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<document id="F0A249E77BB3913FBD56FA73796C8542" ID-DOI="10.1080/00222933.2019.1646338" ID-GBIF-Dataset="9195a7a3-87bd-40a9-9165-2b2363b953b2" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673704" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582028089786" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Eguchi, Katsuyuki &amp; Hwang, Ui W." docDate="2019" docId="2C5787C93D7B5A6DFE4433D671FCFA34" docLanguage="en" docName="JNATHIST.53.21–22.1369–1384.pdf.imf" docOrigin="Journal of Natural History 53 (21)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hyleoglomeris fanxipan Nguyen ı Katsuyuki Eguchi &amp; Hwang 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="1378" masterDocId="D06EFFB13D7D5A67FF8D3612702EFFDB" masterDocTitle="Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam" masterLastPageNumber="1384" masterPageNumber="1369" pageNumber="1374" updateTime="1698719133712" updateUser="plazi">
<mods:mods id="20512CBDA3563AD9342C60F485CB5A84" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="83DC0B261FF41E546CC81B52D0FAAC4B">
<mods:title id="9F759BB1E3DA49567975DA657CD22CE9">Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="2E07047065B4465D0E89C84BFD59EAAB" type="personal">
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<mods:namePart id="E099DE9948E050E497086E7382F2BAB6">Nguyen, Anh D.</mods:namePart>
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<mods:role id="16595AEA45CA0511EB59C0C3A316A490">
<mods:roleTerm id="2FE85C060ABA1DCAFC9FFCF90C6887CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2369D36DD8DE714C51816978ACC20239">Eguchi, Katsuyuki</mods:namePart>
</mods:name>
<mods:name id="B1938F00D17580B70823C218993A0B1A" type="personal">
<mods:role id="59B07EE91B31ED6D9E859D4E817FDA55">
<mods:roleTerm id="DFA95C344DB7CA4A4972B99BF61F282D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="76EA2425E7BBD4B91403BE641D690230">Hwang, Ui W.</mods:namePart>
</mods:name>
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<mods:title id="A30F5E025C45442628AB2B72BFBB6DF0">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="1A52E399BA9C66C4EF4B49F03944EAE2">
<mods:date id="314F7E8B10606FB4163F3DE686B3782E">2019</mods:date>
<mods:detail id="AF5413DBA5AB43518493A333DDB32C2B" type="pubDate">
<mods:number id="36E0B574236E53123F5C1A247310A23A">2019-08-20</mods:number>
</mods:detail>
<mods:detail id="AA83A41D838D106F4867E47C5E4FCC02" type="volume">
<mods:number id="F473D845D61D19CA7D86353B5AA29938">53</mods:number>
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<mods:number id="F3A662F5C8884C312189A84BAD1F6B0D">21</mods:number>
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<mods:start id="1473F22FD604A85CE694B00EF29C10F8">1369</mods:start>
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<mods:classification id="151318F5B2795F1B12FD5A0310D51A75">journal article</mods:classification>
<mods:identifier id="BE12AC5C2BF611F4AD6F9A87F7424740" type="DOI">10.1080/00222933.2019.1646338</mods:identifier>
<mods:identifier id="195732B3D650BDEC9BBE1038B44C1160" type="GBIF-Dataset">9195a7a3-87bd-40a9-9165-2b2363b953b2</mods:identifier>
<mods:identifier id="A89C0426C05F6936E560C58898D61B19" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="F66F12B085F98F035FA7F4BD7A367060" type="Zenodo-Dep">3673704</mods:identifier>
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<treatment id="2C5787C93D7B5A6DFE4433D671FCFA34" ID-DOI="http://doi.org/10.5281/zenodo.3679863" ID-GBIF-Taxon="162162043" ID-Zenodo-Dep="3679863" LSID="urn:lsid:plazi:treatment:2C5787C93D7B5A6DFE4433D671FCFA34" httpUri="http://treatment.plazi.org/id/2C5787C93D7B5A6DFE4433D671FCFA34" lastPageId="10" lastPageNumber="1378" pageId="6" pageNumber="1374">
<subSubSection id="ECE465543D7B5A61FE4433D67374FA06" box="[457,858,1476,1501]" pageId="6" pageNumber="1374" type="nomenclature">
<paragraph id="A44136DF3D7B5A61FE4433D67374FA06" blockId="6.[457,858,1476,1536]" box="[457,858,1476,1501]" pageId="6" pageNumber="1374">
<heading id="FF0981B33D7B5A61FE4433D67374FA06" box="[457,858,1476,1501]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="1374" reason="2">
<taxonomicName id="63FE4D5C3D7B5A61FE4433D672C0FA06" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[457,750,1476,1501]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="fanxipan" status="sp. nov.">
