]" class="Entognatha" family="Campodeidae" genus="Paratachycampa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="BFABF948FFC5FF992F09497D0D28F33C" box="[264,589,804,829]" italics="true" pageId="14" pageNumber="49">Paratachycampa hispanica</emphasis>
</taxonomicName>
; Anillini carabid 
<taxonomicName id="4ADF5ED9FFC5FF992D27497D0B90F33F" authority="Espanol, 1966" authorityName="Espanol" authorityYear="1966" box="[806,1269,804,830]" class="Insecta" family="Carabidae" genus="Speleotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="aurouxi">
<emphasis id="BFABF948FFC5FF992D27497D0B48F33C" box="[806,1069,804,829]" italics="true" pageId="14" pageNumber="49">Speleotyphlus aurouxi</emphasis>
(Español, 1966)
</taxonomicName>
and 
<taxonomicName id="4ADF5ED9FFC5FF992F0949150EF1F367" box="[264,404,844,870]" class="Insecta" family="Pselaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="family">Pselaphidae</taxonomicName>
<taxonomicName id="4ADF5ED9FFC5FF992FAA49150CD4F367" authority="Besuchet, 1974" authorityName="Besuchet" authorityYear="1974" box="[427,945,844,870]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="escolai">
<emphasis id="BFABF948FFC5FF992FAA49150DDBF364" box="[427,702,844,869]" italics="true" pageId="14" pageNumber="49">Tychobythinus escolai</emphasis>
(Besuchet, 1974)
</taxonomicName>
. Five of the specimens of 
<taxonomicName id="4ADF5ED9FFC5FF992F09492D0D6DF38C" box="[264,520,884,909]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC5FF992F09492D0D6DF38C" box="[264,520,884,909]" italics="true" pageId="14" pageNumber="49">Gollumjapyx smeagol</emphasis>
</taxonomicName>
come from this cave which extends for 
<quantity id="4A2788BFFFC5FF992DDF492D0B42F38F" box="[990,1063,884,910]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="49" unit="m" value="110.0">110 m</quantity>
and 
<quantity id="4A2788BFFFC5FF992A61492D0BF9F38F" box="[1120,1180,884,910]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="49" unit="m" value="34.0">34 m</quantity>
depth. Four of the aforementioned specimens were collected in the deepest and dampest areas of the cave, totally covered by calcareous formations, at more than 
<quantity id="4A2788BFFFC5FF992DD1499D0B69F3DF" box="[976,1036,964,990]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="49" unit="m" value="20.0">20 m</quantity>
depth. In the case of the “Avenc del Mas de la Cova” pit no precise data are available about the way the specimen was located. The cave runs for more than 
<quantity id="4A2788BFFFC5FF992D034E4D0C2AF42F" box="[770,847,1044,1070]" metricMagnitude="2" metricUnit="m" metricValue="1.41" pageId="14" pageNumber="49" unit="m" value="141.0">141 m</quantity>
and reaches a depth of 
<specimenCount id="9BD9EED3FFC5FF992A6E4E4D0BD1F42F" box="[1135,1204,1044,1070]" pageId="14" pageNumber="49" type="generic">
<quantity id="4A2788BFFFC5FF992A6E4E4D0BD5F42F" box="[1135,1200,1044,1070]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="14" pageNumber="49" unit="m" value="43.0">43 m</quantity>
.
</specimenCount>
From the base of the first shaft at 
<quantity id="4A2788BFFFC5FF992C2F4E650D09F457" box="[558,620,1084,1110]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="49" unit="m" value="17.0">17 m</quantity>
depth, the cave remains very damp, with an abundance of stalagmitic formations and dripstones.
