treatments-xml/data/F1/82/40/F182407BF9175965A4C8C010C564271F.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="F182407BF9175965A4C8C010C564271F" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria irregulara Miller 2022, sp. nov." docType="treatment" docUuid="58E77BB3-EC2F-4F6A-85E7-E0FCBEF2E331" docUuidSource="ZooBank" docVersion="1" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287389350" updateUser="pensoft">
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<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-16</mods:number>
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<mods:number>1136</mods:number>
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<mods:start>1</mods:start>
<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/58E77BB3-EC2F-4F6A-85E7-E0FCBEF2E331" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria irregulara" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irregulara" status="sp. nov.">Desmopachria irregulara</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">Figures 69-71</figureCitation>
<figureCitation captionStart="Figure 81" captionStartId="F10" captionText="Figure 81. Desmopachria species distributions, Desmopachria portmanni-aldessa species group, Desmopachria portmanni-portmanni species group, and Desmopachria striola species group." figureDoi="10.3897/zookeys.1136.72744.figure81" httpUri="https://binary.pensoft.net/fig/782462" pageId="0" pageNumber="1">, 81</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Venezuela, Zulia State, Perija National Park, Tukuko, 
<normalizedToken originalValue="Río">Rio</normalizedToken>
Manantial, 
<geoCoordinate degrees="9" direction="north" minutes="50.490" orientation="latitude" precision="1" value="9.8415">9°50.490'N</geoCoordinate>
, 
<geoCoordinate degrees="72" direction="west" minutes="49.310" orientation="longitude" precision="1" value="-72.82183">72°49.310'W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is distinctive for the non-iridescent dorsal and ventral surfaces and the characteristic male median and lateral lobes. The median lobe has an extremely large lobe on the dorsal surface of the median lobe medially (Fig. 
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">69</figureCitation>
). The lateral lobe is broad throughout with a short, dense series of setae along the apical and subapical dorsal margins and a cluster of elongate setae subapically on the ventral margin (Fig. 
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">71</figureCitation>
). The elytral punctation is large with few minute punctures interspersed. The pronotal punctation is extremely fine except posteromedially where it is large and dense.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 2.1-2.2 mm, GW = 1.4-1.5 mm, PW = 1.1-1.2 mm, HW = 0.7-0.8 mm, EW = 0.4 mm, TL/GW = 1.4-1.5, HW/EW = 1.8-1.9. Body very broad, laterally rounded, lateral margins continuous between pronotum and elytron; dorsoventrally compressed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration.</emphasis>
Head and pronotum evenly orange-red. Elytron evenly orange-red, similar to pronotum and head, not iridescent. Metaventrites, metacoxae and abdominal ventrites orange-red, other surfaces orange.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad, short; anterior margin of clypeus finely margined with continuous flattened narrow bead; surface of head shiny, impunctate medially, finely punctate posteriorly; eyes large (HW/EW = 1.8-1.9); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, shallowly curved with continuous narrow bead; surface shiny, punctation very fine and sparse over most of surface, punctation large and dense posteromedially, posterior margin sinuate. Elytron broad, laterally broadly curved; surface shiny, punctation dual, mostly large, relatively dense, with fewer fine punctures interspersed. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process in male very slender anteriorly, with low, indistinct medial tubercle, bifid apically with deep medial pit, in female slender anteriorly, with distinctive, small medial tubercle, apically short and broad, medially slightly carinate, apically acutely pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/3 length of metaventrite medially, metacoxal lines sinuate, divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, very finely punctate, not iridescent. Metatrochanter very large, subequal to length of ventral margin of metafemur; male metafemur curved ventrally with conspicuous series of short setae long entire margin. Abdomen with surfaces shiny and smooth, finely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male median lobe in lateral aspect irregular, ventral margin abruptly expanded submedially, evenly convexly curved in apical 2/5, ventral margin with large, apically subtruncate lobe medially, apex slight broadened, apex subtruncate (Fig. 
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">69</figureCitation>
); in ventral aspect slender, lateral margins shallowly sinuate, apex deeply bifid, lateral rami apically broad, directed slightly mesad (Fig. 
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">70</figureCitation>
). Lateral lobe very broad throughout length, slightly curved dorsad, apex broadly rounded, subapically on ventral surface with small cluster of elongate setae, apex and subapically on dorsal margin with dense series of short setae (Fig. 