<emphasis id="968AEACD3D7B5A61FE4433D672C0FA06" bold="true" box="[457,750,1476,1501]" italics="true" pageId="6" pageNumber="1374">Hyleoglomeris fanxipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D7B5A61FD7533D67374FA06" box="[760,858,1476,1501]" pageId="6" pageNumber="1374" rank="species">
<emphasis id="968AEACD3D7B5A61FD7533D6737AFA06" bold="true" box="[760,852,1476,1501]" pageId="6" pageNumber="1374">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7B5A61FDB333F572C8F9DB" box="[574,742,1511,1536]" pageId="6" pageNumber="1374" type="description">
<paragraph id="A44136DF3D7B5A61FDB333F572C8F9DB" blockId="6.[457,858,1476,1536]" box="[574,742,1511,1536]" pageId="6" pageNumber="1374">
<heading id="FF0981B33D7B5A61FDB333F572C8F9DB" box="[574,742,1511,1536]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="1374" reason="5">
(
<figureCitation id="3CC52A5A3D7B5A61FDCB33F572F0F9DB" box="[582,734,1511,1536]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStart-4="Figure 10" captionStartId-0="8.[160,225,838,861]" captionStartId-1="9.[160,225,920,943]" captionStartId-2="10.[160,225,980,1003]" captionStartId-3="11.[160,225,960,983]" captionStartId-4="11.[160,225,1523,1546]" captionTargetBox-0="[170,1148,149,806]" captionTargetBox-1="[165,1152,145,887]" captionTargetBox-2="[166,1150,145,935]" captionTargetBox-3="[258,1058,142,928]" captionTargetBox-4="[181,1135,1103,1489]" captionTargetId-0="figure@8.[163,1153,142,806]" captionTargetId-1="figure@9.[163,1153,142,889]" captionTargetId-2="figure@10.[163,1153,142,948]" captionTargetId-3="figure@11.[258,1058,142,928]" captionTargetId-4="figure@11.[178,1138,1100,1492]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="11" captionText-0="Figure 6. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). A–C, whole body, dorsal view (A), ventral view (B), lateral view (C); collum, anterior view (D)." captionText-1="Figure 7. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). Ocelli &amp; Tömösváry organ (A); Antenna (B); Thoracic shield (C); Anal shield (D)." captionText-2="Figure 8. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673 H). Legs 17, anterior view (A); telopods, posterior view (B); left telopod, posterior view (C); trichosteles, posterior view (D)." captionText-3="Figure 9. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). Telopods, anterior view (A); left telopod, anterior view (B); femoral process and tarsi, anterior view (C); syncoxital horn and lobe, posterior view (D)." captionText-4="Figure 10. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). Syncoxital horns and lobe, posterior view (A), anterior view (B)." figureDoi-0="http://doi.org/10.5281/zenodo.3673716" figureDoi-1="http://doi.org/10.5281/zenodo.3673718" figureDoi-2="http://doi.org/10.5281/zenodo.3673720" figureDoi-3="http://doi.org/10.5281/zenodo.3673722" figureDoi-4="http://doi.org/10.5281/zenodo.3673724" httpUri-0="https://zenodo.org/record/3673716/files/figure.png" httpUri-1="https://zenodo.org/record/3673718/files/figure.png" httpUri-2="https://zenodo.org/record/3673720/files/figure.png" httpUri-3="https://zenodo.org/record/3673722/files/figure.png" httpUri-4="https://zenodo.org/record/3673724/files/figure.png" pageId="6" pageNumber="1374">
Figures 6
<emphasis id="968AEACD3D7B5A61FD3E33F572EFF9DB" box="[691,705,1511,1536]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
10
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7B5A60FF2D302F7125FB7D" lastPageId="7" lastPageNumber="1375" pageId="6" pageNumber="1374" type="materials_examined">
<paragraph id="A44136DF3D7B5A61FF2D302F71AEF98C" blockId="6.[160,1155,1597,1727]" box="[160,384,1597,1623]" pageId="6" pageNumber="1374">
<emphasis id="968AEACD3D7B5A61FF2D302F71AEF98C" bold="true" box="[160,384,1597,1623]" italics="true" pageId="6" pageNumber="1374">Material examined</emphasis>
</paragraph>
<paragraph id="A44136DF3D7B5A61FF2D307272F4F964" blockId="6.[160,1155,1597,1727]" pageId="6" pageNumber="1374">
<materialsCitation id="14963C823D7B5A61FF2D307272F4F964" ID-GBIF-Occurrence="2565952785" collectingDate="2017-01" collectorName="Anh D. Nguyen &amp; K. Eguchi." county="One" elevation="900" location="Pai Hoang Lien National Parki" municipality="Sa Pai Hoang Lien National" pageId="6" pageNumber="1374" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="968AEACD3D7B5A61FF2D3072713CF9A1" bold="true" box="[160,274,1632,1658]" italics="true" pageId="6" pageNumber="1374">
