434 lines
78 KiB
XML
434 lines
78 KiB
XML
<document id="BE69C5AE5B17CF1E92B4F7FFFCC8C197" ID-DOI="10.26879/1209" ID-ISSN="1094-8074" ID-Zenodo-Dep="11063294" ID-ZooBank="1726FDAE-2EE5-4145-A124-6D24287C0514" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1714014612185" checkinUser="felipe" docAuthor="Zazhigin, Vladimir S. & Voyta, Leonid L." docDate="2022" docId="71755174FFA70336CE0DFD79CA83FD83" docLanguage="en" docName="PalaeontolElectron.25.3.a29.pdf" docOrigin="Palaeontologia Electronica (a 29) 25 (3)" docSource="http://dx.doi.org/10.26879/1209" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Paranourosorex intermedius Zazhigin & Voyta 2022" docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="8D4C290CFFAB0339CF1EFFE8C819FF81" masterDocTitle="New Neogene anourosoricin shrews from northern Asia" masterLastPageNumber="36" masterPageNumber="1" pageNumber="13" updateTime="1714663430529" updateUser="julia" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="54512F19C6EC59F8BE2C255AB014189C">
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<mods:title id="38F6C7D2256A912702992F3C121308E3">New Neogene anourosoricin shrews from northern Asia</mods:title>
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<mods:namePart id="8C928B5026D811E43BE4E51BA1B6BDEE">Zazhigin, Vladimir S.</mods:namePart>
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<mods:namePart id="C87F9D0908B81AA52363A9179F20B566">Voyta, Leonid L.</mods:namePart>
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<mods:title id="42C713EE6A7B96A9E1F928C712EFE1BA">Palaeontologia Electronica</mods:title>
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<mods:part id="DB8FB3616E46B4897022E54B086723B7">
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<mods:date id="34A50CBE7BF681B00438A1A773D364DC">2022</mods:date>
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<mods:title id="5E18B5F908DD2ADFFA6477F8E260CAAF">a 29</mods:title>
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<mods:number id="E4BEC5ED1DEAEFE252C432B4CF387C31">25</mods:number>
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<mods:number id="BB9F0E01B44C5D037EB371A4825CC49C">3</mods:number>
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<mods:url id="D44E86E1DF51A1F08F1785452C23F559">http://dx.doi.org/10.26879/1209</mods:url>
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<mods:identifier id="44DC63449024CC1FCF0D2DA09BCB24B2" type="DOI">10.26879/1209</mods:identifier>
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<mods:identifier id="58951DA187FB0FDD83BF20BEA3208639" type="ISSN">1094-8074</mods:identifier>
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<mods:identifier id="167568A32EFB33FC68B4B3F7BB31B8F9" type="Zenodo-Dep">11063294</mods:identifier>
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<treatment id="71755174FFA70336CE0DFD79CA83FD83" LSID="urn:lsid:zoobank.org:act:FA6D006C-2C2A-4860-BE5A-BBAD043A2D49" httpUri="http://treatment.plazi.org/id/71755174FFA70336CE0DFD79CA83FD83" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
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<subSubSection id="B1C6B3E9FFA70335CE0DFD79CADCFD2B" box="[275,709,657,682]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="F963E062FFA70335CE0DFD79CADCFD2B" blockId="12.[239,745,657,714]" box="[275,709,657,682]" pageId="12" pageNumber="13">
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<heading id="A22B570EFFA70335CE0DFD79CADCFD2B" box="[275,709,657,682]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="7">
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<taxonomicName id="3EDC9BE1FFA70335CE0DFD79CA7BFD2B" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[275,610,657,682]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
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<emphasis id="CBA83C70FFA70335CE0DFD79CA7BFD2B" box="[275,610,657,682]" italics="true" pageId="12" pageNumber="13">Paranourosorex intermedius</emphasis>
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</taxonomicName>
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sp. nov.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1C6B3E9FFA70335CFF1FD59CA0FFCBB" pageId="12" pageNumber="13" type="description">
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<paragraph id="F963E062FFA70335CFF1FD59CAF0FD4B" blockId="12.[239,745,657,714]" box="[239,745,688,714]" pageId="12" pageNumber="13">
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<figureCitation id="61E7FCE7FFA70335CFF1FD59C96DFD4B" box="[239,372,689,714]" captionStart="FIGURE 6" captionStartId="11.[196,287,1080,1103]" captionTargetBox="[190,1441,166,1060]" captionTargetId="figure-235@11.[190,1441,166,1060]" captionTargetPageId="11" captionText="FIGURE 6. Teeth and bones of Paranourosorex seletiensis Storch and Zazhigin, 1996 (A, B), Crusafontina sp. 1, (C), Crusafontina sp. 2 (D) and Paranourosorex intermedius sp. nov. (E). A, GIN 951/1000 (holotype) from SLT/1A, left hemimandible fragment (cut image) with m1 and m2 in occlusal view (A2, magnified m2); B, GIN 951/1001 (paratype) from SLT/1A, left P4 in occlusal view; C, GIN 640/1004 from PVL/1A, left M1 in occlusal view; D, GIN 951/1003 SLT/ 1A, left P4 in occlusal view; E, GIN 948/1051 (holotype) from NST/1A, right dentary fragment with m1 and m2 (E1, occlusal view; E2, lateral view E3, medial view;). Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063306" httpUri="https://zenodo.org/record/11063306/files/figure.png" pageId="12" pageNumber="13">Figures 6E</figureCitation>
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,
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<figureCitation id="61E7FCE7FFA70335CE61FD59CAF0FD4B" box="[383,745,688,714]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="12" pageNumber="13">7B, C–D, F–G, I, K, M, O, Q–R</figureCitation>
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</paragraph>
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<paragraph id="F963E062FFA70335CFDEFD33CB14FD6E" blockId="12.[192,781,731,751]" box="[192,781,731,751]" pageId="12" pageNumber="13">
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<emphasis id="CBA83C70FFA70335CFDEFD33CB14FD6E" bold="true" box="[192,781,731,751]" pageId="12" pageNumber="13">
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zoobank.org/
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<uuid id="8D7ADAB7FFA70335CE56FD33CB14FD6E" box="[328,781,731,751]" pageId="12" pageNumber="13">FA6D006C-2C2A-4860-BE5A-BBAD043A2D49</uuid>
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</emphasis>
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</paragraph>
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<paragraph id="F963E062FFA70335CFDEFCE9CA0FFCBB" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
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1996
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<taxonomicName id="3EDC9BE1FFA70335CE3EFCE9CA05FC9B" box="[288,540,769,794]" class="Mammalia" family="Soricidae" genus="Paranourosorex" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="undefined-2">
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<emphasis id="CBA83C70FFA70335CE3EFCE9C9C7FC9B" box="[288,478,769,794]" italics="true" pageId="12" pageNumber="13">Paranourosorex</emphasis>
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sp. 2
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</taxonomicName>
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Storch and Zazhigin, p. 264, figs. 3c, 4l–o.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B1C6B3E9FFA70335CFDEFCAECA7EF95E" pageId="12" pageNumber="13" type="materials_examined">
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||
<paragraph id="F963E062FFA70335CFDEFCAEC999FC41" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
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<emphasis id="CBA83C70FFA70335CFDEFCAEC970FCDE" bold="true" box="[192,361,838,863]" pageId="12" pageNumber="13">
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<typeStatus id="26675EC0FFA70335CFDEFCAEC8E5FCDE" box="[192,252,838,863]" pageId="12" pageNumber="13">Type</typeStatus>
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locality.
