treatments-xml/data/2E/BA/8E/2EBA8EB5F4A818324C85FCE46395A1ED.xml

<document ID-DOI="http://dx.doi.org/10.3897/dez.66.31754" ID-GBIF-Dataset="22c3ecd6-0d0a-443a-ba47-023ac8504bcc" ID-GBIF-Taxon="182223588" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="8A92B765B02355A2A9D1168A93C881F2" ID-Zenodo-Dep="2551893" ID-ZooBank="11B6BFA2262745108C7C5D38945AA4F1" LSID="urn:lsid:zoobank.org:act:BCB33301-A988-465D-A0F5-82060EAD16B0" ModsDocID="1860-1324-66-1-1" checkinTime="1621372408696" checkinUser="pensoft" docAuthor="Lohaj, Roman &amp; Delic, Teo" docDate="2019" docId="2EBA8EB5F4A818324C85FCE46395A1ED" docLanguage="en" docName="DeutEntomolZeit 66(1): 1-15" docOrigin="Deutsche Entomologische Zeitschrift 66 (1)" docPubDate="2019-01-21" docSource="http://dx.doi.org/10.3897/dez.66.31754" docTitle="Derossiella lukici Lohaj &amp; Delić 2019, sp. n." docType="treatment" docUuid="BCB33301-A988-465D-A0F5-82060EAD16B0" docUuidSource="ZooBank" docVersion="3" id="8A92B765B02355A2A9D1168A93C881F2" lastPageNumber="1" masterDocId="8A92B765B02355A2A9D1168A93C881F2" masterDocTitle="Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst" masterLastPageNumber="15" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1643457464602" updateUser="ExternalLinkService">
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<mods:title>Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lohaj, Roman</mods:namePart>
<mods:affiliation>Zahradna 61 A, SK- 900 91 Limbach, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Delic, Teo</mods:namePart>
<mods:affiliation>SubBioLab, Department of Biology, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, SI- 1000 Ljubljana, Slovenia</mods:affiliation>
<mods:nameIdentifier type="email">tejc86@gmail.com</mods:nameIdentifier>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2019</mods:date>
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<mods:number>2019-01-21</mods:number>
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<mods:detail type="volume">
<mods:number>66</mods:number>
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<mods:number>1</mods:number>
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<mods:end>15</mods:end>
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<mods:url>http://dx.doi.org/10.3897/dez.66.31754</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.66.31754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
<mods:identifier type="ZooBank">11B6BFA2262745108C7C5D38945AA4F1</mods:identifier>
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<treatment ID-GBIF-Taxon="182223588" LSID="urn:lsid:zoobank.org:act:BCB33301-A988-465D-A0F5-82060EAD16B0" httpUri="http://treatment.plazi.org/id/2EBA8EB5F4A818324C85FCE46395A1ED" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/BCB33301-A988-465D-A0F5-82060EAD16B0" authority="Lohaj &amp; Delić, 2019" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici" status="sp. n.">Derossiella lukici</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geographical distribution of the Dinaric aphaenopsoid trechines of the genera Derossiella and Adriaphenops. Source of data: http: // subbio. net / db /" figureDoi="10.3897/dez.66.31754.figure2" httpUri="https://binary.pensoft.net/fig/258009" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Derossiella lukici sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. Bregovic)." figureDoi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type series">
<paragraph pageId="0" pageNumber="1">Type series.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype male labelled: &quot;Croatia, Mt Biokovo, Golubinjak, Biokovka, -300 m, 24.6.2017, T. 
<normalizedToken originalValue="Delić">Delic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;DNA extraction RL-07&quot; (orange label, printed) / &quot;HOLOTYPUS 
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CNHM, voucher code 600: ZAG; ZEC2, 4194 Coll. 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
). Paratypes: one female (right posterior tarsus missing) labelled: &quot;Croatia, Mt Biokovo, Golubinjak, Biokovka, 2.9.2007, M. 