<typeStatus id="7B45887D3D7B5A61FF2D30727123F9A1" box="[160,269,1632,1658]" pageId="6" pageNumber="1374" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="4D204E533D7B5A61FEAC3070717EF9A1" box="[289,336,1634,1658]" pageId="6" pageNumber="1374">One</collectingCounty>
male (
<emphasis id="968AEACD3D7B5A61FE2D3070727BF9A1" bold="true" box="[416,597,1634,1658]" pageId="6" pageNumber="1374">IEBR-Myr 673H</emphasis>
)ı
<collectingRegion id="663AF83D3D7B5A61FDE63070732AF9A1" box="[619,772,1634,1658]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="1374">Lao Cai Prov.</collectingRegion>
ı
<collectingMunicipality id="4425ACA53D7B5A61FC833070746AF9A1" box="[782,1092,1634,1658]" pageId="6" pageNumber="1374">
Sa
<location id="A12160043D7B5A61FCBC3070746AF9A1" LSID="urn:lsid:plazi:treatment:2C5787C93D7B5A6DFE4433D671FCFA34:A12160043D7B5A61FCBC3070746AF9A1" box="[817,1092,1634,1658]" county="One" municipality="Sa Pai Hoang Lien National" name="Pai Hoang Lien National Parki" pageId="6" pageNumber="1374" stateProvince="Lao Cai">Paı Hoang Lien National</location>
</collectingMunicipality>
Parkı short bamboo forestsı
<location id="A12160043D7B5A61FE3C3097726DF946" LSID="urn:lsid:plazi:treatment:2C5787C93D7B5A6DFE4433D671FCFA34:A12160043D7B5A61FE3C3097726DF946" box="[433,579,1669,1693]" county="One" municipality="Sa Pai Hoang Lien National" name="Fanxifan Mt." pageId="6" pageNumber="1374" stateProvince="Lao Cai">Fanxifan Mt.</location>
ı 2ı800
<emphasis id="968AEACD3D7B5A61FD02309672B2F947" box="[655,668,1668,1692]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
2ı
<quantity id="63069B3A3D7B5A61FD20309772D2F946" box="[685,764,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="1374" unit="m" value="900.0">
<elevation id="2FD3D1EC3D7B5A61FD20309772D2F946" box="[685,764,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="1374" unit="m" value="900.0">900 m</elevation>
</quantity>
alt.ı 103°52
<emphasis id="968AEACD3D7B5A61FC06309673BFF947" bold="true" box="[907,913,1668,1692]" italics="true" pageId="6" pageNumber="1374">ʹ</emphasis>
42
<emphasis id="968AEACD3D7B5A61FC2030967396F947" bold="true" box="[941,952,1668,1692]" italics="true" pageId="6" pageNumber="1374">″</emphasis>
Eı 22°21
<emphasis id="968AEACD3D7B5A61FB9A30967433F947" bold="true" box="[1047,1053,1668,1692]" italics="true" pageId="6" pageNumber="1374">ʹ</emphasis>
22
<emphasis id="968AEACD3D7B5A61FBB53096746DF947" bold="true" box="[1080,1091,1668,1692]" italics="true" pageId="6" pageNumber="1374">″</emphasis>
Nı 23 September
<date id="D040101F3D7B5A61FEA930B5714CF964" box="[292,354,1703,1727]" pageId="6" pageNumber="1374" value="2017-01">
<collectingDate id="C004E9F73D7B5A61FEA930B5714CF964" box="[292,354,1703,1727]" pageId="6" pageNumber="1374" value="2017-01">2017ı</collectingDate>
</date>
leg.
<collectorName id="090B53093D7B5A61FE1730B57267F964" box="[410,585,1703,1727]" pageId="6" pageNumber="1374">Anh D. Nguyen</collectorName>
&amp;
<collectorName id="090B53093D7B5A61FDE730B572F4F964" box="[618,730,1703,1727]" pageId="6" pageNumber="1374">K. Eguchi.</collectorName>
</materialsCitation>
</paragraph>
<caption id="F08166573D7A5A60FF2D35C070D4FBFF" ID-DOI="http://doi.org/10.5281/zenodo.3673714" ID-Zenodo-Dep="3673714" httpUri="https://zenodo.org/record/3673714/files/figure.png" pageId="7" pageNumber="1375" startId="7.[160,225,978,1001]" targetBox="[163,1153,142,936]" targetPageId="7">
<paragraph id="A44136DF3D7A5A60FF2D35C070D4FBFF" blockId="7.[160,1155,978,1060]" pageId="7" pageNumber="1375">
<emphasis id="968AEACD3D7A5A60FF2D35C070D2FC32" bold="true" box="[160,252,978,1001]" pageId="7" pageNumber="1375">Figure 5.</emphasis>
<taxonomicName id="63FE4D5C3D7A5A60FE8B35C07234FC32" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[262,538,978,1001]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="7" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D7A5A60FE8B35C071DAFC32" box="[262,500,978,1001]" italics="true" pageId="7" pageNumber="1375">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D7A5A60FE7035C07234FC32" box="[509,538,978,1001]" pageId="7" pageNumber="1375" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
nov., paratype (IEBR-Myr 672P). Tibia of left telopod, ventral view (A); lateral surface (B); syncoxital lobe and horns, anterior view (C); distal part of the syncoxital horn (D).