<figureCitation captionStart="Figures 51–71" captionStartId="F5" captionText="Figures 51 - 71. Desmopachria species. (51 - 53) Desmopachria angulata 51 median lobe, right lateral aspect 52 median lobe, ventral aspect 53 right lateral lobe, right lateral aspect (54 - 56) Desmopachria emarginata 54 median lobe, right lateral aspect 55 median lobe, ventral aspect 56 right lateral lobe, right lateral aspect (57 - 59) Desmopachria imparis 57 median lobe, right lateral aspect 58 median lobe, ventral aspect 59 right lateral lobe, right lateral aspect (60 - 62) Desmopachria impunctata 60 median lobe, right lateral aspect 61 median lobe, ventral aspect 62 right lateral lobe, right lateral aspect (63 - 65) Desmopachria truncata 63 median lobe, right lateral aspect 64 median lobe, ventral aspect 65 right lateral lobe, right lateral aspect (66 - 68) Desmopachria bisulcata 66 median lobe, right lateral aspect 67 median lobe, ventral aspect 68 right lateral lobe, right lateral aspect (69 - 71) Desmopachria irregulara 69 median lobe, right lateral aspect 70 median lobe, ventral aspect 71 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1136.72744.figures51-71" httpUri="https://binary.pensoft.net/fig/782457" pageId="0" pageNumber="1">71</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism.</emphasis>
Only males were examined, but male pro- and mesotarsomeres I-III appear to be slightly more broadly expanded and with ventral adhesive setae, and male prosternal process is different as in all 
<taxonomicName baseAuthorityName="Clark" baseAuthorityYear="1862" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria portmanni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="portmanni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria portmanni</emphasis>
</taxonomicName>
group species.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation.</emphasis>
There is little variation between the two male specimens examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria irregulara" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irregulara">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria irregulara</emphasis>
</taxonomicName>
, Latin for irregular, for the unusual shape of the male median lobe.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from one locality in Zulia State, Venezuela (Fig. 
<figureCitation captionStart="Figure 81" captionStartId="F10" captionText="Figure 81. Desmopachria species distributions, Desmopachria portmanni-aldessa species group, Desmopachria portmanni-portmanni species group, and Desmopachria striola species group." figureDoi="10.3897/zookeys.1136.72744.figure81" httpUri="https://binary.pensoft.net/fig/782462" pageId="0" pageNumber="1">81</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in MIZA, male labeled, &quot;VENEZUELA: Zulia State 
<geoCoordinate degrees="9" direction="north" minutes="50.490" orientation="latitude" precision="1" value="9.8415">9°50.490'N</geoCoordinate>
, 
<geoCoordinate degrees="72" direction="west" minutes="49.310" orientation="longitude" precision="1" value="-72.82183">72°49.310'W</geoCoordinate>
, Perija, Nat. Park: Tukuko: 
<normalizedToken originalValue="Río">Rio</normalizedToken>
Manatial 29.i.2009; Short, 
<normalizedToken originalValue="García">Garcia</normalizedToken>
, Camacho VZ09-0129-01B: detrital pool/ SM0845453 KUNHM-ENT [barcode label]/ HOLOTYPE 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria irregulara" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irregulara">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria irregulara</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 1 in SEMC labeled same as holotype except with 
<normalizedToken originalValue="“…SM08454561…">&quot;...SM08454561...</normalizedToken>
[barcode label]&quot; and with 
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
PARATYPE 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria irregulara" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irregulara">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria irregulara</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border].&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="checklist of desmopachria portmanni-portmanni subgroup – includes the non-iridescent desmopachria with forked male prosternal process (= desmopachria (portmannia) young, 1980)">
<paragraph pageId="0" pageNumber="1">
Checklist of 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria portmanni-portmanni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="portmanni-portmanni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria portmanni-portmanni</emphasis>
</taxonomicName>
subgroup - includes the non-iridescent 
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
with forked male prosternal process (= 
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmopachria</taxonomicName>
(
<taxonomicName authorityName="Young" authorityYear="1980" class="Insecta" family="Dytiscidae" genus="Portmannia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Portmannia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Portmannia</taxonomicName>
) Young, 1980)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Guignot" authorityYear="1950" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria basicollis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="basicollis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria basicollis</emphasis>
</taxonomicName>
Guignot, 1950 - Costa Rica
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bisulcata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bisulcata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bisulcata</emphasis>
</taxonomicName>
sp. nov. - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bryanstoni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bryanstoni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bryanstoni</emphasis>
</taxonomicName>
Clark, 1862 - Mexico, Panama, Guatemala
</paragraph>
<paragraph pageId="0" pageNumber="1">
= 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria polita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="polita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria polita</emphasis>
</taxonomicName>
Sharp, 1882
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria carranca" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carranca">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria carranca</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2018 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1995" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria decorosa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="decorosa">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria decorosa</emphasis>
</taxonomicName>
Young, 1995 - Mexico
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria dicrophalica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dicrophalica">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria dicrophalica</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria dispar" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dispar">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria dispar</emphasis>