<normalizedToken originalValue="Lukić">Lukic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;PARATYPUS 
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CNHM, voucher code 600: ZAG; ZEC2, 4195 Coll. 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
)., one female (last three antennomeres of right antenna missing) labelled: &quot;Croatia, Mt Biokovo, Lokva, Pretnerova, -120 m, 19.05.2015, E. Premate lgt.&quot; (white label, printed) / &quot;PARATYPUS 
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CRL).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Medium-sized aphaenopsoid trechine with morphological features fully matching generic description proposed by 
<bibRefCitation author="Queinnec, E" editor="Pavicevic, D" journalOrPublisher="Institute for Nature Conservation of Serbia, Belgrade" pageId="0" pageNumber="1" pagination="157 - 176" refId="B44" refString="Queinnec, E, 2008. Two new genera and species of 'aphaenopsid' cave beetles from the Balkan Peninsula and considerations about the evolutionary trends of the so-called &quot;hyper-specialised&quot; trechines. In: Pavicevic, D, Perreau, M, Eds., Advances in the Studies of the Fauna of the Balkan Peninsula. Institute for Nature Conservation of Serbia, Belgrade: 157 - 176" title="Two new genera and species of ' aphaenopsid' cave beetles from the Balkan Peninsula and considerations about the evolutionary trends of the so-called &quot; hyper-specialised &quot; trechines." volumeTitle="Advances in the Studies of the Fauna of the Balkan Peninsula." year="2008">
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
(2008)
</bibRefCitation>
. Head long, parallel-sided, with complete, deep frontal furrows reaching neck constriction and two pairs of supraorbital setae, posterior setae doubled. Eyes absent, mentum with simple tooth. Aphaenopsoid habitus with very narrow head and pronotum; elytra oblong-oval, 2.6 times wider than head and pronotum. Hind angles of pronotum without setae. Surface completely glabrous, striae absent, elytral chaetotaxy with macrochetae and microchetae. Cuticle depigmented, reddish-yellow, body strongly flattened dorso-ventrally, with very long and slender legs and antennae (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Derossiella lukici sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. Bregovic)." figureDoi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1">3</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Closely related to the type species of the genus, 
<taxonomicName authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
, from which it differs by paired posterior supraorbital setiferous punctures, differently shaped elytra with higher number of macro- and microchetae, narrower pronotum, as well as by the differently shaped aedeagus.
</paragraph>
<caption ID-Zenodo-Dep="2551937" doi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. 
<normalizedToken originalValue="Bregović">Bregovic</normalizedToken>
).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="2551939" doi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Habitus of 
<taxonomicName genus="Derosiella" lsidName="Derosiella" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Derosiella</emphasis>
</taxonomicName>
species. 
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. 
<emphasis bold="true" pageId="0" pageNumber="1">(a)</emphasis>
and 
<taxonomicName authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(b)</emphasis>
. Chetotaxy is presented as white points (photo courtesy of 
<normalizedToken originalValue="Dušan">Dusan</normalizedToken>
<normalizedToken originalValue="Beňo">Beno</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">L: 5.5 mm (PT)-6.0 mm (HT), TL: 4.8 mm (PT)-5.4 mm (HT). Head relatively large, nearly parallel-sided, with maximum width behind middle, distinctly longer than wide (index HL/HW 1.35 (PT)-1.42 (HT), slightly wider than pronotum, sparsely pubescent. Frontal furrows deep, complete, reaching neck constriction, slightly divergent posteriorly. Anterior pair of supraorbital setae situated before middle of head length, posterior supraorbital setae paired, two setae on each side of head situated close to neck constriction. Neck constriction distinct; genae gently convex. Clypeus and labrum with three pairs of setae, outer pairs longer. Antennae length 4.9 mm (HT)-4.3 mm (PT), scape as long as pedicel, almost as long as terminal antennomere.</paragraph>
<paragraph pageId="0" pageNumber="1">Pronotum elongated, slightly longer and narrower than head, with maximum width in anterior third, index PL/PW 1.75 (PT)-1.83 (HT), only slightly narrowed anteriorly, posterior part distinctly narrower than anterior. Surface glabrous, median furrow distinct, visible in whole pronotal length. Propleura visible from dorsal aspect in basal two-thirds. Anterior angles of pronotum not protruding, posterior angles obtuse. Lateral furrows well developed, deep, with one pair of anterolateral setae, situated in the apical fifth of pronotal length.</paragraph>
<paragraph pageId="0" pageNumber="1">
Elytra subovate elongate, distinctly longer than wide, index EL/EW 1.74 (PT)-1.85 (HT), with maximum width in posterior third; elytral surface glabrous, without pubescence; striae absent. Stria 3 with 4-5 (3-4 discal and one preapical) macrochetae and 3 or 4 microchetae situated between macrochetae, stria 5 with 4 or 5 microchetae (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Humeral group of umbilicate pores not aggregated, first anterior pore of humeral group isolated and situated at level of the first discal seta.
</paragraph>
<paragraph pageId="0" pageNumber="1">Legs long, slender, densely pubescent. First two tarsomeres of male protarsi distinctly dilated and protracted at their internal margins. Tarsal claws very long and slender, without traces of denticulation on their internal sides.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male genitalia. Aedeagus (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) 0.58 mm long, relatively robust, regularly wide, lacking apical constriction, laterally flattened. Parameres relatively long and slender, longer than half of the length of aedeagus. Apex very short, tip obtuse. Each paramere at apex with three setae, two long, and one short.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Female genitalia as in Figure 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="2551941" doi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Male and female genitalia of 
<taxonomicName authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella</emphasis>
</taxonomicName>
representatives: aedeagus of 
<taxonomicName class="Insecta" genus="D." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. lukici" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">D. lukici</emphasis>
</taxonomicName>
sp. n., dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">(a)</emphasis>
; and lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">(b)</emphasis>
; female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of 
<taxonomicName class="Insecta" genus="D." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. lukici" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">D. lukici</emphasis>
</taxonomicName>
sp. n. 
<emphasis bold="true" pageId="0" pageNumber="1">(c)</emphasis>
; and 
<taxonomicName class="Insecta" genus="D." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. nonveilleri" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis italics="true" pageId="0" pageNumber="1">D. nonveilleri</emphasis>
</taxonomicName>
, lateral view of male aedeagus with parameres detached 
<emphasis bold="true" pageId="0" pageNumber="1">(d)</emphasis>
(illustration by Fedor 
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Patronymic, dedicated to our dear friend Marko 
<normalizedToken originalValue="Lukić">Lukic</normalizedToken>
(Zagreb, Croatia), enthusiastic speleologist and speleobiologist, taxonomic specialist on subterranean 
<taxonomicName class="Insecta" genus="D." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Collembola</taxonomicName>
, and collector of the first specimen of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. is closely related to the type species of the genus, 
<taxonomicName authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
. However, these two species can be easily recognized using the following key:
</paragraph>
<paragraph pageId="0" pageNumber="1">
<table inLine="true" pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1(2)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head with 2 posterior supraorbital setiferous punctures on each side, which are very close to each other. Elytra with maximum width in the posterior third. Putative stria 3 with 4 or 5 macrochetae and 4-6 microchetae, putative stria 5 with 3 or 4 microchetae (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Pronotum narrower, index PL/PW: 1.75-1.83, anterior angles not protruding. Aedeagus (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) shorter, more robust. L: 5.5-6.0 mm. Croatia, Mt Biokovo
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2(1)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head with only 2 posterior supraorbital setiferous puncture on each side. Elytra with maximum width in middle. Putative stria 3 with 3 macrochetae and 2 microchetae between each macrochetae, putative stria 5 with 1 microcheta in basal fourth (see Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Pronotum wider, index PL/PW: 1.43-1.44, anterior angles slightly protruding. Aedeagus (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) longer and slenderer. L: 5.4-5.8 mm. Croatia, Mt Mosor
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
, 2008
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
So far this species is known from the two pits on Mt Biokovo, the type locality, Biokovka, Golubinjak and Pretnerova jama, Lokva. All three specimens were found in deeper parts of the caves, attaining depths of 120 to 300 m. They were all found walking on the 
<normalizedToken originalValue="“moonmilk”">&quot;moonmilk&quot;</normalizedToken>
, a white, pastelite material consisting of microbiologically transformed microcrystalline calcites with high water content (60-90%), and near the cave hygropetric.
</paragraph>
<paragraph pageId="0" pageNumber="1">Associated subterranean coleopteran fauna observed in the pits:</paragraph>
<paragraph pageId="0" pageNumber="1">1 Biokovka, Golubinjak, Biokovo, Croatia:</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="? arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Miller" baseAuthorityYear="1861" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neotrechus dalmatinus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalmatinus">
<emphasis italics="true" pageId="0" pageNumber="1">Neotrechus dalmatinus</emphasis>
</taxonomicName>
(Miller, 1861)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
: 
<taxonomicName authorityName="Kirby" authorityYear="1837" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cholevinae">Cholevinae</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Leptomeson" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Leptomeson biokovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis italics="true" pageId="0" pageNumber="1">Leptomeson biokovensis</emphasis>
</taxonomicName>
Giachino, 
<normalizedToken originalValue="Bregović">Bregovic</normalizedToken>
&amp; 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
, 2012
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Radziella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Radziella styx" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="styx">
<emphasis italics="true" pageId="0" pageNumber="1">Radziella styx</emphasis>
</taxonomicName>
Casale &amp; 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
, 1988
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Speoplanes giganteus subsp. biocovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis italics="true" pageId="0" pageNumber="1">Speoplanes giganteus biocovensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
, 1934
</paragraph>
<paragraph pageId="0" pageNumber="1">2 Pretnerova jama, Lokva, Biokovo, Croatia:</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="? arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Monguzzi" authorityYear="1993" class="Insecta" family="Carabidae" genus="Dalmataphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Dalmataphaenops chiarae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis italics="true" pageId="0" pageNumber="1">Dalmataphaenops chiarae</emphasis>
</taxonomicName>
Monguzzi, 1993
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="? arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1834" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Schaum" baseAuthorityYear="1858" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Laemostenus cavicola" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">Laemostenus cavicola</emphasis>
</taxonomicName>
(Schaum, 1858)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
: 
<taxonomicName authorityName="Kirby" authorityYear="1837" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cholevinae">Cholevinae</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Leptomeson" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Leptomeson biokovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis italics="true" pageId="0" pageNumber="1">Leptomeson biokovensis</emphasis>
</taxonomicName>
Giachino, 
<normalizedToken originalValue="Bregović">Bregovic</normalizedToken>
&amp; 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
, 2012
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Radziella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Radziella styx" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="styx">
<emphasis italics="true" pageId="0" pageNumber="1">Radziella styx</emphasis>
</taxonomicName>
Casale &amp; 
<normalizedToken originalValue="Jalžić">Jalzic</normalizedToken>
, 1988
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Speoplanes giganteus subsp. biocovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis italics="true" pageId="0" pageNumber="1">Speoplanes giganteus biocovensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
, 1934
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
: 
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
gen.
</paragraph>
</subSubSection>
</treatment>
</document>