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</emphasis>
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<location id="FC03B6B9FFA70335CE6AFCAECA21FCE1" LSID="urn:lsid:plazi:treatment:71755174FFA70336CE0DFD79CA83FD83:FC03B6B9FFA70335CE6AFCAECA21FCE1" box="[372,568,838,864]" country="Russia" county="Omskaya Oblast'" latitude="54.833332" longLatPrecision="1068" longitude="73.4" municipality="Novaya Stanitsa village" name="Novaya Stanitsa" pageId="12" pageNumber="13">Novaya Stanitsa</location>
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1A, right slope of the
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<location id="FC03B6B9FFA70335CFF2FC8EC9D8FCFE" LSID="urn:lsid:plazi:treatment:71755174FFA70336CE0DFD79CA83FD83:FC03B6B9FFA70335CFF2FC8EC9D8FCFE" box="[236,449,870,895]" country="Russia" county="Omskaya Oblast'" latitude="54.833332" longLatPrecision="1068" longitude="73.4" municipality="Novaya Stanitsa village" name="Irtysh River Valley" pageId="12" pageNumber="13">Irtysh River Valley</location>
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near
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<collectingMunicipality id="19077A18FFA70335CD1BFC8ECB0EFCFE" box="[517,791,870,896]" pageId="12" pageNumber="13">Novaya Stanitsa village</collectingMunicipality>
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(ca.
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<geoCoordinate id="9CE886A5FFA70335CFE4FC6EC940FC1E" box="[250,345,902,927]" degrees="54" direction="north" minutes="50" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="54.833332">N54°50'</geoCoordinate>
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<geoCoordinate id="9CE886A5FFA70335CE78FC6EC9D0FC1E" box="[358,457,902,927]" degrees="73" direction="east" minutes="24" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="73.4">E73°24'</geoCoordinate>
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),
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<collectingCounty id="100298EEFFA70335CEFEFC6ECAACFC21" box="[480,693,902,928]" pageId="12" pageNumber="13">Omskaya Oblast’</collectingCounty>
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,
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<collectingCountry id="81CBA0F2FFA70335CDD8FC6ECB0EFC1E" box="[710,791,902,927]" name="Russia" pageId="12" pageNumber="13">Russia</collectingCountry>
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(
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<figureCitation id="61E7FCE7FFA70335CFD8FC4EC935FC3E" box="[198,300,934,959]" captionStart="FIGURE 1" captionStartId="2.[194,285,1060,1083]" captionTargetBox="[196,1434,169,1028]" captionTargetId="figure-256@2.[193,1438,164,1031]" captionTargetPageId="2" captionText="FIGURE 1. Map of Asia (A) showing the location of Russian and Kazakh late Miocene and early Pliocene sites (B): ANR, Andreevka 1A, 2A; BRK, Borki 1A–C; BTK, Biteke; CHR, Cherlak 1A; ISK, Isakovka 1A, 2A; KRN, Krasnokutsk; KRT, Kartashovo; LZN, Lezhanka 1A, 2B; NIL, Nizhneil’inka; NST, Novaya Stanitsa 1A–B; PPL, Petropavlovsk 1A; PSH, Peshnevo 1A–B; PVL, Pavlodar 1A, 2 ‘Quarry’; SLT, Selety 1A (type locality of Paranourosorex seletiensis Storch and Zazhigin, 1996). Age of the deposits is marked by colours, which corresponds to ELMA Neogene zones ‘MN 10, 12–14’; the redeposited fossiliferous layers are marked by ‘?’. Map data from resource ESRI (http:// www.esri.com/) using SASPlanet software (v.160707.9476)." figureDoi="http://doi.org/10.5281/zenodo.11063296" httpUri="https://zenodo.org/record/11063296/files/figure.png" pageId="12" pageNumber="13">Figure 1</figureCitation>
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: NST).
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</paragraph>
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<paragraph id="F963E062FFA70335CFDEFC2ECA25FBBE" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
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<emphasis id="CBA83C70FFA70335CFDEFC2EC962FC5E" bold="true" box="[192,379,966,991]" pageId="12" pageNumber="13">
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<typeStatus id="26675EC0FFA70335CFDEFC2EC8E5FC5E" box="[192,252,966,991]" pageId="12" pageNumber="13">Type</typeStatus>
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horizon.
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</emphasis>
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Stratotypic locality of Novaya Stanitsa Formation (
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<emphasis id="CBA83C70FFA70335CEA5FC0EC9C6FC7E" box="[443,479,998,1023]" italics="true" pageId="12" pageNumber="13">nst</emphasis>
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), late Miocene (Turolian, MN 13). Material was collected by VZ during fieldworks of 1980s, 2000 and 2001.
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</paragraph>
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<paragraph id="F963E062FFA70335CFDEFBAECA19F91E" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
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<emphasis id="CBA83C70FFA70335CFDEFBAEC96EFBDE" bold="true" box="[192,375,1094,1119]" pageId="12" pageNumber="13">Type material.</emphasis>
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<materialsCitation id="49B4EA3FFFA70335CE9BFBAEC9A5FB1E" pageId="12" pageNumber="13">
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<typeStatus id="26675EC0FFA70335CE9BFBAEC9EAFBDE" box="[389,499,1094,1119]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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: GIN 948/1051 — right dentary fragment with moderately worn p4 and heavily worn m1–m3.