</taxonomicName>
Sharp, 1882 - Mexico
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria disticta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="disticta">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria disticta</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria duodentata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="duodentata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria duodentata</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2011 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1995" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria goias" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="goias">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria goias</emphasis>
</taxonomicName>
Young, 1995 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria grammosticta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grammosticta">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria grammosticta</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria grandinigra" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="grandinigra">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria grandinigra</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria irregulara" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="irregulara">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria irregulara</emphasis>
</taxonomicName>
sp. nov. - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria itamontensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="itamontensis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria itamontensis</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria laevis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laevis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria laevis</emphasis>
</taxonomicName>
Sharp, 1882 - Brazil (Braga and Ferreira-Jr. 2014)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria mutata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mutata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria mutata</emphasis>
</taxonomicName>
Sharp, 1882 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">nomen novum</emphasis>
for 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bryanstoni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bryanstoni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bryanstoni</emphasis>
</taxonomicName>
Sharp, 1882
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zimmermann" authorityYear="1923" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria niger" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niger">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria niger</emphasis>
</taxonomicName>
Zimmermann, 1923 - Brazil (
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nigricoxa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nigricoxa">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nigricoxa</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2018 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nigrisphera" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nigrisphera">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nigrisphera</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2018 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zimmermann" authorityYear="1928" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitidissima" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidissima">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitidissima</emphasis>
</taxonomicName>
Zimmermann, 1928 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1995" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria pittieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pittieri">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria pittieri</emphasis>
</taxonomicName>
<bibRefCitation author="Young, FN" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="37 - 45" refId="B24" refString="Young, FN, 1995. The genus Desmopachria Babington, subgenus Portmannia Young (Coleoptera: Dytiscidae). Insecta Mundi 9: 37 - 45" title="The genus Desmopachria Babington, subgenus Portmannia Young (Coleoptera: Dytiscidae)." volume="9" year="1995">Young 1995</bibRefCitation>
- Costa Rica
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Clark" baseAuthorityYear="1862" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria portmanni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="portmanni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria portmanni</emphasis>
</taxonomicName>
Clark, 1862 - USA and N Mexico
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1995" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria sobrina" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sobrina">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria sobrina</emphasis>
</taxonomicName>
Young, 1995 - Panama
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1887" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria specula" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="specula">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria specula</emphasis>
</taxonomicName>
Sharp, 1887 - Panama (probably not part of 
<taxonomicName baseAuthorityName="Clark" baseAuthorityYear="1862" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria portmanni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="portmanni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria portmanni</emphasis>
</taxonomicName>
species group according to 
<bibRefCitation author="Young, FN" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="37 - 45" refId="B24" refString="Young, FN, 1995. The genus Desmopachria Babington, subgenus Portmannia Young (Coleoptera: Dytiscidae). Insecta Mundi 9: 37 - 45" title="The genus Desmopachria Babington, subgenus Portmannia Young (Coleoptera: Dytiscidae)." volume="9" year="1995">Young (1995)</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria ukuki" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ukuki">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria ukuki</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria undulatosterna" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="undulatosterna">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria undulatosterna</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2011 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria variegata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="variegata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria variegata</emphasis>
</taxonomicName>
Sharp, 1882 - Mexico, El Salvador, Honduras
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria youngi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="youngi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria youngi</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1995" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria zetha" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zetha">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria zetha</emphasis>
</taxonomicName>
Young, 1995 - Mexico
</paragraph>
</subSubSection>
</treatment>
</document>