treatments-xml/data/05/4F/DB/054FDB70FE99FEBFB49C1D5D7F34231E.xml

<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FE99FEBFB49C1D5D7F34231E" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax rugosus" docType="treatment" docVersion="4" lastPageNumber="358" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="351" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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<mods:date>2021</mods:date>
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<mods:title>e 11514</mods:title>
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<mods:number>2021-06-30</mods:number>
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<mods:number>9</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102626" ID-GBIF-Taxon="183542645" ID-Zenodo-Dep="5102626" LSID="urn:lsid:plazi:treatment:054FDB70FE99FEBFB49C1D5D7F34231E" httpUri="http://treatment.plazi.org/id/054FDB70FE99FEBFB49C1D5D7F34231E" lastPageId="357" lastPageNumber="358" pageId="350" pageNumber="351">
<subSubSection box="[498,957,1856,1883]" pageId="350" pageNumber="351" type="nomenclature">
<paragraph blockId="350.[498,1203,1856,1923]" box="[498,957,1856,1883]" pageId="350" pageNumber="351">
<heading box="[498,957,1856,1883]" level="2" pageId="350" pageNumber="351">
<taxonomicName authority="(Mackay, 2000)" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,957,1856,1883]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="350" pageNumber="351" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis bold="true" box="[498,957,1856,1883]" pageId="350" pageNumber="351">
<emphasis bold="true" box="[498,759,1856,1883]" italics="true" pageId="350" pageNumber="351">Temnothorax rugosus</emphasis>
(
<bibRefCitation author="Mackay WP" box="[779,947,1856,1883]" journalOrPublisher="Sociobiology" pageId="350" pageNumber="351" pagination="265 - 444" part="36" refId="ref271737" refString="Mackay WP. 2000. A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265 - 444." title="A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae)" type="journal article" year="2000">
<emphasis bold="true" box="[779,947,1856,1883]" italics="true" pageId="350" pageNumber="351">Mackay, 2000</emphasis>
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,1203,1896,1923]" pageId="350" pageNumber="351" type="distribution">
<paragraph blockId="350.[498,1203,1856,1923]" box="[498,1203,1896,1923]" pageId="350" pageNumber="351">
Distribution: 
<figureCitation box="[657,768,1896,1923]" captionStart="Figure 150" captionStartId="345.[524,589,731,753]" captionTargetBox="[498,1541,234,717]" captionTargetId="figure-281@345.[498,1534,234,702]" captionTargetPageId="345" captionText="Figure 150 Geographical and elevational distribution of the rugosus group. (A) Temnothorax parralensis sp. nov. (B) T. rugosus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-150" figureDoi="http://doi.org/10.5281/zenodo.5102440" httpUri="https://zenodo.org/record/5102440/files/figure.png" pageId="350" pageNumber="351">Fig. 150B</figureCitation>
; worker, gyne &amp; variability: 
<figureCitation box="[1104,1197,1896,1923]" captionStart="Figure 152" captionStartId="350.[524,589,723,745]" captionTargetBox="[498,1541,234,710]" captionTargetId="figure-268@350.[498,1534,234,695]" captionTargetPageId="350" captionText="Figure 152 Temnothorax rugosus. (A–C) Worker (LACMENT323296). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323337). (D) Full-face view. (E) Profile view. (F) Dorsal view. Morphological variant, Texas, U.S.A. (G–I) worker (LACMENT323260). (G) Full-face view. (H) Profile view. (I) Dorsal view. (J–L) Gyne (LACMENT323259). (J) Full-face view. (K) Profile view. (L) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-152" figureDoi="http://doi.org/10.5281/zenodo.5102447" httpUri="https://zenodo.org/record/5102447/files/figure.png" pageId="350" pageNumber="351">Fig. 152</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="351" pageNumber="352" type="reference_group">
<paragraph blockId="351.[498,1542,234,1934]" box="[530,1521,234,261]" pageId="351" pageNumber="352">
<treatmentCitationGroup box="[530,1521,234,261]" pageId="351" pageNumber="352">
<treatmentCitation author="Mackay WP" box="[530,1151,234,261]" page="399" pageId="351" pageNumber="352" year="2000">
<taxonomicName authority="Mackay, 2000: 399" authorityName="Mackay" authorityPageNumber="399" authorityYear="2000" box="[530,936,234,260]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="351" pageNumber="352" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[530,936,234,260]" italics="true" pageId="351" pageNumber="352">
Leptothorax rugosus 
<bibRefCitation author="Mackay WP" box="[774,995,234,261]" journalOrPublisher="Sociobiology" pageId="351" pageNumber="352" pagination="265 - 444" part="36" refId="ref271737" refString="Mackay WP. 2000. A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265 - 444." title="A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae)" type="journal article" year="2000">Mackay, 2000: 399</bibRefCitation>
</emphasis>
</taxonomicName>
, figs. 18, 160
</treatmentCitation>
. Worker. 
<collectingRegion box="[1269,1415,234,261]" country="Mexico" name="Nuevo Leon" pageId="351" pageNumber="352">Nuevo León</collectingRegion>
, 
<collectingCountry box="[1428,1517,234,260]" name="Mexico" pageId="351" pageNumber="352">Mexico</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="351.[498,1542,234,1934]" box="[530,1541,274,301]" pageId="351" pageNumber="352">
<treatmentCitationGroup box="[530,1541,274,301]" pageId="351" pageNumber="352">
<taxonomicName authority="(Mackay)" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[530,903,274,301]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="351" pageNumber="352" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[530,781,274,300]" italics="true" pageId="351" pageNumber="352">Temnothorax rugosus</emphasis>
(Mackay)
</taxonomicName>
: 
<treatmentCitation author="Bolton B." box="[913,1116,274,301]" page="272" pageId="351" pageNumber="352" year="2003">
<bibRefCitation author="Bolton B." box="[913,1116,274,301]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="351" pageNumber="352" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[913,1057,274,300]" italics="true" pageId="351" pageNumber="352">
<collectingRegion box="[913,991,274,300]" country="United Kingdom" name="Bolton" pageId="351" pageNumber="352">
<author box="[913,991,274,300]" fns="*" inm="true" npo="*" pageId="351" pageNumber="352">Bolton</author>
</collectingRegion>
, 
<year box="[1002,1057,274,300]" pageId="351" pageNumber="352">2003</year>
</emphasis>
: 272
</bibRefCitation>
</treatmentCitation>
. First combination in 
<emphasis box="[1372,1536,274,300]" italics="true" pageId="351" pageNumber="352">
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[1372,1532,274,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="351" pageNumber="352" phylum="Arthropoda" rank="genus">Temnothorax</taxonomicName>
.
</emphasis>
,
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="352" lastPageNumber="353" pageId="351" pageNumber="352" type="materials_examined">
<paragraph blockId="351.[498,1542,234,1934]" pageId="351" pageNumber="352">
<emphasis bold="true" box="[530,829,314,340]" pageId="351" pageNumber="352">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861521" collectingDate="1988-06-10" collectionCode="UTEP" collectorName="El Salto &amp; W. Mackay" country="Mexico" elevation="1415" location="El Salto" pageId="351" pageNumber="352" specimenCount="1" stateProvince="Nuevo Leon" typeStatus="paratype">
<emphasis box="[836,1031,314,340]" italics="true" pageId="351" pageNumber="352">
<typeStatus box="[836,938,315,340]" pageId="351" pageNumber="352">Paratype</typeStatus>
worker:
</emphasis>
<collectingCountry box="[1038,1153,314,340]" name="Mexico" pageId="351" pageNumber="352">MEXICO</collectingCountry>
: 
<collectingRegion box="[1162,1306,314,340]" country="Mexico" name="Nuevo Leon" pageId="351" pageNumber="352">Nuevo León</collectingRegion>
: 
<collectorName box="[1316,1404,314,340]" pageId="351" pageNumber="352">
<location LSID="urn:lsid:plazi:treatment:054FDB70FE99FEBFB49C1D5D7F34231E:88393CBDFE98FE85B04A1B277C4E2603" box="[1316,1404,314,340]" country="Mexico" name="El Salto" pageId="351" pageNumber="352" stateProvince="Nuevo Leon">El Salto</location>
</collectorName>
(Zaragosa), 
<quantity box="[498,594,354,381]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="351" pageNumber="352" unit="m" value="1415.0">
<elevation box="[498,594,354,381]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="351" pageNumber="352" unit="m" value="1415.0">1,415 m</elevation>
</quantity>
, 
<date box="[607,762,354,380]" pageId="351" pageNumber="352" value="1988-06-10">
<collectingDate box="[607,762,354,380]" pageId="351" pageNumber="352" value="1988-06-10">10 June 1988</collectingDate>
</date>
, 
<collectorName box="[774,912,354,380]" pageId="351" pageNumber="352">W. Mackay</collectorName>
#WM11012-10 (images of UTEPENT05451 examined on antweb.org) [
<collectionCode box="[692,773,393,420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="351" pageNumber="352" type="Museum">UTEP</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="351.[498,1542,234,1934]" lastBlockId="352.[498,1542,234,1934]" lastPageId="352" lastPageNumber="353" pageId="351" pageNumber="352">
<emphasis bold="true" box="[530,888,433,460]" pageId="351" pageNumber="352">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861318" collectingDate="1952-01-14" collectionCode="LACM" collectorName="Creighton" country="Mexico" elevation="1585" location="Jacala" pageId="351" pageNumber="352" specimenCount="22" specimenCount-worker="22" stateProvince="Hidalgo">
<collectingCountry box="[897,1012,434,460]" name="Mexico" pageId="351" pageNumber="352">MEXICO</collectingCountry>
: 
<collectingRegion box="[1024,1120,433,459]" country="Mexico" name="Hidalgo" pageId="351" pageNumber="352">Hidalgo</collectingRegion>
: 
<quantity box="[1132,1206,433,460]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="351" pageNumber="352" unit="km" value="11.0">11 km</quantity>
west of 
<location LSID="urn:lsid:plazi:treatment:054FDB70FE99FEBFB49C1D5D7F34231E:88393CBDFE98FE85B0751BAC7C52269C" box="[1307,1376,433,459]" country="Mexico" name="Jacala" pageId="351" pageNumber="352" stateProvince="Hidalgo">Jacala</location>
, 
<quantity box="[1390,1486,433,460]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="351" pageNumber="352" unit="m" value="1585.0">
<elevation box="[1390,1486,433,460]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="351" pageNumber="352" unit="m" value="1585.0">1,585 m</elevation>
</quantity>
, 
<date box="[498,692,473,500]" pageId="351" pageNumber="352" value="1952-01-14">
<collectingDate box="[498,692,473,500]" pageId="351" pageNumber="352" value="1952-01-14">14 January 1952</collectingDate>
</date>
, W.S. 