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</materialsCitation>
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<materialsCitation id="49B4EA3FFFA70335CEDDFB6ECB0CFB5E" pageId="12" pageNumber="13">
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<typeStatus id="26675EC0FFA70335CEDDFB6ECA26FB1E" box="[451,575,1158,1183]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
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: (n =12): GIN 948/ 1052 — left dentary fragment with i1, m1 and alveoli of a1–p4 (i1 heavily and m1 moderately worn)
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</materialsCitation>
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;
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<materialsCitation id="49B4EA3FFFA70335CFDEFB0ECB00FB7E" box="[192,793,1254,1280]" pageId="12" pageNumber="13">GIN 948/1056 — isolated right m1 (slightly worn);</materialsCitation>
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<materialsCitation id="49B4EA3FFFA70335CFDEFAEECB00FA9E" box="[192,793,1286,1312]" pageId="12" pageNumber="13">GIN 948/1063 — isolated left m2 (slightly worn);</materialsCitation>
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<materialsCitation id="49B4EA3FFFA70335CFDEFACEC91CFAE1" pageId="12" pageNumber="13">GIN 948/1064 — isolated right m3 (slightly worn)</materialsCitation>
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;
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<materialsCitation id="49B4EA3FFFA70335CE16FAAEC912FAFE" pageId="12" pageNumber="13">GIN 948/1065 — isolated left P4 (slightly worn);</materialsCitation>
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<materialsCitation id="49B4EA3FFFA70335CE3BFA8ECAB8FA1E" pageId="12" pageNumber="13">GIN 948/1069 — isolated right M1 (metacone damaged; moderately worn);</materialsCitation>
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<materialsCitation id="49B4EA3FFFA70335CDB5FA6ECA08FA5E" pageId="12" pageNumber="13">GIN 948/ 1070 — isolated right M1 (paracone and metacone damaged; moderately worn)</materialsCitation>
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;
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<materialsCitation id="49B4EA3FFFA70335CD02FA2ECA1BFA7E" pageId="12" pageNumber="13">GIN 948/1073 — isolated left I1 (slightly worn)</materialsCitation>
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||
;
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||
<materialsCitation id="49B4EA3FFFA70335CD0FFA0EC912F9BE" pageId="12" pageNumber="13">GIN 948/1075 — isolated right i1 (tip of root is broken off; moderately worn);</materialsCitation>
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||
<materialsCitation id="49B4EA3FFFA70335CE09F9CEC96AF9DE" pageId="12" pageNumber="13">GIN 948/1078 — isolated left p4 (without roots; unworn)</materialsCitation>
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;
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<materialsCitation id="49B4EA3FFFA70335CE9DF9AECA0AF9FE" pageId="12" pageNumber="13">GIN 948/1081 — isolated left a1 (slightly damaged and worn)</materialsCitation>
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||
;
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<materialsCitation id="49B4EA3FFFA70335CD00F98ECA19F91E" pageId="12" pageNumber="13">GIN 948/1089 — isolated left A1 (slightly worn).</materialsCitation>
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</paragraph>
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||
<paragraph id="F963E062FFA70335CFDEF94ECA7EF95E" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CFDEF94EC9BEF93E" bold="true" box="[192,423,1702,1727]" pageId="12" pageNumber="13">Location of types.</emphasis>
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||
<typeStatus id="26675EC0FFA70335CEAEF94ECA00F93E" box="[432,537,1702,1727]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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||
and
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||
<typeStatus id="26675EC0FFA70335CD46F94ECAD4F93E" box="[600,717,1702,1727]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
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||
in the collection of GIN,
|
||
<collectingRegion id="3B182E80FFA70335CE8CF92EC9EDF95E" box="[402,500,1734,1759]" country="Russia" name="Moscow" pageId="12" pageNumber="13">Moscow</collectingRegion>
|
||
(
|
||
<collectingCountry id="81CBA0F2FFA70335CD1CF92ECA45F95E" box="[514,604,1734,1759]" name="Russia" pageId="12" pageNumber="13">Russia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA70335CFDEF90EC909F8BE" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="F963E062FFA70335CFDEF90EC909F8BE" blockId="12.[192,793,768,1855]" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CFDEF90EC957F97E" bold="true" box="[192,334,1766,1791]" pageId="12" pageNumber="13">Etymology.</emphasis>
|
||
From the Latin
|
||
<taxonomicName id="3EDC9BE1FFA70335CD17F90ECA8AF97E" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[521,659,1766,1791]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="CBA83C70FFA70335CD17F90ECA8AF97E" box="[521,659,1766,1791]" italics="true" pageId="12" pageNumber="13">intermedius</emphasis>
|
||
</taxonomicName>
|
||
, intermediate (between smallest
|
||
<taxonomicName id="3EDC9BE1FFA70335CECEF8EECA73F89E" authorityName="Storch and Zazhigin" authorityYear="1996" box="[464,618,1798,1823]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="seletiensis">
|
||
<emphasis id="CBA83C70FFA70335CECEF8EECA73F89E" box="[464,618,1798,1823]" italics="true" pageId="12" pageNumber="13">P. seletiensis</emphasis>
|
||
</taxonomicName>
|
||
and largest
|
||
<taxonomicName id="3EDC9BE1FFA70335CC1CF8EEC919F8BE" authorityName="Rzebik-Kowalska" authorityYear="1975" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA70335CC1CF8EEC919F8BE" italics="true" pageId="12" pageNumber="13">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA70335CC56FF4ECD66FEBE" pageId="12" pageNumber="13" type="materials_examined">
|
||
<paragraph id="F963E062FFA70335CC56FF4ECD66FEBE" blockId="12.[840,1441,166,1887]" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CC56FF4ECBADFF3E" bold="true" box="[840,948,166,191]" pageId="12" pageNumber="13">Material.</emphasis>
|
||
<typeStatus id="26675EC0FFA70335CCA0FF4ECBE1FF3E" box="[958,1016,166,191]" pageId="12" pageNumber="13">Type</typeStatus>
|
||
material and 27 remains from
|
||
<typeStatus id="26675EC0FFA70335CA73FF4ECD86FF3E" box="[1389,1439,166,191]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality (NST/1A), 60 remains from other 8 localities: NST/1B (n = 1), BRK/1A (n = 26), BRK/1B (n = 18), LZN/2B (n = 8), PVL/2 (n = 1), ISK/1A (n = 1), ISK/2A (n = 3), KRT (n = 1) (see Appendix 2: 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA70335CC56FEAECCCDFEDE" box="[840,1236,326,352]" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="F963E062FFA70335CC56FEAECCCDFEDE" blockId="12.[840,1441,166,1887]" box="[840,1236,326,352]" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CC56FEAECC10FEDE" bold="true" box="[840,1033,326,351]" pageId="12" pageNumber="13">Measurements.</emphasis>
|
||
See Appendix 4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA70335CC56FE8ECC1CFCBE" pageId="12" pageNumber="13" type="diagnosis">
|
||
<paragraph id="F963E062FFA70335CC56FE8ECC1CFCBE" blockId="12.[840,1441,166,1887]" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CC56FE8ECBD7FEFE" bold="true" box="[840,974,358,383]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