</paragraph>
</caption>
<paragraph id="A44136DF3D7A5A60FF2D327B7125FB7D" blockId="7.[160,1155,1129,1190]" pageId="7" pageNumber="1375">
<emphasis id="968AEACD3D7A5A60FF2D327B7132FB58" bold="true" box="[160,284,1129,1155]" italics="true" pageId="7" pageNumber="1375">
<typeStatus id="7B45887D3D7A5A60FF2D327B7139FB58" box="[160,279,1129,1155]" pageId="7" pageNumber="1375" type="paratype">Paratypes</typeStatus>
.
</emphasis>
One maleı three females (
<emphasis id="968AEACD3D7A5A60FDE43279730FFB58" bold="true" box="[617,801,1131,1155]" pageId="7" pageNumber="1375">IEBR-Myr 673P</emphasis>
)ı with the same data as the holotype.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7A5A60FF2D32F3724BFA10" pageId="7" pageNumber="1375" type="diagnosis">
<paragraph id="A44136DF3D7A5A60FF2D32F3713AFB20" blockId="7.[160,1156,1249,1483]" box="[160,276,1249,1275]" pageId="7" pageNumber="1375">
<emphasis id="968AEACD3D7A5A60FF2D32F3713AFB20" bold="true" box="[160,276,1249,1275]" italics="true" pageId="7" pageNumber="1375">Diagnosis</emphasis>
</paragraph>
<paragraph id="A44136DF3D7A5A60FF2D3314724BFA10" blockId="7.[160,1156,1249,1483]" pageId="7" pageNumber="1375">
An entirely blackish or greenish-black body; ocelli 7 + 1; thoracic shield with 10
<emphasis id="968AEACD3D7A5A60FBD733147449FAC5" box="[1114,1127,1286,1310]" italics="true" pageId="7" pageNumber="1375">–</emphasis>
12 vague transverse striae (but only three cross); telopods long and robust; syncoxital lobe roundly trapeziform; both prefemoral and femoral trichosteles present; femoral process directed mesoventrallyı strongly rugulose and folded in anterior side; tip strongly recurved; posterior surface of tip densely micropapillate; tibia without modifications; tarsus curvedı bearing two apical setae.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7A5A60FF2D30157107F9BD" pageId="7" pageNumber="1375" type="etymology">
<paragraph id="A44136DF3D7A5A60FF2D30157133F9FA" blockId="7.[160,1155,1543,1638]" box="[160,285,1543,1569]" pageId="7" pageNumber="1375">
<emphasis id="968AEACD3D7A5A60FF2D30157133F9FA" bold="true" box="[160,285,1543,1569]" italics="true" pageId="7" pageNumber="1375">Etymology</emphasis>
</paragraph>
<paragraph id="A44136DF3D7A5A60FF2D30397107F9BD" blockId="7.[160,1155,1543,1638]" pageId="7" pageNumber="1375">
Named (noun in apposition) after the Fanxipan Mountain where the
<typeStatus id="7B45887D3D7A5A60FC5D3039742CF998" box="[976,1026,1579,1603]" pageId="7" pageNumber="1375">type</typeStatus>
specimens were found.
</paragraph>
</subSubSection>
<caption id="F08166573D755A6FFF2D35547339FCA1" ID-DOI="http://doi.org/10.5281/zenodo.3673716" ID-Zenodo-Dep="3673716" httpUri="https://zenodo.org/record/3673716/files/figure.png" pageId="8" pageNumber="1376" startId="8.[160,225,838,861]" targetBox="[170,1148,149,806]" targetPageId="8">
<paragraph id="A44136DF3D755A6FFF2D35547339FCA1" blockId="8.[160,1155,837,890]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D355470D2FC86" bold="true" box="[160,252,838,861]" pageId="8" pageNumber="1376">Figure 6.</emphasis>
<taxonomicName id="63FE4D5C3D755A6FFE89355471C9FC86" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[260,487,838,861]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="8" pageNumber="1376" phylum="Arthropoda" rank="species" species="fanxipan" status="sp. nov.">
<emphasis id="968AEACD3D755A6FFE89355471C9FC86" box="[260,487,838,861]" italics="true" pageId="8" pageNumber="1376">Hyleoglomeris fanxipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D755A6FFE623554726EFC86" box="[495,576,838,861]" pageId="8" pageNumber="1376" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IEBR-Myr 673H). A–C, whole body, dorsal view (A), ventral view (B), lateral view (C); collum, anterior view (D).