<collectorName box="[768,888,473,499]" pageId="351" pageNumber="352">Creighton</collectorName>
, in 
<taxonomicName box="[933,1175,473,500]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis box="[933,1175,473,500]" italics="true" pageId="351" pageNumber="352">Quercus polymorpha</emphasis>
</taxonomicName>
, 
<specimenCount box="[1190,1307,473,499]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323267) [
<collectionCode box="[754,842,512,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,513,539]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323268) [
<collectionCode box="[1237,1325,512,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,513,539]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323269) [
<collectionCode box="[754,842,552,579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,553,579]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323270) [
<collectionCode box="[1237,1325,552,579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,553,579]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323272) [
<collectionCode box="[754,842,592,619]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,593,619]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
(LACMENT323333) [
<collectionCode box="[1237,1326,592,619]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861609" collectingDate="1952-01-14" collectionCode="LACM" collectorName="Creighton" country="Mexico" elevation="1585" location="Jacala" pageId="351" pageNumber="352" specimenCount="17" specimenCount-worker="17" stateProvince="Hidalgo">
same data as previous, except: in oak twig, 
<specimenCount box="[849,965,633,659]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
(LACMENT323280) [
<collectionCode box="[1230,1318,632,659]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1333,1448,633,659]" pageId="351" pageNumber="352" type="worker">2 workers</specimenCount>
(LACMENT323324) [
<collectionCode box="[754,842,672,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,975,673,699]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323326) [
<collectionCode box="[1420,1508,672,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[498,616,712,738]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323327) [
<collectionCode box="[1061,1149,712,739]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1164,1282,712,738]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323328) [
<collectionCode box="[754,842,752,779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,975,752,778]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323271) [
<collectionCode box="[1420,1509,752,779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861756" collectingDate="1952-01-14" collectionCode="LACM" collectorName="Creighton" country="Mexico" elevation="1585" location="Jacala" pageId="351" pageNumber="352" specimenCount="8" specimenCount-worker="8" stateProvince="Hidalgo">
same data as previous, except: in oak limb, 
<specimenCount box="[1001,1117,792,818]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
, 1 dealate gyne (LACMENT323329) [
<collectionCode box="[505,593,831,858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[605,720,832,858]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
, 1 dealate gyne (LACMENT323330) [
<collectionCode box="[1151,1241,831,858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3321861472" collectingDate="1952-01-21" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1280" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="23" specimenCount-worker="23" stateProvince="Nuevo Leon">
<collectingRegion box="[1258,1402,832,858]" country="Mexico" name="Nuevo Leon" pageId="351" pageNumber="352">Nuevo León</collectingRegion>
: 
<collectorName box="[1411,1537,832,858]" pageId="351" pageNumber="352">Monterrey</collectorName>
, 
<collectorName box="[498,691,872,898]" pageId="351" pageNumber="352">Chipinque Mesa</collectorName>
, 
<quantity box="[701,795,872,898]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="351" pageNumber="352" unit="m" value="1280.0">
<elevation box="[701,795,872,898]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="351" pageNumber="352" unit="m" value="1280.0">1,280 m</elevation>
</quantity>
, 
<date box="[805,993,872,899]" pageId="351" pageNumber="352" value="1952-01-21">
<collectingDate box="[805,993,872,899]" pageId="351" pageNumber="352" value="1952-01-21">21 January 1952</collectingDate>
</date>
, W.S. 
<collectorName box="[1064,1184,872,898]" pageId="351" pageNumber="352">Creighton</collectorName>
, in 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[1224,1416,872,898]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="clivicola">
<emphasis box="[1224,1416,872,898]" italics="true" pageId="351" pageNumber="352">Quercus clivicola</emphasis>
</taxonomicName>
, 
<specimenCount box="[1429,1542,872,898]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
(LACMENT323302) [
<collectionCode box="[754,842,911,938]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,912,938]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323303) [
<collectionCode box="[1237,1325,911,938]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,912,938]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323306) [
<collectionCode box="[754,842,951,978]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,951,978]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323307) [
<collectionCode box="[1237,1325,951,978]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,951,978]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323314) [
<collectionCode box="[754,842,991,1018]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,991,1018]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323325) [
<collectionCode box="[1237,1326,991,1018]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861697" collectingDate="1952-01-22" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1280" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="39" specimenCount-worker="39" stateProvince="Nuevo Leon">
same data as previous, except: in 
<taxonomicName box="[734,913,1031,1058]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="canbyi">
<emphasis box="[734,913,1031,1058]" italics="true" pageId="351" pageNumber="352">Quercus canbyi</emphasis>
</taxonomicName>
, 
<specimenCount box="[928,1044,1031,1057]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323308) [
<collectionCode box="[1308,1396,1031,1058]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1411,1527,1031,1057]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323309) [
<collectionCode box="[754,842,1070,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1071,1097]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323310) [
<collectionCode box="[1237,1325,1070,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1071,1097]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323311) [
<collectionCode box="[754,843,1110,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]; 
<date box="[862,1055,1111,1138]" pageId="351" pageNumber="352" value="1952-01-22">
<collectingDate box="[862,1055,1111,1138]" pageId="351" pageNumber="352" value="1952-01-22">22 January 1952</collectingDate>
</date>
, in 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[1099,1294,1111,1137]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="clivicola">
<emphasis box="[1099,1294,1111,1137]" italics="true" pageId="351" pageNumber="352">Quercus clivicola</emphasis>
</taxonomicName>
, 
<specimenCount box="[1309,1426,1111,1137]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323286) [
<collectionCode box="[818,906,1150,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[921,1036,1151,1177]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323287) [
<collectionCode box="[1300,1388,1150,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1403,1519,1151,1177]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323288) [
<collectionCode box="[754,842,1190,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1191,1217]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323289) [
<collectionCode box="[1237,1325,1190,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1191,1217]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323290) [
<collectionCode box="[754,842,1230,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1230,1257]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323291) [
<collectionCode box="[1237,1326,1230,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861854" collectingDate="1952-02-18" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1280" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="24" specimenCount-worker="24" stateProvince="Nuevo Leon">
same data as previous, except: 
<quantity box="[702,798,1270,1297]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="351" pageNumber="352" unit="m" value="1280.0">
<elevation box="[702,798,1270,1297]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="351" pageNumber="352" unit="m" value="1280.0">1,280 m</elevation>
</quantity>
, 
<date box="[812,1020,1270,1297]" pageId="351" pageNumber="352" value="1952-02-18">
<collectingDate box="[812,1020,1270,1297]" pageId="351" pageNumber="352" value="1952-02-18">18 February 1952</collectingDate>
</date>
, in 
<taxonomicName box="[1066,1244,1270,1297]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="canbyi">
<emphasis box="[1066,1244,1270,1297]" italics="true" pageId="351" pageNumber="352">Quercus canbyi</emphasis>
</taxonomicName>
, 
<specimenCount box="[1259,1375,1270,1296]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323297) [
<collectionCode box="[754,842,1309,1336]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1310,1336]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323298) [
<collectionCode box="[1237,1325,1309,1336]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1310,1336]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323299) [
<collectionCode box="[754,842,1349,1376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1350,1376]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323300) [
<collectionCode box="[1237,1325,1349,1376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1350,1376]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323301) [
<collectionCode box="[754,842,1389,1416]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1390,1416]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323304) [
<collectionCode box="[1237,1326,1389,1416]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861588" collectingDate="1952-02-19" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="975" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="19" specimenCount-worker="19" stateProvince="Nuevo Leon">
same data as previous, except: 
<quantity box="[702,778,1429,1456]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="351" pageNumber="352" unit="m" value="975.0">
<elevation box="[702,778,1429,1456]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="351" pageNumber="352" unit="m" value="975.0">975 m</elevation>
</quantity>
, 
<date box="[792,1000,1429,1456]" pageId="351" pageNumber="352" value="1952-02-19">
<collectingDate box="[792,1000,1429,1456]" pageId="351" pageNumber="352" value="1952-02-19">19 February 1952</collectingDate>
</date>
, in 
<taxonomicName box="[1045,1182,1430,1456]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1045,1140,1430,1456]" italics="true" pageId="351" pageNumber="352">Quercus</emphasis>
sp.