|
||
Large-sized
|
||
<taxonomicName id="3EDC9BE1FFA70335CB6AFE8ECDB8FEFE" box="[1140,1441,358,383]" class="Mammalia" family="Soricidae" genus="Paranourosorex" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="CBA83C70FFA70335CB6AFE8ECD28FEFE" box="[1140,1329,358,383]" italics="true" pageId="12" pageNumber="13">Paranourosorex</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Lower teeth, mostly p4 and m1, are inflated. Upper antemolar row consists of three teeth with vestigial A3. P4 has trapeziform shape with a deep anterior notch and very shallow the posterior emargination. M1 has subquadrate shape with the expressed Wshaped buccal crests, moderately expressed short metaloph without the metacone base contact. p4 is moderately exaenodont, bulbous-like in shape with the spot-shaped wear facet; the posterolingual basin of p4 is very weak. The entocristid of m1 is separated from the metaconid base by a distinctly marked transverse groove; the hypolohid is low; the small entostylid is weakly separated from the entoconid base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA70337CC56FCAECD8CFB3E" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="F963E062FFA70337CC56FCAEC960FA1E" blockId="12.[840,1441,166,1887]" lastBlockId="14.[192,793,166,1888]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
|
||
<emphasis id="CBA83C70FFA70335CC56FCAECBFBFCDE" bold="true" box="[840,994,838,863]" pageId="12" pageNumber="13">Description.</emphasis>
|
||
I1 crown proportional in size to the root; the talon has an oblique tilted cutting edge without any notch in the base; the ectocingulum is moderately developed and rises up to the middle of the crown (
|
||
<figureCitation id="61E7FCE7FFA70335CCCEFC2ECC57FC5E" box="[976,1102,966,991]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="12" pageNumber="13">Figure 7B</figureCitation>
|
||
). The upper antemolars are dramatically decreased in size from the largest A1 to middle-sized A2 (less than 1/2 of A1) and vestigial A3. The addition of cingular cusps (usually one cusp) on the lingual part of the crown is developed on A1; the entocingulum of A2 is smooth. A1 and A2 have developed both anterior and central crests; A1 displays a wrinkled enamel (
|
||
<figureCitation id="61E7FCE7FFA70335CA0BFB4ECD8CFB3E" box="[1301,1429,1190,1215]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="12" pageNumber="13">Figure 7M</figureCitation>
|
||
). P4 has trapeziform crown, developed anterior notch and the shallow posterior emargination. The parastyle is separated from the paracone base by a shallow groove. P4 has a short ectocingulum, well-developed postcingulum and wrinkled enamel on the paracone buccal side (
|
||
<figureCitation id="61E7FCE7FFA70335CBC9FA8ECD44FA01" box="[1239,1373,1382,1408]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="12" pageNumber="13">Figure 7C</figureCitation>
|
||
); the entocingulum is absent. M1 has subquadrate occlusal shape with the expressed W-shaped line of the buccal crests. The metaconule is absent; the metaloph is short and separated from the base of the metacone (
|
||
<figureCitation id="61E7FCE7FFA70335CCE8F9EECC6BF9A1" box="[1014,1138,1542,1568]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="12" pageNumber="13">Figure 7G</figureCitation>
|
||
). The posterior emargination of M1 is shallow. M2 has well-developed cones; the parastyle is longest, the mesostyle is buccally prominent, the metastyle undeveloped; the hypocone round in the base with a spot-shaped wear facet. M3 unknown. i1 has a smooth cutting edge and distinctly upturned tip. The ectocingulid of i1 is undeveloped. The medial groove of the root is short and did not overreach the crown-root junction. a1 has a bulbous-like crown and massive root; the crown with well-developed ecto- and entocingulids, spot-shaped wear facet and short anterior crest. The central crests of a1 is undeveloped. The inflated p4 bears the spot-shaped wear facet, extremely weak developed the crown structures and distinctly wrinkled enamel (
|
||
<figureCitation id="61E7FCE7FFA50337CD59FECECACFFEC1" box="[583,726,294,320]" captionStart="FIGURE 4" captionStartId="7.[196,287,1008,1031]" captionTargetBox="[199,1432,178,975]" captionTargetId="figure-328@7.[190,1439,168,984]" captionTargetPageId="7" captionText="FIGURE 4. Schematic image of the fourth lower premolar of Ishimosorex ishimiensis gen. et sp. nov. (A, GIN 952/ 1153, PPL/1A), Crusafontina kormosi (B, Polgárdi 4), Paranourosorex intermedius sp. nov. (C, GIN 948/1051, NST/ 1A), Paranourosorex gigas (D, GIN 1118/1011, PSH/1B), Anourosorex squamipes (E, ZIN 98253, Recent), Crusafontina fastigata (F, AG5A from Los Aguanaces 5A, Teruel Basin, Spain by van Dam, 2004: figs. 4-1), Amblycoptus oligodon (G, Polgárdi 4) and Amblycoptus jessiae (H, KS 3157 from Las Casiones, Teruel Basin, Spain by van Dam, 2004: figs. 5-11). Abbreviations: a, concavity; add, additional crown elements (1, crest; 2, cusplet); ccr, central crest; eccd, ectocingulid; encd, entocingulid; pcd, postcingulid; plb, posterolingual basin (1, shallow, weak; 2, expressed); wen, wrinkled enamel; wft, wear facet (1, spot-like; 2, comma-like). Star indicates reversed image. Scale bars are 1 mm; F, H unscaled." figureDoi="http://doi.org/10.5281/zenodo.11063302" httpUri="https://zenodo.org/record/11063302/files/figure.png" pageId="14" pageNumber="15">Figures 4C</figureCitation>
|
||
,
|
||
<figureCitation id="61E7FCE7FFA50337CDF9FECECB14FEBE" box="[743,781,294,319]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">7K</figureCitation>
|
||
). m1 is relatively short and wide and so seems slightly inflated. The enamel surface of the trigonid buccal side is slightly wrinkled. The protoconid and metaconid of m1 are brought closer, the hypoconid and entoconid are widely separated. The hypoflexid of m1 is a deep almost vertical groove that does slightly not reach the ectocingulid. The hypolophid is worn in different degrees in all examined specimens, but despite this, we can see two morphotypes: the hypolophid is weakly separated from the entoconid, the entostylid is absent and the posterolingual groove very weak (Morphotype I, GIN 948/1056, NST/1A) (
|
||
<figureCitation id="61E7FCE7FFA50337CEA5FD2ECA34FD5E" box="[443,557,710,735]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7I</figureCitation>
|
||
); the hypolophid is clearly separated from the entoconid, the small entostylid and the shallow and short posterolingual groove are present (Morphotype II, GIN 1115/1126, BRK/1B). The entoconid is high and relatively narrow; the entocristid is separated from the metaconid base by a narrow transverse groove. The ectocingulid is well-distinguished along the base of the tooth; entocingulid is absent. m2 notably less than m1; the crown bears the high trigonid and the low talonid. The entostylid is absent; the hypolophid contacts the entoconid without the groove. The entocristid is separated the base of the metaconid by a narrow slot. The ectocingulid m2 is very well-developed along the tooth base with the plate-like extension in the first third (paraconid level). The small m3 has the well-developed trigonid and the reduced talonid without a talonid basin; the talonid consists of the small hypoconid and entoconid that separated by the shallow posterolingual groove (
|
||
<figureCitation id="61E7FCE7FFA50337CE7AFAAEC9FBFADE" box="[356,482,1350,1375]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7F</figureCitation>
|
||
). The ectocingulid of m3 bears the plate-like extension in the first third that is similar to m2.