</paragraph>
<paragraph id="8D60255AFFC5FF992F394ED50E3BF56F" blockId="14.[264,1323,284,1710]" pageId="14" pageNumber="49">
<taxonomicName id="4ADF5ED9FFC5FF992F394ED50D5AF4A4" box="[312,575,1164,1189]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC5FF992F394ED50D5AF4A4" box="[312,575,1164,1189]" italics="true" pageId="14" pageNumber="49">Gollumjapyx smeagol</emphasis>
</taxonomicName>
has been found in mesovoid deep substratum, in caves of average extension and little depth, excavated in limestone from the Lower Cretaceous, in altitudes between 300–500 metres. Ground temperature ranges from 14ºC to 17ºC, always in humid areas. It is therefore likely to be stenoterm and stenohygrobious, although not strictly, which would presumably allow it to survive in mesovoid shallow substratum (MSS).
</paragraph>
<paragraph id="8D60255AFFC5FF992F394F250C9EF6AF" blockId="14.[264,1323,284,1710]" pageId="14" pageNumber="49">
It is well known that the most common feeding habit in 
<taxonomicName id="4ADF5ED9FFC5FF992DEC4F250B07F597" box="[1005,1122,1404,1430]" class="Entognatha" family="Japygidae" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="family">Japygidae</taxonomicName>
is to predate on smaller invertebrates. It has been stated that japygids refuse to feed on dead organisms (
<bibRefCitation id="E94E58ABFFC5FF992F114F950ECDF5E7" author="Pages" box="[272,424,1484,1510]" pageId="14" pageNumber="52" refString="Pages, J. (1967) Donnees sur la Biologie de Dipljapyx humberti (Grassi). Revue d'Ecologie et de Biologie du Sol, 4, 187 - 281." type="journal article" year="1967">Pagés, 1967</bibRefCitation>
). Even though feeding in 
<taxonomicName id="4ADF5ED9FFC5FF992CEA4F950C95F5E4" box="[747,1008,1484,1509]" class="Entognatha" family="Japygidae" genus="Gollumjapyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diplura" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="smeagol">
<emphasis id="BFABF948FFC5FF992CEA4F950C95F5E4" box="[747,1008,1484,1509]" italics="true" pageId="14" pageNumber="49">Gollumjapyx smeagol</emphasis>
</taxonomicName>
has not been observed, a thorough examination of the digestive content has been performed. Two specimens from the “Avenc d’en Serenge” showed fragments of Acari (legs and sclerites) inside the digestive tract. A third specimen from the same cave showed fragments of 
<taxonomicName id="4ADF5ED9FFC5FF992A894C1D0E07F684" class="Insecta" family="Carabidae" genus="Speleotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="aurouxi">
<emphasis id="BFABF948FFC5FF992A894C1D0E07F684" italics="true" pageId="14" pageNumber="49">Speleotyphlus aurouxi</emphasis>
</taxonomicName>
(5 antennomeres, a leg, part of the ring of the genital segment and the whole aedeagus- median lobe and parameres still articulated) (
<figureCitation id="15E439DFFFC5FF992D974CCD0CB9F6AF" box="[918,988,1684,1710]" captionStart="FIGURE 2. a" captionStartId="5.[264,367,1746,1770]" captionTargetBox="[264,1323,313,1734]" captionTargetId="figure@5.[264,1323,313,1734]" captionTargetPageId="5" captionText="FIGURE 2. a. Gollumjapyx smeagol Sendra &amp; Ortuño sp. n., Avenc d’en Serenge cave from Cabanes, Castellón (Spain); b. detail of the contents from the digestive tract belonging to a specimen from Avenc d’en Serenge; all the fragments belong to a male of Speleotyphlus aurouxi (Carabidae, Anillini). Terms: a, antennomeres; rgs, ring of the genital segment; ml, median lobe; p, paramere; t, tarsomeres. (scales, a: 3 mm, b: 0.3 mm)." httpUri="https://zenodo.org/record/174945/files/figure.png" pageId="14" pageNumber="49">Fig. 2</figureCitation>
b).
</paragraph>
</subSubSection>
</treatment>
</document>