</taxonomicName>
, 
<specimenCount box="[1196,1314,1430,1456]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 alate gyne (LACMENT323337) [
<collectionCode box="[754,842,1469,1496]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1470,1496]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323334) [
<collectionCode box="[1237,1325,1469,1496]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1470,1496]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323335) [
<collectionCode box="[754,842,1509,1536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1509,1535]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323336) [
<collectionCode box="[1237,1325,1509,1536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1509,1535]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323338) [
<collectionCode box="[754,843,1549,1576]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861651" collectingDate="1952-02-19" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="975" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="14" specimenCount-worker="14" stateProvince="Nuevo Leon">
same data as previous, except: in 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[1258,1453,1549,1576]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="clivicola">
<emphasis box="[1258,1453,1549,1576]" italics="true" pageId="351" pageNumber="352">Quercus clivicola</emphasis>
</taxonomicName>
, 
<specimenCount pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323273) [
<collectionCode box="[1038,1126,1588,1615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1141,1257,1589,1615]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323274) [
<collectionCode box="[505,593,1628,1655]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[608,726,1629,1655]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323276) [
<collectionCode box="[1170,1258,1628,1655]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1273,1389,1629,1655]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323279) [
<collectionCode box="[752,841,1668,1695]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861743" collectingDate="1952-02-19" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="975" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="11" specimenCount-worker="11" stateProvince="Nuevo Leon">
same data as previous, except: in 
<taxonomicName box="[1239,1415,1669,1695]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="canbyi">
<emphasis box="[1239,1415,1669,1695]" italics="true" pageId="351" pageNumber="352">Quercus canbyi</emphasis>
</taxonomicName>
, 
<specimenCount box="[1429,1542,1669,1695]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323275) [
<collectionCode box="[754,842,1708,1735]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1709,1735]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323277) [
<collectionCode box="[1237,1325,1708,1735]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1709,1735]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
(LACMENT323278) [
<collectionCode box="[752,841,1748,1775]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861473" collectingDate="1952-02-19" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1100" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="24" specimenCount-worker="24" stateProvince="Nuevo Leon">
same data as previous, except: 
<quantity box="[1207,1301,1749,1775]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="351" pageNumber="352" unit="m" value="1100.0">
<elevation box="[1207,1301,1749,1775]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="351" pageNumber="352" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
, in 
<taxonomicName box="[1342,1535,1749,1775]" class="Magnoliopsida" family="Fabaceae" genus="Cercis" kingdom="Plantae" order="Fabales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="reniformis">
<emphasis box="[1342,1535,1749,1775]" italics="true" pageId="351" pageNumber="352">Cercis reniformis</emphasis>
</taxonomicName>
, 
<specimenCount box="[498,614,1788,1814]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323292) [
<collectionCode box="[878,966,1788,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[981,1097,1788,1814]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323293) [
<collectionCode box="[1361,1449,1788,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323294) [
<collectionCode box="[852,940,1828,1855]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[953,1067,1828,1854]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323295) [
<collectionCode box="[1327,1415,1828,1855]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1428,1542,1828,1854]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323296) [
<collectionCode box="[754,842,1867,1894]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1868,1894]" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323361) [
<collectionCode box="[1237,1326,1867,1894]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861874" collectingDate="1952-02-19" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1220" lastPageId="352" lastPageNumber="353" location="Nuevo Leon" pageId="351" pageNumber="352" specimenCount="22" specimenCount-worker="22" stateProvince="Nuevo Leon">
same data as previous, except: 
<quantity box="[702,798,1908,1934]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="351" pageNumber="352" unit="m" value="1220.0">
<elevation box="[702,798,1908,1934]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="351" pageNumber="352" unit="m" value="1220.0">1,220 m</elevation>
</quantity>
, in 
<taxonomicName box="[844,1023,1908,1934]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="351" pageNumber="352" phylum="Tracheophyta" rank="species" species="canbyi">
<emphasis box="[844,1023,1908,1934]" italics="true" pageId="351" pageNumber="352">Quercus canbyi</emphasis>
</taxonomicName>
, 
<specimenCount box="[1038,1154,1908,1934]" pageId="351" pageNumber="352" type="worker">3 workers</specimenCount>
(LACMENT323281) [
<collectionCode box="[1418,1506,1907,1934]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="351" pageNumber="352" type="Museum">LACM</collectionCode>
] 
<specimenCount lastPageId="352" lastPageNumber="353" pageId="351" pageNumber="352" type="worker">4 workers</specimenCount>
(LACMENT323282) [
<collectionCode box="[852,940,234,261]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[953,1067,234,260]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323283) [
<collectionCode box="[1327,1415,234,261]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1428,1542,234,260]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323284) [
<collectionCode box="[754,842,273,300]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,274,300]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323285) [
<collectionCode box="[1237,1325,273,300]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1458,274,300]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323322) [
<collectionCode box="[930,1019,313,340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861570" collectingDate="1952-04-21" collectionCode="LACM" collectorName="W.S. Creighton" country="Mexico" elevation="1280" location="Nuevo Leon" pageId="352" pageNumber="353" specimenCount="12" specimenCount-worker="12" stateProvince="Nuevo Leon">
same data as previous, except: 
<quantity box="[1402,1498,314,340]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="352" pageNumber="353" unit="m" value="1280.0">
<elevation box="[1402,1498,314,340]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="352" pageNumber="353" unit="m" value="1280.0">1,280 m</elevation>
</quantity>
, 
<date box="[498,663,354,381]" pageId="352" pageNumber="353" value="1952-04-21">
<collectingDate box="[498,663,354,381]" pageId="352" pageNumber="353" value="1952-04-21">21 April 1952</collectingDate>
</date>
, in 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[708,903,354,380]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="clivicola">
<emphasis box="[708,903,354,380]" italics="true" pageId="352" pageNumber="353">Quercus clivicola</emphasis>
</taxonomicName>
, (LACMENT323305) [
<collectionCode box="[1174,1262,353,380]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1277,1393,354,380]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323318) [
<collectionCode box="[754,842,393,420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,394,420]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323319) [
<collectionCode box="[1237,1325,393,420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,394,420]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323320) [
<collectionCode box="[754,843,433,460]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861861" collectingDate="1952-02-16" collectionCode="LACM" collectorName="W. S. Creighton" country="Mexico" elevation="1890" location="Iturbide" municipality="Iturbide" pageId="352" pageNumber="353" specimenCount="63" specimenCount-worker="63" stateProvince="Nuevo Leon">
same data as previous, except: in 
<taxonomicName box="[1258,1436,433,460]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="canbyi">
<emphasis box="[1258,1436,433,460]" italics="true" pageId="352" pageNumber="353">Quercus canbyi</emphasis>
</taxonomicName>
, 
<specimenCount box="[498,614,473,499]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323312) [
<collectionCode box="[878,966,473,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[981,1097,473,499]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323313) [
<collectionCode box="[1361,1449,473,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323315) [
<collectionCode box="[852,940,512,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[953,1067,513,539]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323316) [
<collectionCode box="[1327,1415,512,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1428,1542,513,539]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323317) [
<collectionCode box="[754,842,552,579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,553,579]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323321) [
<collectionCode box="[1237,1325,552,579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,553,579]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323323) [
<collectionCode box="[754,843,592,619]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]; 
<quantity box="[863,938,593,619]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="352" pageNumber="353" unit="km" value="10.0">10 km</quantity>
west of 
<collectingMunicipality box="[1039,1135,593,619]" pageId="352" pageNumber="353">Iturbide</collectingMunicipality>
, 
<quantity box="[1148,1244,593,619]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="352" pageNumber="353" unit="m" value="1890.0">
<elevation box="[1148,1244,593,619]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="352" pageNumber="353" unit="m" value="1890.0">1,890 m</elevation>
</quantity>
, 
<date box="[1258,1466,593,620]" pageId="352" pageNumber="353" value="1952-02-16">
<collectingDate box="[1258,1466,593,620]" pageId="352" pageNumber="353" value="1952-02-16">16 February 1952</collectingDate>
</date>
, 
<collectorName box="[498,682,633,659]" pageId="352" pageNumber="353">W.S. Creighton</collectorName>
, in 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[727,922,633,659]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="clivicola">
<emphasis box="[727,922,633,659]" italics="true" pageId="352" pageNumber="353">Quercus clivicola</emphasis>
</taxonomicName>
, 
<specimenCount box="[938,1053,633,659]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323358) [
<collectionCode box="[1318,1406,632,659]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1421,1536,633,659]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323360) [
<collectionCode box="[754,842,672,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,673,699]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323362) [
<collectionCode box="[1237,1325,672,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,673,699]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323363) [
<collectionCode box="[754,842,712,739]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,712,738]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323364) [
<collectionCode box="[1237,1325,712,739]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,712,738]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323365) [
<collectionCode box="[754,842,752,779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,752,778]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323366) [
<collectionCode box="[1237,1325,752,779]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1458,752,778]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323367) [
<collectionCode box="[924,1013,792,819]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]; 
<quantity box="[1032,1091,792,818]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="352" pageNumber="353" unit="km" value="3.0">3 km</quantity>
west of 
<location LSID="urn:lsid:plazi:treatment:054FDB70FE99FEBFB49C1D5D7F34231E:88393CBDFEA7FEBAB1CD19057C312465" box="[1187,1283,792,818]" country="Mexico" municipality="Iturbide" name="Iturbide" pageId="352" pageNumber="353" stateProvince="Nuevo Leon">Iturbide</location>
, 
<quantity box="[1295,1390,792,819]" metricMagnitude="3" metricUnit="m" metricValue="1.675" pageId="352" pageNumber="353" unit="m" value="1675.0">
<elevation box="[1295,1390,792,819]" metricMagnitude="3" metricUnit="m" metricValue="1.675" pageId="352" pageNumber="353" unit="m" value="1675.0">1,675 m</elevation>
</quantity>
, 
<date pageId="352" pageNumber="353" value="1952-02-16">
<collectingDate pageId="352" pageNumber="353" value="1952-02-16">16 February 1952</collectingDate>
</date>
, 
<collectorName box="[567,747,832,858]" pageId="352" pageNumber="353">W.S. Creighton</collectorName>
, in hollow stem of 
<taxonomicName box="[962,1059,832,858]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[962,1019,832,858]" italics="true" pageId="352" pageNumber="353">Rhus</emphasis>
sp.