|
||
</paragraph>
|
||
<caption id="ADA3B0EAFFA60334CFDCFA2CCCA9F8E7" ID-DOI="http://doi.org/10.5281/zenodo.11063308" ID-Zenodo-Dep="11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="13" pageNumber="14" startId="13.[194,285,1476,1499]" targetBox="[305,1326,164,1454]" targetPageId="13" targetType="figure">
|
||
<paragraph id="F963E062FFA60334CFDCFA2CCCA9F8E7" blockId="13.[194,1435,1476,1894]" pageId="13" pageNumber="14">
|
||
<emphasis id="CBA83C70FFA60334CFDCFA2CC922FA5D" bold="true" box="[194,315,1476,1500]" pageId="13" pageNumber="14">FIGURE 7.</emphasis>
|
||
Teeth and bones of
|
||
<taxonomicName id="3EDC9BE1FFA60334CD3DFA2DCB4DFA5D" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[547,852,1477,1500]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA60334CD3DFA2DCB4DFA5D" box="[547,852,1477,1500]" italics="true" pageId="13" pageNumber="14">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA60334CC43FA2DCBADFA5D" box="[861,948,1477,1500]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and
|
||
<taxonomicName id="3EDC9BE1FFA60334CE55FA0ACB23FA78" authority="Rzebik-Kowalska, 1975" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[331,826,1506,1529]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA60334CE55FA0ACA2FFA78" box="[331,566,1506,1529]" italics="true" pageId="13" pageNumber="14">Paranourosorex gigas</emphasis>
|
||
Rzebik-Kowalska, 1975
|
||
</taxonomicName>
|
||
(A, E, H, J, L, N, P, S).
|
||
<emphasis id="CBA83C70FFA60334CB21FA0ACC49FA78" bold="true" box="[1087,1104,1506,1529]" pageId="13" pageNumber="14">A</emphasis>
|
||
, GIN 1118/1044 from PSH/1B, right I1 in lateral view;
|
||
<emphasis id="CBA83C70FFA60334CEA6FA17C9D0F997" bold="true" box="[440,457,1535,1558]" pageId="13" pageNumber="14">B</emphasis>
|
||
, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view;
|
||
<emphasis id="CBA83C70FFA60334CB70FA17CC66F997" bold="true" box="[1134,1151,1535,1558]" pageId="13" pageNumber="14">C</emphasis>
|
||
, GIN 948/1065 (paratype) from NST/1A, left P4 (
|
||
<emphasis id="CBA83C70FFA60334CEA5F9F4C9CEF9B9" bold="true" box="[443,471,1564,1592]" pageId="13" pageNumber="14">
|
||
C
|
||
<subScript id="6558E227FFA60334CED2F9CEC9CEF9B9" attach="left" box="[460,471,1574,1592]" fontSize="7" pageId="13" pageNumber="14">1</subScript>
|
||
</emphasis>
|
||
, occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD94F9F4CABFF9B9" bold="true" box="[650,678,1564,1592]" pageId="13" pageNumber="14">
|
||
C
|
||
<subScript id="6558E227FFA60334CD85F9CECABFF9B9" attach="left" box="[667,678,1574,1592]" fontSize="7" pageId="13" pageNumber="14">2</subScript>
|
||
</emphasis>
|
||
, tilted occlusal view);
|
||
<emphasis id="CBA83C70FFA60334CC80F9F5CBB6F9B5" bold="true" box="[926,943,1565,1588]" pageId="13" pageNumber="14">D</emphasis>
|
||
, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD25F9D6CA52F9D4" bold="true" box="[571,587,1598,1621]" pageId="13" pageNumber="14">E</emphasis>
|
||
, GIN 1118/1028 from PSH/1B, left P4 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CBBFF9D6CCB4F9D4" bold="true" box="[1185,1197,1598,1621]" pageId="13" pageNumber="14">F</emphasis>
|
||
, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (
|
||
<emphasis id="CBA83C70FFA60334CDB7F9B4CADAF9F6" bold="true" box="[681,707,1628,1655]" pageId="13" pageNumber="14">
|
||
F
|
||
<subScript id="6558E227FFA60334CDA6F98DCADAF9F6" attach="left" box="[696,707,1637,1655]" fontSize="7" pageId="13" pageNumber="14">1</subScript>
|
||
</emphasis>
|
||
, SEM image;
|
||
<emphasis id="CBA83C70FFA60334CC47F9B4CB6AF9F6" bold="true" box="[857,883,1628,1655]" pageId="13" pageNumber="14">
|
||
F
|
||
<subScript id="6558E227FFA60334CC76F98DCB6AF9F6" attach="left" box="[872,883,1637,1655]" fontSize="7" pageId="13" pageNumber="14">2</subScript>
|
||
</emphasis>
|
||
, diagrammatic image);
|
||
<emphasis id="CBA83C70FFA60334CB73F9B3CC64F9F3" bold="true" box="[1133,1149,1627,1650]" pageId="13" pageNumber="14">G</emphasis>
|
||
, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD6AF995CA9CF915" bold="true" box="[628,645,1661,1684]" pageId="13" pageNumber="14">H</emphasis>
|
||
, GIN 1118/1005 from PSH/1B, right M1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CBF0F995CCECF915" bold="true" box="[1262,1269,1661,1684]" pageId="13" pageNumber="14">I</emphasis>
|
||
, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CDF6F973CAECF933" bold="true" box="[744,757,1691,1714]" pageId="13" pageNumber="14">J</emphasis>
|
||
, GIN 1118/1013 from PSH/1B, left m1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CA4EF973CD78F933" bold="true" box="[1360,1377,1691,1714]" pageId="13" pageNumber="14">K</emphasis>
|
||
, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CC21F950CB57F94E" bold="true" box="[831,846,1720,1743]" pageId="13" pageNumber="14">L</emphasis>
|
||
, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD1EF93DCA0DF96D" bold="true" box="[512,532,1749,1772]" pageId="13" pageNumber="14">M</emphasis>
|
||
, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CBC1F93DCCE9F96D" bold="true" box="[1247,1264,1749,1772]" pageId="13" pageNumber="14">N</emphasis>
|
||
, GIN 11181036 from PSH/1B, left A1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD4BF91ACA71F888" bold="true" box="[597,616,1778,1801]" pageId="13" pageNumber="14">O</emphasis>
|
||
, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CA09F91BCD3DF88B" bold="true" box="[1303,1316,1779,1802]" pageId="13" pageNumber="14">P</emphasis>
|
||
, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view;
|
||
<emphasis id="CBA83C70FFA60334CD90F8F8CAB8F8A6" bold="true" box="[654,673,1808,1831]" pageId="13" pageNumber="14">Q</emphasis>
|
||
, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view;
|
||
<emphasis id="CBA83C70FFA60334CE25F8C5C955F8C5" bold="true" box="[315,332,1837,1860]" pageId="13" pageNumber="14">R</emphasis>
|
||
, GIN 948/1075 (paratype) from NST/1A, right i1 (
|
||
<emphasis id="CBA83C70FFA60334CC4DF8C5CB76F8C8" bold="true" box="[851,879,1837,1865]" pageId="13" pageNumber="14">
|
||
R
|
||
<subScript id="6558E227FFA60334CC7AF8DFCB76F8C8" attach="left" box="[868,879,1847,1865]" fontSize="7" pageId="13" pageNumber="14">1</subScript>
|
||
</emphasis>
|
||
, SEM image in lateral view;
|
||
<emphasis id="CBA83C70FFA60334CB84F8C5CCAFF8C8" bold="true" box="[1178,1206,1837,1865]" pageId="13" pageNumber="14">
|
||
R
|
||
<subScript id="6558E227FFA60334CBB5F8DFCCAFF8C8" attach="left" box="[1195,1206,1847,1865]" fontSize="7" pageId="13" pageNumber="14">2</subScript>
|
||
</emphasis>
|
||
, diagrammatic image in medial view);
|
||
<emphasis id="CBA83C70FFA60334CE70F8A7C967F8E7" bold="true" box="[366,382,1871,1894]" pageId="13" pageNumber="14">S</emphasis>
|
||
, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F963E062FFA50337CFE4FA4ECABDF9DE" blockId="14.[192,793,166,1888]" pageId="14" pageNumber="15">
|
||
A single left premaxilla-maxilla fragment with I1 and A2 from
|
||
<collectionCode id="9FCD78A7FFA50337CE82FA2EC9C1FA61" box="[412,472,1478,1504]" pageId="14" pageNumber="15">PSH</collectionCode>
|
||
/1B (GIN 1118/1027) was examined. I1 is moderately worn; double-rooted A2 is slightly worn. Two alveoli of A1 indicate the presence of the small separated anterior root of A1. Thus, A1 and A2 are both double-rooted.