</paragraph>
</caption>
<subSubSection id="ECE465543D755A6DFF2D35DF71E1FB01" lastPageId="10" lastPageNumber="1378" pageId="8" pageNumber="1376" type="description">
<paragraph id="A44136DF3D755A6FFF2D35DF7108FC3C" blockId="8.[160,1155,973,1069]" box="[160,294,973,999]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D35DF7108FC3C" bold="true" box="[160,294,973,999]" italics="true" pageId="8" pageNumber="1376">Description</emphasis>
</paragraph>
<paragraph id="A44136DF3D755A6FFF2D35E27124FBF6" blockId="8.[160,1155,973,1069]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D35E270FAFBD1" bold="true" box="[160,212,1008,1034]" italics="true" pageId="8" pageNumber="1376">Size.</emphasis>
<typeStatus id="7B45887D3D755A6FFF6835E07162FBD1" box="[229,332,1010,1034]" pageId="8" pageNumber="1376" type="holotype">Holotype</typeStatus>
male (
<emphasis id="968AEACD3D755A6FFE1335E0727AFBD1" bold="true" box="[414,596,1010,1034]" pageId="8" pageNumber="1376">IEBR-Myr 673H</emphasis>
). Width of second tergum
<quantity id="63069B3A3D755A6FFC1D35E073C4FBD1" box="[912,1002,1010,1034]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="1376" unit="mm" value="4.9">4.9 mm</quantity>
; body length
<quantity id="63069B3A3D755A6FFF2D32077128FBF6" box="[160,262,1045,1069]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="1376" unit="mm" value="11.8">11.8 mm</quantity>
.
</paragraph>
<paragraph id="A44136DF3D755A6FFF2D327372F8FB46" blockId="8.[160,1156,1121,1181]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D327370DBFBA0" bold="true" box="[160,245,1121,1147]" italics="true" pageId="8" pageNumber="1376">Colour.</emphasis>
Entirely blackish or greenish-black except for two distolateral brownish-yellow marbled oval spots and lateral margins yellowish.
</paragraph>
<paragraph id="A44136DF3D755A6FFF2D32C370C6FA88" blockId="8.[160,1155,1233,1363]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D32C370CBFB30" bold="true" box="[160,229,1233,1259]" italics="true" pageId="8" pageNumber="1376">Head.</emphasis>
Ocelli 7 + 1; lenses convexı shining black as background of the head. Tömösváry organs transverse ovalı ca. 2
<emphasis id="968AEACD3D755A6FFE6A32E471DAFAD5" box="[487,500,1270,1294]" italics="true" pageId="8" pageNumber="1376">–</emphasis>
2.2X wider than long. Antenna clavate apically; antennomere six largeı ca. 2.5
<emphasis id="968AEACD3D755A6FFE1D330A71B3FAEB" box="[400,413,1304,1328]" italics="true" pageId="8" pageNumber="1376">–</emphasis>
3.0X longer than wide. Antennal tip with four largeı apical sensory cones.
</paragraph>
<paragraph id="A44136DF3D755A6FFF2D3395717EF994" blockId="8.[160,1156,1415,1615]" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D339570CFFA7A" bold="true" box="[160,225,1415,1441]" italics="true" pageId="8" pageNumber="1376">Body.</emphasis>
Collum semicircuları with a very large-marbled yellowish-brown spot in the middleı and two transverse striae; other parts of collum brownish-black. Thoracic shield (= tergum 2) with a narrow hyposchismı reaching caudal margin; with 10
<emphasis id="968AEACD3D755A6FFBD733DC7449FA3D" box="[1114,1127,1486,1510]" italics="true" pageId="8" pageNumber="1376">–</emphasis>
12 vague transverse striae; only 3 of which cross the dorsum. Other terga with 3
<emphasis id="968AEACD3D755A6FFBE533E3745BF9D2" box="[1128,1141,1521,1545]" italics="true" pageId="8" pageNumber="1376">–</emphasis>
4 short vague transverse striae. Anal shield strongly rounded emarginate mediocaudally.
</paragraph>
<paragraph id="A44136DF3D755A6EFF2D3091714BFB95" blockId="8.[160,1155,1667,1727]" lastBlockId="9.[160,1155,1044,1102]" lastPageId="9" lastPageNumber="1377" pageId="8" pageNumber="1376">
<emphasis id="968AEACD3D755A6FFF2D309170F5F946" bold="true" box="[160,219,1667,1693]" italics="true" pageId="8" pageNumber="1376">Legs.</emphasis>
Leg-pair 17 strongly reducedı 4-segmented; coxite with a rounded outer lobeı with a setiferous tubercle distomesally; podomere 1 with a small distomesal setiferous knob. Leg-pair 18 less strongly reducedı 4-segmentedı with a simple syncoxital notch.