</taxonomicName>
, 
<specimenCount box="[1070,1183,832,858]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323359) [
<collectionCode box="[1441,1531,831,858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="352.[498,1542,234,1934]" pageId="352" pageNumber="353">
<materialsCitation ID-GBIF-Occurrence="3321861835" collectingDate="1953-05-11" collectionCode="LACM" collectorName="W. S. Creighton" country="United States of America" county="Jeff Davis County" elevation="1525" location="Davis Mountains State Park" pageId="352" pageNumber="353" specimenCount="3" specimenCount-worker="3" stateProvince="Texas">
<collectingCountry box="[530,607,872,898]" name="United States of America" pageId="352" pageNumber="353">U.S.A.</collectingCountry>
: 
<collectingRegion box="[620,690,872,898]" country="United States of America" name="Texas" pageId="352" pageNumber="353">Texas</collectingRegion>
: 
<collectingCounty box="[702,914,872,898]" pageId="352" pageNumber="353">Jeff Davis County</collectingCounty>
: 
<location LSID="urn:lsid:plazi:treatment:054FDB70FE99FEBFB49C1D5D7F34231E:88393CBDFEA7FEBAB6F019757DDB24D5" box="[926,1257,872,898]" country="United States of America" county="Jeff Davis County" name="Davis Mountains State Park" pageId="352" pageNumber="353" stateProvince="Texas">Davis Mountains State Park</location>
, 
<quantity box="[1270,1366,872,899]" metricMagnitude="3" metricUnit="m" metricValue="1.525" pageId="352" pageNumber="353" unit="m" value="1525.0">
<elevation box="[1270,1366,872,899]" metricMagnitude="3" metricUnit="m" metricValue="1.525" pageId="352" pageNumber="353" unit="m" value="1525.0">1,525 m</elevation>
</quantity>
, 
<date box="[1380,1536,872,899]" pageId="352" pageNumber="353" value="1953-05-11">
<collectingDate box="[1380,1536,872,899]" pageId="352" pageNumber="353" value="1953-05-11">11 May 1953</collectingDate>
</date>
, 
<collectorName box="[498,682,912,938]" pageId="352" pageNumber="353">W.S. Creighton</collectorName>
, in 
<taxonomicName authorityName="Liebmann" authorityYear="1854" box="[727,896,912,938]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="grisea">
<emphasis box="[727,896,912,938]" italics="true" pageId="352" pageNumber="353">Quercus grisea</emphasis>
</taxonomicName>
, 
<specimenCount box="[912,1027,912,938]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323257) [
<collectionCode box="[1292,1381,911,938]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861896" collectingDate="1953-05-11" collectionCode="LACM" collectorName="W. S. Creighton" country="United States of America" county="Jeff Davis County" elevation="1525" location="Davis Mountains State Park" pageId="352" pageNumber="353" specimenCount="2" specimenCount-worker="2" stateProvince="Texas">
same data as previous, except: in deciduous oak, 
<specimenCount box="[942,1056,951,978]" pageId="352" pageNumber="353" type="worker">2 workers</specimenCount>
(LACMENT323258) [
<collectionCode box="[1318,1407,951,978]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861903" collectingDate="1953-05-16" collectionCode="LACM" collectorName="W. S. Creighton" country="United States of America" county="Jeff Davis County" elevation="1525" location="Davis Mountains State Park" pageId="352" pageNumber="353" specimenCount="34" specimenCount-worker="34" stateProvince="Texas">
same data as previous, except: 
<date box="[734,890,991,1018]" pageId="352" pageNumber="353" value="1953-05-16">
<collectingDate box="[734,890,991,1018]" pageId="352" pageNumber="353" value="1953-05-16">16 May 1953</collectingDate>
</date>
, 
<collectorName box="[903,1087,991,1018]" pageId="352" pageNumber="353">W.S. Creighton</collectorName>
, in 
<taxonomicName authorityName="Liebmann" authorityYear="1854" box="[1131,1301,992,1018]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="grisea">
<emphasis box="[1131,1301,992,1018]" italics="true" pageId="352" pageNumber="353">Quercus grisea</emphasis>
</taxonomicName>
, 
<specimenCount box="[1316,1432,991,1018]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323252) [
<collectionCode box="[754,842,1031,1058]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1031,1057]" pageId="352" pageNumber="353" type="worker">2 workers</specimenCount>
(LACMENT323253) [
<collectionCode box="[1237,1325,1031,1058]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1031,1057]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323254) [
<collectionCode box="[754,842,1070,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1071,1097]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323255) [
<collectionCode box="[1237,1325,1070,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1071,1097]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323256) [
<collectionCode box="[754,842,1110,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,975,1111,1137]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
, 1 dealate gyne (LACMENT323259) [
<collectionCode box="[1420,1508,1110,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[498,614,1151,1177]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323260) [
<collectionCode box="[878,966,1150,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[981,1097,1151,1177]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323261) [
<collectionCode box="[1361,1449,1150,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323262) [
<collectionCode box="[856,944,1190,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[958,1074,1191,1217]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323263) [
<collectionCode box="[1339,1428,1190,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3321861767" collectingDate="1952-05-20" collectionCode="LACM" collectorName="W. S. Creighton" country="United States of America" county="Jeff Davis County" elevation="1525" location="Davis Mountains State Park" pageId="352" pageNumber="353" specimenCount="20" specimenCount-worker="20" stateProvince="Texas">
same data as previous, except: 
<date box="[793,948,1230,1257]" pageId="352" pageNumber="353" value="1952-05-20">
<collectingDate box="[793,948,1230,1257]" pageId="352" pageNumber="353" value="1952-05-20">20 May 1952</collectingDate>
</date>
, 
<collectorName box="[961,1145,1230,1257]" pageId="352" pageNumber="353">W.S. Creighton</collectorName>
, in 
<taxonomicName authorityName="Liebmann" authorityYear="1854" box="[1190,1359,1231,1257]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="352" pageNumber="353" phylum="Tracheophyta" rank="species" species="grisea">
<emphasis box="[1190,1359,1231,1257]" italics="true" pageId="352" pageNumber="353">Quercus grisea</emphasis>
</taxonomicName>
, 
<specimenCount box="[1375,1490,1230,1257]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323264) [
<collectionCode box="[754,842,1270,1297]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1270,1296]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323265) [
<collectionCode box="[1237,1325,1270,1297]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1270,1296]" pageId="352" pageNumber="353" type="worker">4 workers</specimenCount>
(LACMENT323266) [
<collectionCode box="[754,842,1309,1336]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1310,1336]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323331) [
<collectionCode box="[1237,1325,1309,1336]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[1340,1456,1310,1336]" pageId="352" pageNumber="353" type="worker">2 workers</specimenCount>
(LACMENT323332) [
<collectionCode box="[754,842,1349,1376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[857,973,1350,1376]" pageId="352" pageNumber="353" type="worker">3 workers</specimenCount>
(LACMENT323339) [
<collectionCode box="[1237,1327,1349,1376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="352" pageNumber="353" type="Museum">LACM</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="352" pageNumber="353" type="distribution">
<paragraph blockId="352.[498,1542,234,1934]" pageId="352" pageNumber="353">
<emphasis bold="true" box="[530,755,1390,1416]" pageId="352" pageNumber="353">Geographic range:</emphasis>
Mid-to-high elevations of southwestern Texas to 
<collectingRegion box="[1344,1504,1390,1416]" country="Mexico" name="Hidalgo" pageId="352" pageNumber="353">Hidalgo State</collectingRegion>
, 
<collectingCountry box="[498,584,1430,1456]" name="Mexico" pageId="352" pageNumber="353">Mexico</collectingCountry>
(
<figureCitation box="[603,714,1429,1456]" captionStart="Figure 150" captionStartId="345.[524,589,731,753]" captionTargetBox="[498,1541,234,717]" captionTargetId="figure-281@345.[498,1534,234,702]" captionTargetPageId="345" captionText="Figure 150 Geographical and elevational distribution of the rugosus group. (A) Temnothorax parralensis sp. nov. (B) T. rugosus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-150" figureDoi="http://doi.org/10.5281/zenodo.5102440" httpUri="https://zenodo.org/record/5102440/files/figure.png" pageId="352" pageNumber="353">Fig. 150B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="353" lastPageNumber="354" pageId="352" pageNumber="353" type="diagnosis">
<paragraph blockId="352.[498,1542,234,1934]" pageId="352" pageNumber="353">
<emphasis bold="true" box="[530,753,1469,1496]" pageId="352" pageNumber="353">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[760,1011,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[760,1011,1470,1496]" italics="true" pageId="352" pageNumber="353">Temnothorax rugosus</emphasis>
</taxonomicName>
can be separated from all other species in the 
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[498,573,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[498,573,1509,1535]" italics="true" pageId="352" pageNumber="353">salvini</emphasis>
</taxonomicName>
clade by the following character combination: anterior clypeal margin emarginate; metanotal groove absent; propodeum not depressed; hind femora weakly to strongly incrassate; petiole cuneiform to subcuneiform; dorsum of head, mesosoma, waist segments and gaster with erect, short, blunt-tipped setae; legs without erect setae; integument testaceous yellow, with legs lighter yellow.