|
||
</paragraph>
|
||
<paragraph id="F963E062FFA50337CFE4F98ECCE7FEBE" blockId="14.[192,793,166,1888]" lastBlockId="14.[840,1441,166,1887]" pageId="14" pageNumber="15">
|
||
The mandibular body is long and relatively low without a visible inward curve of the lower outline. The mental foramen is positioned under the protoconid of m1 without any grooves in a wide and shallow depression. The mandibular ramus is massive with a small internal fossa for the temporal muscle; its maximal length is lesser than the condylar height (HC). The coronoid process includes a well-developed long coronoid spicule. The single mandibular foramen is large. The lower articular facet of the condylar process shifts far anteriorly; an interarticular area is relatively narrow (
|
||
<figureCitation id="61E7FCE7FFA50337CA4CFEEECB69FEC1" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7Q</figureCitation>
|
||
). An angular process is unknown.
|
||
</paragraph>
|
||
<paragraph id="F963E062FFA50337CC56FEAECD8CFB3E" blockId="14.[840,1441,166,1887]" pageId="14" pageNumber="15">
|
||
<emphasis id="CBA83C70FFA50337CC56FEAECBE0FEE1" bold="true" box="[840,1017,326,352]" pageId="14" pageNumber="15">Comparisons.</emphasis>
|
||
<taxonomicName id="3EDC9BE1FFA50337CB13FEAECD70FEDE" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[1037,1385,326,351]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CB13FEAECD70FEDE" box="[1037,1385,326,351]" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CA62FEAECB60FEFE" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="3EDC9BE1FFA50337CB11FE8ECCB1FEFE" authorityName="Storch and Zazhigin" authorityYear="1996" box="[1039,1192,358,383]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="seletiensis">
|
||
<emphasis id="CBA83C70FFA50337CB11FE8ECCB1FEFE" box="[1039,1192,358,383]" italics="true" pageId="14" pageNumber="15">P. seletiensis</emphasis>
|
||
</taxonomicName>
|
||
in a larger size (
|
||
<figureCitation id="61E7FCE7FFA50337CA6EFE8ECB97FE1E" captionStart="FIGURE 5" captionStartId="9.[194,285,954,977]" captionTargetBox="[247,1385,168,925]" captionTargetId="figure-340@9.[246,1389,168,929]" captionTargetPageId="9" captionText="FIGURE 5. Bivariate plot of linear differences between five Asian species, Paranourosorex seletiensis Storch and Zazhigin, 1996 (PSt.), Paranourosorex gigas Rzebik-Kowalska, 1975 (P. gigas), Paranourosorex intermedius sp. nov. (PIm.) and Ishimosorex ishimiensis gen. et sp. nov. (PPl) based on the second lower molar measurements (L vs TRL), mm). Studied remains from Asian Neogene localities are marked by colours, which corresponds to ELMA (see Figure 1); numbers are given in the Appendix 2." figureDoi="http://doi.org/10.5281/zenodo.11063304" httpUri="https://zenodo.org/record/11063304/files/figure.png" pageId="14" pageNumber="15">Figure 5</figureCitation>
|
||
; Appendix 4) and some dental characters (the first species has longer hypoconal flange of P4 and more inflated m1 and m2 than has the second species;
|
||
<figureCitation id="61E7FCE7FFA50337CCACFE0ECC3FFE7E" box="[946,1062,486,511]" captionStart="FIGURE 6" captionStartId="11.[196,287,1080,1103]" captionTargetBox="[190,1441,166,1060]" captionTargetId="figure-235@11.[190,1441,166,1060]" captionTargetPageId="11" captionText="FIGURE 6. Teeth and bones of Paranourosorex seletiensis Storch and Zazhigin, 1996 (A, B), Crusafontina sp. 1, (C), Crusafontina sp. 2 (D) and Paranourosorex intermedius sp. nov. (E). A, GIN 951/1000 (holotype) from SLT/1A, left hemimandible fragment (cut image) with m1 and m2 in occlusal view (A2, magnified m2); B, GIN 951/1001 (paratype) from SLT/1A, left P4 in occlusal view; C, GIN 640/1004 from PVL/1A, left M1 in occlusal view; D, GIN 951/1003 SLT/ 1A, left P4 in occlusal view; E, GIN 948/1051 (holotype) from NST/1A, right dentary fragment with m1 and m2 (E1, occlusal view; E2, lateral view E3, medial view;). Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063306" httpUri="https://zenodo.org/record/11063306/files/figure.png" pageId="14" pageNumber="15">Figure 6B</figureCitation>
|
||
cf.
|
||
<figureCitation id="61E7FCE7FFA50337CB4AFE0ECCD7FE7E" box="[1108,1230,486,511]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7D</figureCitation>
|
||
).