</paragraph>
<caption id="F08166573D745A6EFF2D358A72ADFC16" ID-DOI="http://doi.org/10.5281/zenodo.3673718" ID-Zenodo-Dep="3673718" httpUri="https://zenodo.org/record/3673718/files/figure.png" pageId="9" pageNumber="1377" startId="9.[160,225,920,943]" targetBox="[165,1152,145,887]" targetPageId="9">
<paragraph id="A44136DF3D745A6EFF2D358A72ADFC16" blockId="9.[160,1156,920,973]" pageId="9" pageNumber="1377">
<emphasis id="968AEACD3D745A6EFF2D358A70D2FC74" bold="true" box="[160,252,920,943]" pageId="9" pageNumber="1377">Figure 7.</emphasis>
<taxonomicName id="63FE4D5C3D745A6EFE89358A71C8FC74" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[260,486,920,943]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="9" pageNumber="1377" phylum="Arthropoda" rank="species" species="fanxipan" status="sp. nov.">
<emphasis id="968AEACD3D745A6EFE89358A71C8FC74" box="[260,486,920,943]" italics="true" pageId="9" pageNumber="1377">Hyleoglomeris fanxipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D745A6EFE62358A726EFC74" box="[495,576,920,943]" pageId="9" pageNumber="1377" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IEBR-Myr 673H). Ocelli &amp; Tömösváry organ (A); Antenna (B); Thoracic shield (C); Anal shield (D).
</paragraph>
</caption>
<paragraph id="A44136DF3D745A6EFF2D3298711FFA07" blockId="9.[160,1156,1162,1500]" pageId="9" pageNumber="1377">
<emphasis id="968AEACD3D745A6EFF2D329871E1FB7F" bold="true" box="[160,463,1162,1188]" pageId="9" pageNumber="1377">
<emphasis id="968AEACD3D745A6EFF2D32987122FB7F" bold="true" box="[160,268,1162,1188]" italics="true" pageId="9" pageNumber="1377">Genitalia</emphasis>
(
<figureCitation id="3CC52A5A3D745A6EFEAD329971EEFB7F" box="[288,448,1163,1188]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStart-2="Figure 10" captionStartId-0="10.[160,225,980,1003]" captionStartId-1="11.[160,225,960,983]" captionStartId-2="11.[160,225,1523,1546]" captionTargetBox-0="[166,1150,145,935]" captionTargetBox-1="[258,1058,142,928]" captionTargetBox-2="[181,1135,1103,1489]" captionTargetId-0="figure@10.[163,1153,142,948]" captionTargetId-1="figure@11.[258,1058,142,928]" captionTargetId-2="figure@11.[178,1138,1100,1492]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionText-0="Figure 8. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673 H). Legs 17, anterior view (A); telopods, posterior view (B); left telopod, posterior view (C); trichosteles, posterior view (D)." captionText-1="Figure 9. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). Telopods, anterior view (A); left telopod, anterior view (B); femoral process and tarsi, anterior view (C); syncoxital horn and lobe, posterior view (D)." captionText-2="Figure 10. Hyleoglomeris fanxipan sp. nov., holotype (IEBR-Myr 673H). Syncoxital horns and lobe, posterior view (A), anterior view (B)." figureDoi-0="http://doi.org/10.5281/zenodo.3673720" figureDoi-1="http://doi.org/10.5281/zenodo.3673722" figureDoi-2="http://doi.org/10.5281/zenodo.3673724" httpUri-0="https://zenodo.org/record/3673720/files/figure.png" httpUri-1="https://zenodo.org/record/3673722/files/figure.png" httpUri-2="https://zenodo.org/record/3673724/files/figure.png" pageId="9" pageNumber="1377">Figures 8–10</figureCitation>
).
</emphasis>
Telopods long and robust. Syncoxital lobe (
<emphasis id="968AEACD3D745A6EFC79329E7438FB7F" bold="true" box="[1012,1046,1164,1188]" pageId="9" pageNumber="1377">syl</emphasis>
) roundly trapeziform. Syncoxital horns (
<emphasis id="968AEACD3D745A6EFD8F32BC7202FB1D" bold="true" box="[514,556,1198,1222]" pageId="9" pageNumber="1377">syh</emphasis>
) highly elevatedı directed ventrallyı with numerous setae; tips truncated. Prefemur (
<emphasis id="968AEACD3D745A6EFDBA32C37247FB32" bold="true" box="[567,617,1233,1257]" pageId="9" pageNumber="1377">pref</emphasis>
) longı densely micropapillate laterally and posterolaterallyı with a longı prominentı finger-shaped trichostele (
<emphasis id="968AEACD3D745A6EFC5032E6743EFAD7" bold="true" box="[989,1040,1268,1292]" pageId="9" pageNumber="1377">pret</emphasis>
). Femora (
<emphasis id="968AEACD3D745A6EFF25330470EEFAF5" bold="true" box="[168,192,1302,1326]" pageId="9" pageNumber="1377">fe</emphasis>
) broadı enlarged distallyı also bearing a longı prominentı finger-shaped trichostele (
<emphasis id="968AEACD3D745A6EFF6F332B712AFA8A" bold="true" box="[226,260,1337,1361]" pageId="9" pageNumber="1377">fet</emphasis>
)ı and with densely micropapillate apical cingulum. Femoral process (
<emphasis id="968AEACD3D745A6EFBEF332B7452FA8A" bold="true" box="[1122,1148,1337,1361]" pageId="9" pageNumber="1377">fp</emphasis>
) directed mesoventrallyı strongly rugulose and folded in anterior side; tip strongly recurved; posterior surface of tip densely micropapillate. Tibia (
<emphasis id="968AEACD3D745A6EFC1D336C738CFA4D" bold="true" box="[912,930,1406,1430]" pageId="9" pageNumber="1377">ti</emphasis>
) without modificationsı but the tibial process (
<emphasis id="968AEACD3D745A6EFD8A33B3720FFA62" bold="true" box="[519,545,1441,1465]" pageId="9" pageNumber="1377">tb</emphasis>
) short and stout. Tarsus (
<emphasis id="968AEACD3D745A6EFCEE33B37355FA62" bold="true" box="[867,891,1441,1465]" pageId="9" pageNumber="1377">ta</emphasis>
) curvedı bearing two apical setae.