</paragraph>
<paragraph blockId="352.[498,1542,234,1934]" lastBlockId="353.[498,1542,234,1934]" lastPageId="353" lastPageNumber="354" pageId="352" pageNumber="353">
<emphasis bold="true" box="[530,717,1708,1734]" pageId="352" pageNumber="353">Similar species:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[725,1024,1709,1735]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis box="[725,1024,1709,1735]" italics="true" pageId="352" pageNumber="353">Temnothorax acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1032,1124,1709,1735]" pageId="352" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis box="[1136,1417,1709,1734]" italics="true" pageId="352" pageNumber="353">
<taxonomicName box="[1136,1238,1709,1734]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="nitens">T. nitens</taxonomicName>
, 
<taxonomicName box="[1250,1417,1709,1734]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="tricarinatus">T. tricarinatus</taxonomicName>
</emphasis>
, species of the 
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[544,640,1749,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[544,640,1749,1774]" italics="true" pageId="352" pageNumber="353">annexus</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Santschi" baseAuthorityYear="1908" box="[701,795,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis box="[701,795,1748,1774]" italics="true" pageId="352" pageNumber="353">silvestrii</emphasis>
</taxonomicName>
groups. 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[898,1151,1748,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[898,1151,1748,1774]" italics="true" pageId="352" pageNumber="353">Temnothorax rugosus</emphasis>
</taxonomicName>
may be separated from 
<taxonomicName box="[1439,1541,1749,1774]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="nitens">
<emphasis box="[1439,1541,1749,1774]" italics="true" pageId="352" pageNumber="353">T. nitens</emphasis>
</taxonomicName>
and 
<taxonomicName box="[549,714,1789,1814]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis box="[549,714,1789,1814]" italics="true" pageId="352" pageNumber="353">T. tricarinatus</emphasis>
</taxonomicName>
by the moderately incrassate femora (not apparent in smaller workers), and the weakly sculptured first gastral tergite (which is smooth and shining in 
<taxonomicName box="[1430,1532,1829,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="nitens">
<emphasis box="[1430,1532,1829,1854]" italics="true" pageId="352" pageNumber="353">T. nitens</emphasis>
</taxonomicName>
and 
<taxonomicName box="[551,719,1869,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis box="[551,719,1869,1894]" italics="true" pageId="352" pageNumber="353">T. tricarinatus</emphasis>
</taxonomicName>
). Many of the preceding characters, in combination, are shared with members of the 
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[684,780,1909,1934]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[684,780,1909,1934]" italics="true" pageId="352" pageNumber="353">annexus</emphasis>
</taxonomicName>
species group, which 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1032,1149,1909,1934]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="352" pageNumber="353" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[1032,1149,1909,1934]" italics="true" pageId="352" pageNumber="353">T. rugosus</emphasis>
</taxonomicName>
is closely related to. 
<taxonomicName lastPageId="353" lastPageNumber="354" pageId="352" pageNumber="353">
<emphasis box="[1385,1542,1908,1934]" italics="true" pageId="352" pageNumber="353">Temnothorax</emphasis>
<emphasis box="[498,585,235,260]" italics="true" pageId="353" pageNumber="354">rugosus</emphasis>
</taxonomicName>
can be separated from the species of the 
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1065,1161,235,260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="353" pageNumber="354" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1065,1161,235,260]" italics="true" pageId="353" pageNumber="354">annexus</emphasis>
</taxonomicName>
group by the cuneiform petiolar node, which is laterally compressed in dorsal view; all species in the 
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1308,1404,275,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="353" pageNumber="354" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1308,1404,275,300]" italics="true" pageId="353" pageNumber="354">annexus</emphasis>
</taxonomicName>
group have quadrate to subquadrate petiolar nodes. Finally, the areolate first gastral tergite is similar to members of the 
<taxonomicName baseAuthorityName="Santschi" baseAuthorityYear="1908" box="[800,894,354,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="353" pageNumber="354" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis box="[800,894,354,380]" italics="true" pageId="353" pageNumber="354">silvestrii</emphasis>
</taxonomicName>
group, but members of that group never have incrassate femora, and always have subquadrate petiolar nodes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="357" lastPageNumber="358" pageId="353" pageNumber="354" type="description">
<paragraph blockId="353.[498,1542,234,1934]" pageId="353" pageNumber="354">
<emphasis bold="true" box="[530,1039,433,460]" pageId="353" pageNumber="354">
Worker measurements &amp; indices (
<emphasis bold="true" box="[950,966,434,460]" italics="true" pageId="353" pageNumber="354">n</emphasis>
= 27)
</emphasis>
: SL = 
<geoCoordinate box="[1118,1180,433,460]" degrees="0.465" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.465">0.465</geoCoordinate>
<geoCoordinate box="[1180,1257,433,459]" degrees="0.602" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.602">–0.602</geoCoordinate>
(0.527); FRS = 
<geoCoordinate box="[579,641,473,499]" degrees="0.207" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.207">0.207</geoCoordinate>
<geoCoordinate box="[642,719,473,499]" degrees="0.279" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.279">–0.279</geoCoordinate>
(0.228); CW = 
<geoCoordinate box="[907,969,473,499]" degrees="0.610" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.61">0.610</geoCoordinate>
<geoCoordinate box="[969,1046,473,499]" degrees="0.800" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.8">–0.800</geoCoordinate>
(0.700); CWb = 
<geoCoordinate box="[1249,1311,473,500]" degrees="0.563" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.563">0.563</geoCoordinate>
<geoCoordinate box="[1311,1388,473,499]" degrees="0.733" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.733">–0.733</geoCoordinate>
(0.642); PoOC = 
<geoCoordinate box="[604,666,513,540]" degrees="0.265" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.265">0.265</geoCoordinate>
<geoCoordinate box="[667,744,513,540]" degrees="0.325" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.325">–0.325</geoCoordinate>
(0.288); CL = 
<geoCoordinate box="[919,981,513,539]" degrees="0.701" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.701">0.701</geoCoordinate>
<geoCoordinate box="[981,1058,513,539]" degrees="0.874" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.874">–0.874</geoCoordinate>
(0.773); EL = 
<geoCoordinate box="[1231,1293,513,540]" degrees="0.157" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.157">0.157</geoCoordinate>
<geoCoordinate box="[1294,1370,513,540]" degrees="0.255" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.255">–0.255</geoCoordinate>
(0.198); EW = 
<geoCoordinate box="[577,639,553,579]" degrees="0.116" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.116">0.116</geoCoordinate>
<geoCoordinate box="[640,716,553,579]" degrees="0.166" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.166">–0.166</geoCoordinate>
(0.142); MD = 
<geoCoordinate box="[905,967,553,579]" degrees="0.173" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.173">0.173</geoCoordinate>
<geoCoordinate box="[967,1044,553,579]" degrees="0.217" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.217">–0.217</geoCoordinate>
(0.197); WL = 
<geoCoordinate box="[1228,1290,553,579]" degrees="0.824" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.824">0.824</geoCoordinate>
<geoCoordinate box="[1291,1368,553,580]" degrees="1.095" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-1.095">–1.095</geoCoordinate>
(0.948); SPST = 
<geoCoordinate box="[594,656,593,619]" degrees="0.141" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.141">0.141</geoCoordinate>
<geoCoordinate box="[657,734,593,619]" degrees="0.219" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.219">–0.219</geoCoordinate>
(0.182); MPST = 
<geoCoordinate box="[948,1010,593,619]" degrees="0.203" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.203">0.203</geoCoordinate>
<geoCoordinate box="[1011,1088,593,619]" degrees="0.334" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.334">–0.334</geoCoordinate>
(0.281); PEL = 
<geoCoordinate box="[1277,1339,593,619]" degrees="0.242" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.242">0.242</geoCoordinate>
<geoCoordinate box="[1339,1416,593,619]" degrees="0.362" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.362">–0.362</geoCoordinate>
(0.291); NOL = 
<geoCoordinate box="[591,653,633,659]" degrees="0.147" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.147">0.147</geoCoordinate>
<geoCoordinate box="[654,731,633,659]" degrees="0.230" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.23">–0.230</geoCoordinate>
(0.178); NOH = 
<geoCoordinate box="[937,999,632,659]" degrees="0.105" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.105">0.105</geoCoordinate>
<geoCoordinate box="[999,1076,633,659]" degrees="0.180" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.18">–0.180</geoCoordinate>
(0.133); PEH = 
<geoCoordinate box="[1272,1334,633,659]" degrees="0.226" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.226">0.226</geoCoordinate>
<geoCoordinate box="[1334,1411,633,659]" degrees="0.303" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.303">–0.303</geoCoordinate>
(0.260); PPL = 
<geoCoordinate box="[581,643,673,699]" degrees="0.170" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.17">0.170</geoCoordinate>
<geoCoordinate box="[643,720,673,699]" degrees="0.289" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.289">–0.289</geoCoordinate>
(0.211); PPH = 
<geoCoordinate box="[916,978,673,699]" degrees="0.210" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.21">0.210</geoCoordinate>
<geoCoordinate box="[978,1055,672,699]" degrees="0.306" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.306">–0.306</geoCoordinate>
(0.244); PW = 
<geoCoordinate box="[1241,1303,672,698]" degrees="0.368" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.368">0.368</geoCoordinate>
<geoCoordinate box="[1303,1380,672,699]" degrees="0.502" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.502">–0.502</geoCoordinate>
(0.428); SBPA = 
<geoCoordinate box="[600,662,712,739]" degrees="0.135" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.135">0.135</geoCoordinate>
<geoCoordinate box="[662,739,712,738]" degrees="0.213" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.213">–0.213</geoCoordinate>
(0.160); SPTI = 
<geoCoordinate box="[938,1000,712,738]" degrees="0.148" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.148">0.148</geoCoordinate>
<geoCoordinate box="[1000,1077,712,738]" degrees="0.241" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.241">–0.241</geoCoordinate>
(0.187); PEW = 
<geoCoordinate box="[1279,1341,712,738]" degrees="0.149" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.149">0.149</geoCoordinate>
<geoCoordinate box="[1341,1418,712,738]" degrees="0.217" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.217">–0.217</geoCoordinate>
(0.178); PNW = 
<geoCoordinate box="[599,661,752,778]" degrees="0.066" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.066">0.066</geoCoordinate>
<geoCoordinate box="[661,738,752,778]" degrees="0.117" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.117">–0.117</geoCoordinate>
(0.090); PPW = 
<geoCoordinate box="[940,1002,752,779]" degrees="0.245" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.245">0.245</geoCoordinate>
<geoCoordinate box="[1002,1079,752,778]" degrees="0.393" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.393">–0.393</geoCoordinate>
(0.290); HFL = 
<geoCoordinate box="[1273,1335,752,778]" degrees="0.484" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.484">0.484</geoCoordinate>
<geoCoordinate box="[1335,1412,752,778]" degrees="0.680" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.68">–0.680</geoCoordinate>
(0.578); HFWmax = 
<geoCoordinate box="[648,710,792,818]" degrees="0.148" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.148">0.148</geoCoordinate>
<geoCoordinate box="[711,788,792,818]" degrees="0.209" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.209">–0.209</geoCoordinate>
(0.177); HFWmin = 
<geoCoordinate box="[1041,1103,792,818]" degrees="0.049" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.049">0.049</geoCoordinate>
<geoCoordinate box="[1104,1181,792,818]" degrees="0.081" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.081">–0.081</geoCoordinate>
(0.059); CS = 
<geoCoordinate box="[1354,1416,792,818]" degrees="0.914" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.914">0.914</geoCoordinate>
<geoCoordinate box="[1417,1494,792,818]" degrees="1.170" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-1.17">–1.170</geoCoordinate>
(1.029); ES = 
<geoCoordinate box="[660,722,832,858]" degrees="0.219" direction="north" orientation="latitude" pageId="353" pageNumber="354" precision="55" value="0.219">0.219</geoCoordinate>
<geoCoordinate box="[723,799,832,858]" degrees="0.327" direction="west" orientation="longitude" pageId="353" pageNumber="354" precision="55" value="-0.327">–0.327</geoCoordinate>
(0.269); SI = 77–89 (82); OI = 24–29 (26); CI = 80–87 (83); WLI = 139–157 (148); SBI = 22–29 (25); PSI = 17–22 (19); PWI = 140–181 (163); PLI = 113–176 (139); NI = 105–173 (135); PNWI = 38–67 (50); NLI = 49–75 (61); FI = 225–354 (301).