|
||
<taxonomicName id="3EDC9BE1FFA50337CBFCFE0ECBCBFD9E" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CBFCFE0ECBCBFD9E" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CCC7FDEECC2DFD9E" box="[985,1076,518,543]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="3EDC9BE1FFA50337CBDBFDEECD9DFD9E" baseAuthorityName="Schlosser" baseAuthorityYear="1924" box="[1221,1412,518,543]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="inexspectatus">
|
||
<emphasis id="CBA83C70FFA50337CBDBFDEECD9DFD9E" box="[1221,1412,518,543]" italics="true" pageId="14" pageNumber="15">P. inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
in a slightly larger size; in the characters of P4 such as a more developed anterior notch and a more shallow posterior emargination, absent the entocingulum; in a more developed metastyle of M1 (the metastyle of
|
||
<taxonomicName id="3EDC9BE1FFA50337CCF6FD4ECCB4FD3E" baseAuthorityName="Schlosser" baseAuthorityYear="1924" box="[1000,1197,678,703]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="inexspectatus">
|
||
<emphasis id="CBA83C70FFA50337CCF6FD4ECCB4FD3E" box="[1000,1197,678,703]" italics="true" pageId="14" pageNumber="15">P. inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
is relatively shorter and narrower); in a more lingual position of the hypocone and a slightly deeper posterior emargination of M1 (for
|
||
<taxonomicName id="3EDC9BE1FFA50337CB18FCEECCDFFC9E" baseAuthorityName="Schlosser" baseAuthorityYear="1924" box="[1030,1222,774,799]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="inexspectatus">
|
||
<emphasis id="CBA83C70FFA50337CB18FCEECCDFFC9E" box="[1030,1222,774,799]" italics="true" pageId="14" pageNumber="15">P. inexspectatus</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="9D4D9D93FFA50337CA1CFCEECD85FC9E" author="Storch, G." box="[1282,1436,774,800]" pageId="14" pageNumber="15" pagination="221 - 251" refId="ref17471" refString="Storch, G. 1995. The Neogene mammalian faunas of Ertemte and Harr Obo in Inner Mongolia (Nei Mongol), China. - 11. Soricidae (Insectivora). Senckenbergiana Lethaea, 75: 221 - 251." type="journal article" year="1995">Storch, 1995</bibRefCitation>
|
||
: plate 4).
|
||
<taxonomicName id="3EDC9BE1FFA50337CCA8FCCECD11FCBE" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[950,1288,806,831]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CCA8FCCECD11FCBE" box="[950,1288,806,831]" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CA0CFCCECD68FCBE" box="[1298,1393,806,831]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="3EDC9BE1FFA50337CCDCFCAECC3DFCDE" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[962,1060,838,863]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA50337CCDCFCAECC3DFCDE" box="[962,1060,838,863]" italics="true" pageId="14" pageNumber="15">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
in a slightly smaller size; in a more shallow anterior notch and posterior emargination of P4 (
|
||
<figureCitation id="61E7FCE7FFA50337CCCBFC6ECC25FC1E" box="[981,1084,902,927]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7</figureCitation>
|
||
: C cf. E); in a more developed metastyle of M1 (the metastyle of
|
||
<taxonomicName id="3EDC9BE1FFA50337CBFBFC4ECD5DFC3E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[1253,1348,934,959]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA50337CBFBFC4ECD5DFC3E" box="[1253,1348,934,959]" italics="true" pageId="14" pageNumber="15">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
is relatively narrower); in a more lingual position of the hypocone, a slightly longer hypoconal flange and a slightly deeper posterior emargination of M1; in absence of the entocingulum of M1 (
|
||
<figureCitation id="61E7FCE7FFA50337CBEBFBCECD45FBBE" box="[1269,1372,1062,1087]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7</figureCitation>
|
||
: G cf. H); in strongly exaenodont buccal half of a1 (
|
||
<figureCitation id="61E7FCE7FFA50337CA4CFBAECB43FBFE" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7</figureCitation>
|
||
: O cf. P); in presence of a weak posterolingual groove of m1 (
|
||
<taxonomicName id="3EDC9BE1FFA50337CB18FB6ECC73FB1E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[1030,1130,1158,1183]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA50337CB18FB6ECC73FB1E" box="[1030,1130,1158,1183]" italics="true" pageId="14" pageNumber="15">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
has not the groove, the hypolophid contacts the entoconid; Morphotype I).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1C6B3E9FFA50336CC56FB2ECA83FD83" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="F963E062FFA50337CC56FB2ECC8BF93E" blockId="14.[840,1441,166,1887]" pageId="14" pageNumber="15">
|
||
<emphasis id="CBA83C70FFA50337CC56FB2ECBD9FB5E" bold="true" box="[840,960,1222,1247]" pageId="14" pageNumber="15">Remarks.</emphasis>
|
||
The fossils of
|
||
<taxonomicName id="3EDC9BE1FFA50337CB69FB2ECB60FB7E" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CB69FB2ECB60FB7E" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CC9CFB0ECBC6FB7E" box="[898,991,1254,1279]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
correspond to ‘
|
||
<taxonomicName id="3EDC9BE1FFA50337CB87FB0ECD84FB7E" box="[1177,1437,1254,1279]" class="Mammalia" family="Soricidae" genus="Paranourosorex" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undefined-2">
|
||
<emphasis id="CBA83C70FFA50337CB87FB0ECD4EFB7E" box="[1177,1367,1254,1279]" italics="true" pageId="14" pageNumber="15">Paranourosorex</emphasis>
|
||
sp. 2
|
||
</taxonomicName>
|
||
’ in
|
||
<bibRefCitation id="9D4D9D93FFA50337CC79FAEECCEDFA9E" author="Storch, G. & Zazhigin, V. S." box="[871,1268,1286,1312]" pageId="14" pageNumber="15" pagination="257 - 268" refId="ref17571" refString="Storch, G. and Zazhigin, V. S. 1996. Taxonomy and phylogeny of the Paranourosorex lineage, Neogene of Eurasia (Mammalia: Soricidae: Anourooricini). Palaontologische Zeitschrift, 70: 257 - 268. https: // doi. org / 10.1007 / BF 02988282" type="journal article" year="1996">Storch and Zazhigin (1996: 264)</bibRefCitation>
|
||
. That description comprises several discordant features that require clarification. Authors stated: “its [p4] apex is weakly two-cusped.” We re-evaluated the apex of p4 as a distinctly single cusp with a spot-shaped wear facet (