</paragraph>
<paragraph id="A44136DF3D745A6EFF2D300570DCF942" blockId="9.[160,1155,1559,1689]" pageId="9" pageNumber="1377">
<emphasis id="968AEACD3D745A6EFF2D3005713BF9EA" bold="true" box="[160,277,1559,1585]" italics="true" pageId="9" pageNumber="1377">Variation.</emphasis>
In other specimensı only 10
<emphasis id="968AEACD3D745A6EFDEB300B725DF9EA" box="[614,627,1561,1585]" italics="true" pageId="9" pageNumber="1377">–</emphasis>
11 vague transverse striae are present on the dorsal surface of the thoracic shield. Width of second tergum
<quantity id="63069B3A3D745A6EFCE1302E73DEF98F" box="[876,1008,1595,1620]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.2" metricValueMin="5.4" pageId="9" pageNumber="1377" unit="mm" value="5.8" valueMax="6.2" valueMin="5.4">
5.4
<emphasis id="968AEACD3D745A6EFC00302973B4F988" box="[909,922,1595,1619]" italics="true" pageId="9" pageNumber="1377">–</emphasis>
6.2 mm
</quantity>
in the maleı
<quantity id="63069B3A3D745A6EFF2D304C710DF9AD" box="[160,291,1630,1654]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.9" metricValueMin="5.1" pageId="9" pageNumber="1377" unit="mm" value="5.5" valueMax="5.9" valueMin="5.1">
5.1
<emphasis id="968AEACD3D745A6EFF4C304C70E0F9AD" box="[193,206,1630,1654]" italics="true" pageId="9" pageNumber="1377">–</emphasis>
5.9 mm
</quantity>
in the female; body length 11.0
<emphasis id="968AEACD3D745A6EFD1F304C72B1F9AD" box="[658,671,1630,1654]" italics="true" pageId="9" pageNumber="1377">–</emphasis>
<quantity id="63069B3A3D745A6EFD12304C732CF9AD" box="[671,770,1630,1654]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="9" pageNumber="1377" unit="mm" value="11.4">11.4 mm</quantity>
in the maleı 12.0
<emphasis id="968AEACD3D745A6EFC45304C73FBF9AD" box="[968,981,1630,1654]" italics="true" pageId="9" pageNumber="1377">–</emphasis>
14.0 mm in the female.
</paragraph>
<caption id="F08166573D775A6DFF2D35C67414FBD3" ID-DOI="http://doi.org/10.5281/zenodo.3673720" ID-Zenodo-Dep="3673720" httpUri="https://zenodo.org/record/3673720/files/figure.png" pageId="10" pageNumber="1378" startId="10.[160,225,980,1003]" targetBox="[166,1150,145,935]" targetPageId="10">
<paragraph id="A44136DF3D775A6DFF2D35C67414FBD3" blockId="10.[160,1156,980,1032]" pageId="10" pageNumber="1378">
<emphasis id="968AEACD3D775A6DFF2D35C670D0FC30" bold="true" box="[160,254,980,1003]" pageId="10" pageNumber="1378">Figure 8.</emphasis>
<taxonomicName id="63FE4D5C3D775A6DFE8535C671C2FC30" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[264,492,980,1003]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="10" pageNumber="1378" phylum="Arthropoda" rank="species" species="fanxipan" status="sp. nov.">
<emphasis id="968AEACD3D775A6DFE8535C671C2FC30" box="[264,492,980,1003]" italics="true" pageId="10" pageNumber="1378">Hyleoglomeris fanxipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D775A6DFE7B35C67267FC30" box="[502,585,980,1003]" pageId="10" pageNumber="1378" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IEBR-Myr 673 H). Legs 17, anterior view (A); telopods, posterior view (B); left telopod, posterior view (C); trichosteles, posterior view (D).
</paragraph>
</caption>
<paragraph id="A44136DF3D775A6DFF2D324A7111FBA9" blockId="10.[160,1155,1112,1242]" box="[160,319,1112,1138]" pageId="10" pageNumber="1378">
<emphasis id="968AEACD3D775A6DFF2D324A7111FBA9" bold="true" box="[160,319,1112,1138]" italics="true" pageId="10" pageNumber="1378">DNA barcode</emphasis>
</paragraph>
<paragraph id="A44136DF3D775A6DFF2D326F71E1FB01" blockId="10.[160,1155,1112,1242]" pageId="10" pageNumber="1378">
COI barcode data (partial) is for the
<typeStatus id="7B45887D3D775A6DFDB5326F72B2FB4E" box="[568,668,1149,1173]" pageId="10" pageNumber="1378" type="paratype">paratype</typeStatus>
uploaded to GenBank under the accession number MH248039. The new species shares the identity of 87% with
<emphasis id="968AEACD3D775A6DFC3D328D7134FB01" italics="true" pageId="10" pageNumber="1378">
<taxonomicName id="63FE4D5C3D775A6DFC3D328D7451FB6C" box="[944,1151,1183,1207]" class="Diplopoda" family="Glomeridae" genus="Glomeris" kingdom="Animalia" order="Glomerida" pageId="10" pageNumber="1378" phylum="Arthropoda" rank="species" species="pustulata">Glomeris pustulata</taxonomicName>
,
<taxonomicName id="63FE4D5C3D775A6DFF2D32D07134FB01" box="[160,282,1218,1242]" class="Diplopoda" family="Glomeridae" genus="Glomeris" kingdom="Animalia" order="Glomerida" pageId="10" pageNumber="1378" phylum="Arthropoda" rank="species" species="connexa">G. connexa</taxonomicName>
</emphasis>
and
<taxonomicName id="63FE4D5C3D775A6DFEDA32D071E7FB01" authorityName="Latzel" authorityYear="1884" box="[343,457,1218,1242]" class="Diplopoda" family="Glomeridae" genus="Glomeris" kingdom="Animalia" order="Glomerida" pageId="10" pageNumber="1378" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="968AEACD3D775A6DFEDA32D071E7FB01" box="[343,457,1218,1242]" italics="true" pageId="10" pageNumber="1378">G. minima</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D775A6DFF2D331771FCFA34" pageId="10" pageNumber="1378" type="discussion">
<paragraph id="A44136DF3D775A6DFF2D33177128FAC4" blockId="10.[160,1156,1285,1519]" box="[160,262,1285,1311]" pageId="10" pageNumber="1378">
<emphasis id="968AEACD3D775A6DFF2D33177128FAC4" bold="true" box="[160,262,1285,1311]" italics="true" pageId="10" pageNumber="1378">Remarks</emphasis>
</paragraph>
<paragraph id="A44136DF3D775A6DFF2D333871FCFA34" blockId="10.[160,1156,1285,1519]" pageId="10" pageNumber="1378">
According to
<bibRefCitation id="C06F4B2E3D775A6DFEB633387213FA99" author="Golovatch SIi Geoffroy JJi Mauries JP" box="[315,573,1322,1346]" pageId="10" pageNumber="1378" pagination="887 - 915" refId="ref5134" refString="Golovatch SIi Geoffroy JJi Mauries JP. 2006. Review of the millipede genus Hyleoglomeris Verhoeffi 1910 (Diplopodai Glomeridai Glomeridae) i with descriptions of new species from caves in Southeast Asia. Zoosystema. 28 (4): 887 - 915." type="journal article" year="2006">Golovatch et al. (2006)</bibRefCitation>
ı the new species seems to belong to the
<emphasis id="968AEACD3D775A6DFB9033387455FA99" box="[1053,1147,1322,1346]" italics="true" pageId="10" pageNumber="1378">modesta</emphasis>
- group characterised by the dark background and lighter pattern/markings. This group consists of eight speciesı mainly distributed in the Himalayas of
<collectingCountry id="DCE9764F3D775A6DFC31337D73DDFA5C" box="[956,1011,1391,1415]" name="India" pageId="10" pageNumber="1378">India</collectingCountry>
and
<collectingCountry id="DCE9764F3D775A6DFBB1337D74AEFA5C" box="[1084,1152,1391,1415]" name="Nepal" pageId="10" pageNumber="1378">Nepal</collectingCountry>
. Howeverı the new species totally differs from the known
<emphasis id="968AEACD3D775A6DFCD533807398FA71" box="[856,950,1426,1450]" italics="true" pageId="10" pageNumber="1378">modesta</emphasis>
-group species in lacking axial spots on terga 2
<emphasis id="968AEACD3D775A6DFE7333A67225FA17" box="[510,523,1460,1484]" italics="true" pageId="10" pageNumber="1378">–</emphasis>
11ı and in the presence of two distolateral brownishyellow marbled oval spots.
</paragraph>
</subSubSection>
</treatment>
</document>
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