</paragraph>
<paragraph blockId="353.[498,1542,234,1934]" pageId="353" pageNumber="354">
<emphasis bold="true" box="[530,775,991,1018]" pageId="353" pageNumber="354">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 80–87). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about two times the maximum width of the antennal scape (SI 77–89). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin concave medially, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="353.[498,1542,234,1934]" lastBlockId="354.[498,1542,234,1934]" lastPageId="354" lastPageNumber="355" pageId="353" pageNumber="354">
In profile view, compound eyes ovular and moderately large (OI 24–29), with 13 ommatidia in longest row. Pronotal declivity distinct: dorsal margin of anterior face of pronotum marked by a weak carina and change in sculpture; neck and anterior face of pronotum forming a ~120 
<emphasis box="[801,811,1624,1642]" italics="true" pageId="353" pageNumber="354">
<superScript attach="left" box="[801,811,1624,1642]" fontSize="8" pageId="353" pageNumber="354">°</superScript>
</emphasis>
angle. Mesosoma very weakly sinuate: promesonotum forming an even convexity from where it joins the pronotal declivity; propodeum weakly depressed below the level of the promesonotum and descending sharply to the base of the propodeal spines on the posterior quarter. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines weakly developed and short (PSI 17–22), about a third as long as the propodeal declivity, flared at the base, stout, triangular, and acute. Propodeal declivity straight, forming a rounded ~120 
<emphasis box="[557,567,389,407]" italics="true" pageId="354" pageNumber="355">
<superScript attach="left" box="[557,567,389,407]" fontSize="8" pageId="354" pageNumber="355">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion a third of the way to the propodeal spiracle. Petiole moderately long (PLI 113–176), with tubercles anterodorsally. Subpetiolar process in the form of a blunt, triangular tooth which grades evenly into the ventral petiole margin posteriorly. Petiolar peduncle short: comprising about a quarter of the total petiole length. Petiolar node truncate but dorsum is extremely angled, so that in many specimens it appears cuneiform: anterior face forming a sharp ~90 
<emphasis box="[634,644,668,686]" italics="true" pageId="354" pageNumber="355">
<superScript attach="left" box="[634,644,668,686]" fontSize="8" pageId="354" pageNumber="355">°</superScript>
</emphasis>
angle with the dorsal face, which is flat; dorsal face meeting the posterior face at a sharp ~120 
<emphasis box="[682,692,708,726]" italics="true" pageId="354" pageNumber="355">
<superScript attach="left" box="[682,692,708,726]" fontSize="8" pageId="354" pageNumber="355">°</superScript>
</emphasis>
angle; very short posterior face forms a ~110 
<emphasis box="[1232,1242,708,726]" italics="true" pageId="354" pageNumber="355">
<superScript attach="left" box="[1232,1242,708,726]" fontSize="8" pageId="354" pageNumber="355">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly convex anterodorsally; weakly lobed ventrally.
</paragraph>
<paragraph blockId="354.[498,1542,234,1934]" pageId="354" pageNumber="355">In dorsal view, humeri developed: angulate anterolaterally and slightly wider than the rest of the mesosoma; mesothoracic spiracles protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture represented by a weak sulcus and disruption in the ground sculpture. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about twice their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node, when viewed at posterodorsal angle, cuneiform: narrowed apically and flat dorsally; node narrower than the peduncle and the caudal cylinder. Postpetiole narrow (PWI ~ 163) and campaniform. Anterior margin of the postpetiole convex and rounds evenly into the lateral margins; lateral margins parallel to each other; posterior corners rounded; posterior margin flat. Metafemur moderately to strongly incrassate (FI 225–354).</paragraph>
<paragraph blockId="354.[498,1542,234,1934]" lastBlockId="355.[498,1542,234,1934]" lastPageId="355" lastPageNumber="356" pageId="354" pageNumber="355">Sculpture: median clypeal carina present, extending posteriorly to the frontal triangle, and flanked on either side by three weaker carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture shining through weak areolae. Antennal scapes areolate. Cephalic dorsum areolate, with costa over the ground sculpture, which become rugose on the posterior sixth of the head; concentric rugulae surrounding the antennal insertions. Lateral surfaces of head areolate, with coarse rugae over the ground sculpture. Ventral surface of head smooth and shining medially, but otherwise weakly areolate with weak costulae. Pronotal neck areolate. Lateral surfaces of mesosoma weakly areolate, with coarse, costae over the ground sculpture; region between the propodeal spiracle and propodeal spines areolate, without rugae. Propodeal declivity areolate, with fine strigulae. Dorsal surface of mesosoma weakly areolate, with coarse costae that are rugose on the pronotum and the propodeum. Femora finely, weakly areolate. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the petiolar spiracle to the caudal cylinder; rugae on lateral faces of the node. Postpetiole uniformly areolate, with rugose sculpture on the dorsal and lateral surfaces. First gastral tergite shining through weak areolate sculpture; without spectral iridescence. First gastral sternite smooth and shining, without spectral iridescence.</paragraph>
<paragraph blockId="355.[498,1542,234,1934]" pageId="355" pageNumber="356">Setae: antennal scapes and funiculi with short, subdecumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about half the width of the compound eye. The head bears ~50, mesosoma ~26, petiole 8, postpetiole ~32, and first gastral tergite ~72 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
<paragraph blockId="355.[498,1542,234,1934]" box="[530,1288,553,580]" pageId="355" pageNumber="356">Color: predominantly testaceous yellow, with legs lighter yellow.</paragraph>
<paragraph blockId="355.[498,1542,234,1934]" pageId="355" pageNumber="356">
<emphasis bold="true" box="[530,1001,593,619]" pageId="355" pageNumber="356">
Gyne measurements &amp; indices (
<emphasis bold="true" box="[919,935,593,619]" italics="true" pageId="355" pageNumber="356">n</emphasis>
= 5):
</emphasis>
SL = 
<geoCoordinate box="[1073,1135,593,620]" degrees="0.518" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.518">0.518</geoCoordinate>
<geoCoordinate box="[1136,1213,593,619]" degrees="0.600" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.6">–0.600</geoCoordinate>
(0.556); FRS = 
<geoCoordinate box="[1400,1462,593,619]" degrees="0.264" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.264">0.264</geoCoordinate>
<geoCoordinate box="[1463,1539,593,619]" degrees="0.309" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.309">–0.309</geoCoordinate>
(0.291); CW = 
<geoCoordinate box="[678,740,633,659]" degrees="0.783" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.783">0.783</geoCoordinate>
<geoCoordinate box="[740,817,633,659]" degrees="0.831" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.831">–0.831</geoCoordinate>
(0.802); CWb = 
<geoCoordinate box="[1020,1082,633,659]" degrees="0.726" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.726">0.726</geoCoordinate>
<geoCoordinate box="[1082,1159,633,659]" degrees="0.782" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.782">–0.782</geoCoordinate>
(0.746); PoOC = 
<geoCoordinate box="[1372,1434,633,659]" degrees="0.304" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.304">0.304</geoCoordinate>
<geoCoordinate box="[1434,1511,633,659]" degrees="0.319" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.319">–0.319</geoCoordinate>
(0.312); CL = 
<geoCoordinate box="[665,727,673,699]" degrees="0.824" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.824">0.824</geoCoordinate>
<geoCoordinate box="[727,804,673,699]" degrees="0.880" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.88">–0.880</geoCoordinate>
(0.851); EL = 
<geoCoordinate box="[977,1039,673,699]" degrees="0.233" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.233">0.233</geoCoordinate>
<geoCoordinate box="[1039,1116,673,699]" degrees="0.277" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.277">–0.277</geoCoordinate>
(0.253); EW = 
<geoCoordinate box="[1302,1364,673,699]" degrees="0.171" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.171">0.171</geoCoordinate>
<geoCoordinate box="[1364,1441,673,699]" degrees="0.207" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.207">–0.207</geoCoordinate>
(0.191); MD = 
<geoCoordinate box="[580,642,712,738]" degrees="0.169" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.169">0.169</geoCoordinate>
<geoCoordinate box="[642,719,712,738]" degrees="0.196" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.196">–0.196</geoCoordinate>
(0.183); WL = 
<geoCoordinate box="[904,966,712,738]" degrees="1.324" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="1.324">1.324</geoCoordinate>
<geoCoordinate box="[966,1043,712,738]" degrees="1.477" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-1.477">–1.477</geoCoordinate>
(1.401); SPST = 
<geoCoordinate box="[1245,1307,712,738]" degrees="0.202" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.202">0.202</geoCoordinate>
<geoCoordinate box="[1308,1385,712,738]" degrees="0.233" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.233">–0.233</geoCoordinate>
(0.223); MPST = 
<geoCoordinate box="[607,669,752,778]" degrees="0.322" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.322">0.322</geoCoordinate>
<geoCoordinate box="[669,746,752,778]" degrees="0.383" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.383">–0.383</geoCoordinate>
(0.354); PEL = 
<geoCoordinate box="[935,997,752,778]" degrees="0.312" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.312">0.312</geoCoordinate>
<geoCoordinate box="[997,1074,752,778]" degrees="0.382" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.382">–0.382</geoCoordinate>
(0.355); NOL = 
<geoCoordinate box="[1274,1336,752,778]" degrees="0.204" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.204">0.204</geoCoordinate>
<geoCoordinate box="[1336,1413,752,778]" degrees="0.228" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.228">–0.228</geoCoordinate>
(0.213); NOH = 
<geoCoordinate box="[598,660,792,819]" degrees="0.145" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.145">0.145</geoCoordinate>
<geoCoordinate box="[660,737,792,818]" degrees="0.168" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.168">–0.168</geoCoordinate>
(0.155); PEH = 
<geoCoordinate box="[933,995,792,818]" degrees="0.300" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.3">0.300</geoCoordinate>
<geoCoordinate box="[995,1072,792,819]" degrees="0.351" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.351">–0.351</geoCoordinate>
(0.319); PPL = 
<geoCoordinate box="[1262,1324,792,819]" degrees="0.215" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.215">0.215</geoCoordinate>
<geoCoordinate box="[1324,1401,792,818]" degrees="0.238" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.238">–0.238</geoCoordinate>
(0.224); PPH = 
<geoCoordinate box="[588,650,832,858]" degrees="0.314" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.314">0.314</geoCoordinate>
<geoCoordinate box="[650,727,832,858]" degrees="0.376" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.376">–0.376</geoCoordinate>
(0.345); PW = 
<geoCoordinate box="[912,974,832,858]" degrees="0.783" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.783">0.783</geoCoordinate>
<geoCoordinate box="[975,1052,832,858]" degrees="0.838" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.838">–0.838</geoCoordinate>
(0.802); SBPA = 
<geoCoordinate box="[1260,1322,832,858]" degrees="0.300" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.3">0.300</geoCoordinate>
<geoCoordinate box="[1322,1399,832,858]" degrees="0.391" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.391">–0.391</geoCoordinate>
(0.338); SPTI = 
<geoCoordinate box="[590,652,872,898]" degrees="0.268" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.268">0.268</geoCoordinate>
<geoCoordinate box="[653,730,872,898]" degrees="0.340" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.34">–0.340</geoCoordinate>
(0.310); PEW = 
<geoCoordinate box="[932,994,872,898]" degrees="0.222" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.222">0.222</geoCoordinate>
<geoCoordinate box="[994,1071,872,898]" degrees="0.269" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.269">–0.269</geoCoordinate>
(0.237); PNW = 
<geoCoordinate box="[1278,1340,872,898]" degrees="0.120" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate box="[1340,1417,872,899]" degrees="0.185" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.185">–0.185</geoCoordinate>
(0.146); PPW = 
<geoCoordinate box="[590,652,912,938]" degrees="0.378" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.378">0.378</geoCoordinate>
<geoCoordinate box="[653,730,911,938]" degrees="0.465" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.465">–0.465</geoCoordinate>
(0.413); HFL = 
<geoCoordinate box="[917,979,911,937]" degrees="0.644" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.644">0.644</geoCoordinate>
<geoCoordinate box="[979,1056,912,938]" degrees="0.718" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.718">–0.718</geoCoordinate>
(0.667); HFWmax = 
<geoCoordinate box="[1306,1368,911,937]" degrees="0.166" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.166">0.166</geoCoordinate>
<geoCoordinate box="[1368,1445,912,938]" degrees="0.189" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.189">–0.189</geoCoordinate>
(0.178); HFWmin = 
<geoCoordinate box="[645,707,951,978]" degrees="0.053" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.053">0.053</geoCoordinate>
<geoCoordinate box="[708,785,951,978]" degrees="0.069" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.069">–0.069</geoCoordinate>
(0.062); CS = 
<geoCoordinate box="[958,1020,952,978]" degrees="1.138" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="1.138">1.138</geoCoordinate>
<geoCoordinate box="[1021,1098,951,978]" degrees="1.216" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-1.216">–1.216</geoCoordinate>
(1.171); ES = 
<geoCoordinate box="[1269,1331,952,978]" degrees="0.319" direction="north" orientation="latitude" pageId="355" pageNumber="356" precision="55" value="0.319">0.319</geoCoordinate>
<geoCoordinate box="[1331,1408,952,978]" degrees="0.381" direction="west" orientation="longitude" pageId="355" pageNumber="356" precision="55" value="-0.381">–0.381</geoCoordinate>
(0.349); SI = 71–77 (75); OI = 28–33 (30); CI = 85–90 (88); WLI = 182–200 (188); SBI = 41–52 (45); PSI = 15–17 (16); PWI = 165–186 (174); PLI = 145–176 (158); NI = 121–146 (138); PNWI = 52–79 (62); NLI = 53–67 (60); FI = 242–313 (289).
</paragraph>
<paragraph blockId="355.[498,1542,234,1934]" pageId="355" pageNumber="356">
<emphasis bold="true" box="[530,747,1111,1137]" pageId="355" pageNumber="356">Gyne description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 85–90). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about two times the maximum width of the scape (SI 71–77). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two and a half times the maximum width of the antennal scape. Compound eyes strongly protruding past the lateral margins of the head capsule. Lateral margins of head convex behind the compound eyes, but parallel to each other between the compound eyes and the mandibular insertions. Posterior head margin weakly concave medially, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="355.[498,1542,234,1934]" lastBlockId="356.[498,1542,230,1934]" lastPageId="356" lastPageNumber="357" pageId="355" pageNumber="356">
In profile view, compound eyes ovular and large (OI 28–33), with 17 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum on the same plane as the mesoscutum and flat dorsally. Posterior margin of metanotum extending slightly past the posterior margin of the mesoscutum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout, weakly developed, and short (PSI 15–17), about a third as long as the propodeal declivity, flared at the base, triangular, and blunt. Propodeal declivity straight and flat, forming a rounded ~120 
<emphasis box="[789,799,230,248]" italics="true" pageId="356" pageNumber="357">
<superScript attach="left" box="[789,799,230,248]" fontSize="8" pageId="356" pageNumber="357">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 145–176), with tubercles anterodorsally. Subpetiolar process in the form of a blunt, triangular tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle short: comprising about a quarter of the total petiole length. Petiolar node truncate but dorsum is extremely angled, so that in many specimens it appears cuneiform: anterior face forming a sharp ~90 
<emphasis box="[616,626,509,527]" italics="true" pageId="356" pageNumber="357">
<superScript attach="left" box="[616,626,509,527]" fontSize="8" pageId="356" pageNumber="357">°</superScript>
</emphasis>
angle with the dorsal face, which is flat; dorsal face meeting the posterior face at a sharp ~120 
<emphasis box="[682,692,548,566]" italics="true" pageId="356" pageNumber="357">
<superScript attach="left" box="[682,692,548,566]" fontSize="8" pageId="356" pageNumber="357">°</superScript>
</emphasis>
angle; very short posterior face forms a ~110 
<emphasis box="[1232,1242,548,566]" italics="true" pageId="356" pageNumber="357">
<superScript attach="left" box="[1232,1242,548,566]" fontSize="8" pageId="356" pageNumber="357">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly convex anterodorsally, flat posterodorsally; weakly lobed ventrally.
</paragraph>
<paragraph blockId="356.[498,1542,230,1934]" pageId="356" pageNumber="357">
In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as rounded sclerites. Propodeal spines weakly diverging apically, their apices separated from each other by about three times their length. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node, when viewed posterodorsally, tapering apically so the apex is about half the width of the base; dorsal margin emarginate medially. Postpetiole narrow (PWI 165–186), anteroposteriorly compressed, and subquadrate. Anterior margin of postpetiole weakly convex but emarginated medially; anterior corners marked by rounded ~100 
<emphasis box="[994,1004,947,965]" italics="true" pageId="356" pageNumber="357">
<superScript attach="left" box="[994,1004,947,965]" fontSize="8" pageId="356" pageNumber="357">°</superScript>
</emphasis>
angles as it transitions to the lateral margins; lateral margins parallel to each other; posterior corners rounded; posterior margin weakly concave. Metafemur weakly to strongly incrassate (FI 242–313).
</paragraph>
<paragraph blockId="356.[498,1542,230,1934]" pageId="356" pageNumber="357">Sculpture: median clypeal carina present, extending from the anterior margin nearly to frontal triangle, and flanked on either side by three equally strong carinae. Lateral clypeal lobes with additional weaker carinae; ground sculpture weakly areolate. Antennal scapes finely areolate. Cephalic dorsum areolate, with costae over the ground sculpture, which become rugose on the posterior sixth of the head; concentric rugulae surrounding the antennal insertions. Lateral surfaces of head areolate, with coarse rugae over the ground sculpture. Ventral surface of head smooth and shining medially, but otherwise weakly areolate with weak costulae. Pronotal neck areolate. Anterior face of pronotum weakly areolate. Lateral surfaces of the mesosoma sculptured similarly to the dorsum of the head. Propodeal declivity areolate-strigulate. Mesoscutum and mesoscutellum with fine costulae over finely areolate ground sculpture. Metanotum finely areolate. Propodeum areolate, with coarse costae. Femora shining through weak areolate sculpture. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the petiolar spiracle to the caudal cylinder; coarse rugae on lateral faces of the node. Postpetiole uniformly areolate, with weak rugose sculpture on the dorsal and lateral faces. First gastral tergite shining through weak areolate sculpture; without spectral iridescence. First gastral sternite smooth and shining.</paragraph>
<paragraph blockId="356.[498,1542,230,1934]" pageId="356" pageNumber="357">Setae: antennal scapes and funiculi with short, subdecumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about a third of the width of the compound eye. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
<paragraph blockId="357.[498,1542,234,1098]" pageId="357" pageNumber="358">Color: predominantly testaceous yellow, with legs lighter yellow. Wing bases dark brown.</paragraph>
</subSubSection>
<subSubSection pageId="357" pageNumber="358" type="etymology">
<paragraph blockId="357.[498,1542,234,1098]" pageId="357" pageNumber="358">
<emphasis bold="true" box="[530,660,314,340]" pageId="357" pageNumber="358">Etymology</emphasis>
: Morphological, from the Latin ‘rugosus’ (= wrinkled), in reference to the rugose head and mesosoma (
<bibRefCitation author="Mackay WP" box="[842,1004,354,380]" journalOrPublisher="Sociobiology" pageId="357" pageNumber="358" pagination="265 - 444" part="36" refId="ref271737" refString="Mackay WP. 2000. A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265 - 444." title="A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae)" type="journal article" year="2000">
<emphasis box="[842,1004,354,380]" italics="true" pageId="357" pageNumber="358">Mackay, 2000</emphasis>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="357" pageNumber="358" type="discussion">
<paragraph blockId="357.[498,1542,234,1098]" pageId="357" pageNumber="358">
<emphasis bold="true" box="[530,670,393,419]" pageId="357" pageNumber="358">Comments:</emphasis>
From 
<bibRefCitation author="Mackay WP" box="[752,926,394,420]" journalOrPublisher="Sociobiology" pageId="357" pageNumber="358" pagination="265 - 444" part="36" refId="ref271737" refString="Mackay WP. 2000. A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265 - 444." title="A review of the new world ants of the subgenus Myrafant (genus Leptothorax) (Hymenoptera: Formicidae)" type="journal article" year="2000">
<emphasis box="[752,926,394,420]" italics="true" pageId="357" pageNumber="358">Mackay (2000)</emphasis>
</bibRefCitation>
: “The 
<typeStatus box="[1007,1057,394,420]" pageId="357" pageNumber="358">type</typeStatus>
series was collected in a trap baited with Vienna sausage, placed in oak trees, about 2-meter height. The habitat was a very steep, south-facing slope of a mountain, covered with oak trees. The specimens were collected within 20 meters distant from one another.” After inspecting the collections of the LACM, I found that W.S. Creighton collected 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1046,1299,553,579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[1046,1299,553,579]" italics="true" pageId="357" pageNumber="358">Temnothorax rugosus</emphasis>
</taxonomicName>
in a number of localities spanning northern 
<collectingCountry box="[837,923,593,619]" name="Mexico" pageId="357" pageNumber="358">Mexico</collectingCountry>
and southwestern Texas, in addition to the 
<typeStatus box="[1446,1496,593,619]" pageId="357" pageNumber="358">type</typeStatus>
locality near Monterrey. Like members of the 
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1038,1134,633,658]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1038,1134,633,658]" italics="true" pageId="357" pageNumber="358">annexus</emphasis>
</taxonomicName>
group, which it is closely affiliated with, 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[565,685,673,698]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[565,685,673,698]" italics="true" pageId="357" pageNumber="358">T. rugosus</emphasis>
</taxonomicName>
nests arboreally in the branches a number of different 
<taxonomicName box="[1338,1489,673,699]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1338,1433,673,699]" italics="true" pageId="357" pageNumber="358">Quercus</emphasis>
spp.
</taxonomicName>
(including 
<emphasis box="[627,1000,712,739]" italics="true" pageId="357" pageNumber="358">
<taxonomicName box="[627,741,712,739]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="canbyi">Q. canbyi</taxonomicName>
, 
<taxonomicName authorityName="Trelease &amp; C.H.Muller" authorityYear="1936" box="[754,885,712,739]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="clivicola">Q. clivicola</taxonomicName>
, 
<taxonomicName authorityName="Liebmann" authorityYear="1854" box="[898,1000,713,739]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="grisea">Q. grisea</taxonomicName>
</emphasis>
, and 
<taxonomicName box="[1068,1242,712,739]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="polymorpha">
<emphasis box="[1068,1242,712,739]" italics="true" pageId="357" pageNumber="358">Q. polymorpha</emphasis>
</taxonomicName>
), as well as 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Cercis" kingdom="Plantae" order="Fabales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="reniformis">
<emphasis italics="true" pageId="357" pageNumber="358">Cercis reniformis</emphasis>
</taxonomicName>
and 
<taxonomicName box="[674,786,752,779]" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" kingdom="Plantae" order="Sapindales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[674,731,752,778]" italics="true" pageId="357" pageNumber="358">Rhus</emphasis>
spp.
</taxonomicName>
at mid elevations. 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1003,1254,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[1003,1254,752,778]" italics="true" pageId="357" pageNumber="358">Temnothorax rugosus</emphasis>
</taxonomicName>
is also recorded as being collected from 
<taxonomicName authorityName="Matuda" authorityYear="1973" box="[675,949,792,818]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="357" pageNumber="358" phylum="Tracheophyta" rank="species" species="carlos-hankii">
<emphasis box="[675,949,792,818]" italics="true" pageId="357" pageNumber="358">Tillandsia carlos-hankii</emphasis>
</taxonomicName>
in oak-pine forest near Ixtepeji, 
<collectingRegion box="[1338,1424,792,818]" country="Mexico" name="Oaxaca" pageId="357" pageNumber="358">Oaxaca</collectingRegion>
(
<bibRefCitation author="Franco Mendez AD" journalOrPublisher="Masters thesis, Instituto Politecnico Nacional, CIIDIR, Unidad Oaxaca" pageId="357" pageNumber="358" refId="ref271092" refString="Franco Mendez AD. 2008. Diversidad de macroartropodos en Tillandsia carlos-hankii Matuda y Tillandsia oaxacana L. B. Smith en un bosque de encino pino de Oaxaca. Masters thesis, Instituto Politecnico Nacional, CIIDIR, Unidad Oaxaca." title="Diversidad de macroartropodos en Tillandsia carlos-hankii Matuda y Tillandsia oaxacana L. B. Smith en un bosque de encino pino de Oaxaca" type="book" year="2008">
<emphasis italics="true" pageId="357" pageNumber="358">Franco Méndez, 2008</emphasis>
</bibRefCitation>
). The femora of 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[852,1102,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[852,1102,832,858]" italics="true" pageId="357" pageNumber="358">Temnothorax rugosus</emphasis>
</taxonomicName>
, as noted above, tend to be incrassate, but the degree of swollenness appears to scale with the size of the worker, with smaller workers having less incrassate femora. 
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[957,1210,912,938]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="357" pageNumber="358" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[957,1210,912,938]" italics="true" pageId="357" pageNumber="358">Temnothorax rugosus</emphasis>
</taxonomicName>
shows some variability in petiolar node shape, eye size, and sculpture among populations (
<figureCitation box="[1252,1448,951,978]" captionStart="Figure 152" captionStartId="350.[524,589,723,745]" captionTargetBox="[498,1541,234,710]" captionTargetId="figure-268@350.[498,1534,234,695]" captionTargetPageId="350" captionText="Figure 152 Temnothorax rugosus. (A–C) Worker (LACMENT323296). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323337). (D) Full-face view. (E) Profile view. (F) Dorsal view. Morphological variant, Texas, U.S.A. (G–I) worker (LACMENT323260). (G) Full-face view. (H) Profile view. (I) Dorsal view. (J–L) Gyne (LACMENT323259). (J) Full-face view. (K) Profile view. (L) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-152" figureDoi="http://doi.org/10.5281/zenodo.5102447" httpUri="https://zenodo.org/record/5102447/files/figure.png" pageId="357" pageNumber="358">Figs. 152G–152L</figureCitation>
). In the petiolar node, there appears to be a complete range of variation between completely cuneiform, with no differentiation between the dorsal and posterior declivitous face, to subcuneiform, with a very short but distinct posterior face and a deeply sloping dorsal face.
</paragraph>
</subSubSection>
</treatment>
</document>