|
||
<figureCitation id="61E7FCE7FFA50337CCC4FA4ECC43FA3E" box="[986,1114,1446,1471]" captionStart="FIGURE 7" captionStartId="13.[194,285,1476,1499]" captionTargetBox="[305,1326,164,1454]" captionTargetId="figure-21@13.[305,1326,164,1454]" captionTargetPageId="13" captionText="FIGURE 7. Teeth and bones of Paranourosorex intermedius sp. nov. (B, C–D, F–G, I, K, M, O, Q–R; with dotted frames) and Paranourosorex gigas Rzebik-Kowalska, 1975 (A, E, H, J, L, N, P, S). A, GIN 1118/1044 from PSH/1B, right I1 in lateral view; B, GIN 948/1073 (paratype) from NST/1A, left I1 in lateral view; C, GIN 948/1065 (paratype) from NST/1A, left P4 (C1, occlusal view; C2, tilted occlusal view); D, GIN 1130/1104 from LZN/2B, diagrammatic image of right P4 in occlusal view; E, GIN 1118/1028 from PSH/1B, left P4 in occlusal view; F, GIN 948/1064 (paratype) from NST/1A, right m3 in occlusal view (F1, SEM image; F2, diagrammatic image); G, GIN 948/1069 (paratype) from NST/1A, right M1 in occlusal view; H, GIN 1118/1005 from PSH/1B, right M1 in occlusal view; I, GIN 948/1056 (paratype) from NST/1A, right m1 in occlusal view; J, GIN 1118/1013 from PSH/1B, left m1 in occlusal view; K, GIN 948/1078 (paratype) from NST/1A, left p4 in occlusal view; L, GIN 1118/1011 from PSH/1B, left p4 and m1 fragment (cut image) in occlusal view; M, GIN 948/1089 (paratype) from NST/1A, left A1 in occlusal view; N, GIN 11181036 from PSH/1B, left A1 in occlusal view; O, GIN 948/1081 (paratype) from NST/1A, left a1 in occlusal view; P, GIN 1118/ 1008 from PSH/1B, left a1 in occlusal view; Q, GIN 1115/1123 from BRK/1A, left fragment of mandibular ramus in posterior view; R, GIN 948/1075 (paratype) from NST/1A, right i1 (R1, SEM image in lateral view; R2, diagrammatic image in medial view); S, 1118 951/1021 from PSH/1B, right i1 in lateral view. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063308" httpUri="https://zenodo.org/record/11063308/files/figure.png" pageId="14" pageNumber="15">Figure 7K</figureCitation>
|
||
). Authors also stated: “the hypolophid on M/1 is separated from the entoconid by a groove (even in worn specimens).” We found two morphotypes of hypolophid/entoconid of m1 state when the hypolophid is weakly separated from the entoconid with a very weak groove, either the hypolophid is clearly separated from the entoconid with the small entostylid and the shallow and short posterolingual groove.
|
||
</paragraph>
|
||
<paragraph id="F963E062FFA50336CC9CF92EC973FEC3" blockId="14.[840,1441,166,1887]" lastBlockId="15.[192,793,169,514]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
Much material of
|
||
<taxonomicName id="3EDC9BE1FFA50337CB70F92ECB60F97E" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CB70F92ECB60F97E" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CC96F90ECBF5F97E" box="[904,1004,1766,1791]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
originates from NST/1A (n = 39), BRK/1A (n = 26) and BRK/1B (n = 18). The species is more poorly represented in the other nine sites (Appendix 2).
|
||
<taxonomicName id="3EDC9BE1FFA50337CCF0F8AECD24F8DE" authority="Zazhigin & Voyta, 2022" authorityName="Zazhigin & Voyta" authorityYear="2022" box="[1006,1341,1862,1887]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
|
||
<emphasis id="CBA83C70FFA50337CCF0F8AECD24F8DE" box="[1006,1341,1862,1887]" italics="true" pageId="14" pageNumber="15">Paranourosorex intermedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D09B810BFFA50337CA5AF8AECDB8F8DE" box="[1348,1441,1862,1887]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
probably coexisted with larger shrew
|
||
<taxonomicName id="3EDC9BE1FFA40336CD8DFF41CAEFFF43" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[659,758,169,194]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="CBA83C70FFA40336CD8DFF41CAEFFF43" box="[659,758,169,194]" italics="true" pageId="15" pageNumber="16">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
in CHR/1A (Turolian, MN 13, Rytov Formation),
|
||
<collectionCode id="9FCD78A7FFA40336CDFBFF20CB16FF63" box="[741,783,200,226]" pageId="15" pageNumber="16">ISK</collectionCode>
|
||
/ 1A (Ruscinian, MN 14, Isakov Formation),
|
||
<collectionCode id="9FCD78A7FFA40336CDA7FF00CAEDFE83" box="[697,756,232,258]" pageId="15" pageNumber="16">PSH</collectionCode>
|
||
/1A and
|
||
<collectionCode id="9FCD78A7FFA40336CFE8FEE0C92BFEA3" box="[246,306,264,290]" pageId="15" pageNumber="16">PSH</collectionCode>
|
||
/1B (Ruscinian, MN 14, Peshnev Formation) localities.
|
||
</paragraph>
|
||
<paragraph id="F963E062FFA40336CFDEFEA0CA83FD83" blockId="15.[192,793,169,514]" pageId="15" pageNumber="16">
|
||
<emphasis id="CBA83C70FFA40336CFDEFEA0CA94FEE3" bold="true" box="[192,653,328,354]" pageId="15" pageNumber="16">Stratigraphic and geographic range.</emphasis>
|
||
At present, the distribution includes the
|
||
<typeStatus id="26675EC0FFA40336CD3DFE81CA4CFE03" box="[547,597,361,386]" pageId="15" pageNumber="16">type</typeStatus>
|
||
locality and 11 localities from
|
||
<collectingCountry id="81CBA0F2FFA40336CE6AFE61C9E6FE23" box="[372,511,393,418]" name="Kazakhstan" pageId="15" pageNumber="16">Kazakhstan</collectingCountry>
|
||
and
|
||
<collectingCountry id="81CBA0F2FFA40336CD5DFE61CA8DFE23" box="[579,660,393,418]" name="Russia" pageId="15" pageNumber="16">Russia</collectingCountry>
|
||
(Appendix 2) between the late
|
||
<location id="FC03B6B9FFA40336CEB1FE41C909FE63" LSID="urn:lsid:plazi:treatment:71755174FFA70336CE0DFD79CA83FD83:FC03B6B9FFA40336CEB1FE41C909FE63" country="Kazakhstan" municipality="Pliocene Peshnev Formation" name="Miocene Novaya Stanitsa Formation" pageId="15" pageNumber="16">Miocene Novaya Stanitsa Formation</location>
|
||
(
|
||
<collectorName id="542985B4FFA40336CE3DFE21C9D8FE63" box="[291,449,457,482]" pageId="15" pageNumber="16">Turolian, MN</collectorName>
|
||
13) and the early
|
||
<collectingMunicipality id="19077A18FFA40336CDACFE21C9BEFD83" pageId="15" pageNumber="16">Pliocene Peshnev Formation</collectingMunicipality>
|
||
(
|
||
<collectorName id="542985B4FFA40336CEA9FE01CA7DFD83" box="[439,612,489,514]" pageId="15" pageNumber="16">Ruscinian, MN</collectorName>
|
||
14).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |