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<document id="56AD262359D55B958719636BE5B72491" ID-CLB-Dataset="7750" ID-DOI="10.11646/zootaxa.4938.4.1" ID-GBIF-Dataset="e5d1920d-c769-4aea-a8cb-237a0e58ae4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4574809" ID-ZooBank="6FEECE37-B2D3-4AE3-9878-CF212420AF9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614750655839" checkinUser="plazi" docAuthor="Babenko, Anatoly, Kuznetsova, Natalia, Nakamori, Taizo &amp; Shveenkova, Yulia" docDate="2021" docId="BF05878D232CFFFCFF498D9EFB26C9D6" docLanguage="en" docName="zootaxa.4938.4.1.pdf" docOrigin="Zootaxa 4938 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudachorutes morulifer Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="368" masterDocId="433CFFF52323FFEDFFDE8D09FF9DCA57" masterDocTitle="A review of Pseudachorutes Tullberg, 1871 (Collembola, Neanuridae) from the East Asia, with description of six new species" masterLastPageNumber="391" masterPageNumber="351" pageNumber="366" updateTime="1698931878726" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="AD8496E14B906112962E2A87A0C433C2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6E03E9B12314237F67CFF24995784D33">A review of Pseudachorutes Tullberg, 1871 (Collembola, Neanuridae) from the East Asia, with description of six new species</mods:title>
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<mods:name id="DB97926175FF455778BFCDD02146DB17" type="personal">
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<mods:roleTerm id="37E71D348AD882D3D4BC6AA09A352999">Author</mods:roleTerm>
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<mods:namePart id="021B4EE8787C6A9AC3BA3BBEA9676B73">Babenko, Anatoly</mods:namePart>
</mods:name>
<mods:name id="8EE372B4086AB1E6EE67AE1050AA6C50" type="personal">
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</mods:role>
<mods:namePart id="F650B8A9D50B79DC2E1241FD609E1C0D">Kuznetsova, Natalia</mods:namePart>
<mods:affiliation id="E7A7B1C3A982485010ACAF6FE53CDE0D">Moscow State Pedagogical University, Moscow 129278, Russia</mods:affiliation>
</mods:name>
<mods:name id="738E49A28C70811E55DB14F35BEF9E2D" type="personal">
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<mods:namePart id="0A2056350168D9561BD12A978C60D7A3">Nakamori, Taizo</mods:namePart>
<mods:affiliation id="432E806D14426F3384C68865AA27C696">Department of Environment and Natural Sciences, Graduate School of Environment and Information Sciences, Yokohama National University, Yokohama 240 - 8501, Japan</mods:affiliation>
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<mods:roleTerm id="D47D70BDEACE12402F2EB1BB66016FDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86729B61AF9D92FAF09B6613C29C14B7">Shveenkova, Yulia</mods:namePart>
<mods:affiliation id="8A66043F6AE4A775ECE30A5E2A1391AC">State Natural Reserve Privolzhskaya lesostep, Penza 440031, Russia</mods:affiliation>
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<mods:title id="E39EC1B42513283A5EC92BFF92859D1F">Zootaxa</mods:title>
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<mods:date id="8319AFF65DD2E770A2D9243996DD7F3B">2021</mods:date>
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<mods:number id="C42F7F350DAC02CFD7554647A5E6E60A">2021-03-02</mods:number>
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<mods:number id="AE87C43B3952E44DD1F85ED71AD3DCB4">4938</mods:number>
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<mods:classification id="2D41AAE4F2A7D01C947FF27D56BB6F4E">journal article</mods:classification>
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<mods:identifier id="0CACAD6ADB7F785C0C437C41B3243A5A" type="DOI">10.11646/zootaxa.4938.4.1</mods:identifier>
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<treatment id="BF05878D232CFFFCFF498D9EFB26C9D6" ID-DOI="http://doi.org/10.5281/zenodo.4574823" ID-GBIF-Taxon="180568725" ID-Zenodo-Dep="4574823" LSID="urn:lsid:plazi:treatment:BF05878D232CFFFCFF498D9EFB26C9D6" httpUri="http://treatment.plazi.org/id/BF05878D232CFFFCFF498D9EFB26C9D6" lastPageId="17" lastPageNumber="368" pageId="15" pageNumber="366">
<subSubSection id="7FB66510232CFFE2FF498D9EFCC9CAE6" box="[151,852,151,178]" pageId="15" pageNumber="366" type="nomenclature">
<paragraph id="3713369B232CFFE2FF498D9EFCC9CAE6" blockId="15.[151,852,151,214]" box="[151,852,151,178]" pageId="15" pageNumber="366">
<heading id="6C5B81F7232CFFE2FF498D9EFCC9CAE6" bold="true" box="[151,852,151,178]" fontSize="11" level="1" pageId="15" pageNumber="366" reason="1">
<emphasis id="05D8EA89232CFFE2FF498D9EFCC9CAE6" bold="true" box="[151,852,151,178]" pageId="15" pageNumber="366">
<taxonomicName id="F0AC4D18232CFFE2FF498D9EFD72CAE6" authority="Babenko &amp; Nakamori" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[151,751,151,178]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA89232CFFE2FF498D9EFE4ECAE5" bold="true" box="[151,467,151,178]" italics="true" pageId="15" pageNumber="366">Pseudachorutes morulifer</emphasis>
Babenko &amp; Nakamori
</taxonomicName>
<taxonomicNameLabel id="1EEB57F2232CFFE2FD288D9EFCC9CAE6" box="[758,852,151,177]" pageId="15" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232CFFE2FF498DB2FEE8CA82" box="[151,373,187,214]" pageId="15" pageNumber="366" type="description">
<paragraph id="3713369B232CFFE2FF498DB2FEE8CA82" blockId="15.[151,852,151,214]" box="[151,373,187,214]" pageId="15" pageNumber="366">
<figureCitation id="AF972A1E232CFFE2FF498DB2FE8ACA81" box="[151,279,187,214]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Figs 47–52</figureCitation>
,
<tableCitation id="7A2E0320232CFFE2FEFC8DB2FEE8CA82" box="[290,373,187,213]" captionStart="TABLE 1" captionStartId="34.[151,239,587,613]" captionText="TABLE 1. Main morphological characters of the East Asian species of the genus Pseudachorutes" pageId="15" pageNumber="366">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232CFFE2FF498C0AFB51CB86" pageId="15" pageNumber="366" type="materials_examined">
<paragraph id="3713369B232CFFE2FF498C0AFB51CB86" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
<emphasis id="05D8EA89232CFFE2FF498C0AFEDACB4A" bold="true" box="[151,327,259,285]" pageId="15" pageNumber="366">Type material.</emphasis>
<materialsCitation id="87C43CC6232CFFE2FE8E8C0DFD11CB16" ID-GBIF-Occurrence="3053021304" collectingDate="2016-08-25" country="Japan" latitude="34.686165" location="Park" longLatPrecision="11" longitude="135.857" municipality="Honshu Island" pageId="15" pageNumber="366" specimenCount="1" stateProvince="Nara" typeStatus="holotype">
<collectingCountry id="4FBB760B232CFFE2FE8E8C0DFE09CB49" box="[336,404,260,286]" name="Japan" pageId="15" pageNumber="366">Japan</collectingCountry>
:
<typeStatus id="E8178839232CFFE2FE7F8C0AFD9BCB4A" box="[417,518,259,285]" pageId="15" pageNumber="366" type="holotype">holotype</typeStatus>
, female,
<collectingMunicipality id="D777ACE1232CFFE2FDB18C0AFC8BCB4A" box="[623,790,259,285]" pageId="15" pageNumber="366">Honshu Island</collectingMunicipality>
, near
<collectingRegion id="F568F879232CFFE2FC838C0DFC0BCB49" box="[861,918,260,286]" country="Japan" name="Nara" pageId="15" pageNumber="366">Nara</collectingRegion>
,
<collectingRegion id="F568F879232CFFE2FC7D8C0DFC44CB49" box="[931,985,260,286]" country="Japan" name="Nara" pageId="15" pageNumber="366">Nara</collectingRegion>
<location id="32736040232CFFE2FC3C8C0AFB88CB4A" LSID="urn:lsid:plazi:treatment:BF05878D232CFFFCFF498D9EFB26C9D6:32736040232CFFE2FC3C8C0AFB88CB4A" box="[994,1045,259,285]" country="Japan" latitude="34.686165" longLatPrecision="11" longitude="135.857" municipality="Honshu Island" name="Park" pageId="15" pageNumber="366" stateProvince="Nara">Park</location>
(forest-park), debris on river bed,
<geoCoordinate id="5298505C232CFFE2FF498C2EFEBFCB15" box="[151,290,295,322]" degrees="34" direction="north" minutes="41.17" orientation="latitude" pageId="15" pageNumber="366" precision="9" value="34.686165">34°41.17’ N</geoCoordinate>
<geoCoordinate id="5298505C232CFFE2FEF28C2EFE59CB15" box="[300,452,295,322]" degrees="135" direction="east" minutes="51.42" orientation="longitude" pageId="15" pageNumber="366" precision="9" value="135.857">135°51.42’ E</geoCoordinate>
,
<date id="4312105B232CFFE2FE0D8C2EFD11CB16" box="[467,652,295,322]" pageId="15" pageNumber="366" value="2016-08-25">
<collectingDate id="5356E9B3232CFFE2FE0D8C2EFD11CB16" box="[467,652,295,322]" pageId="15" pageNumber="366" value="2016-08-25">25 August 2016</collectingDate>
</date>
</materialsCitation>
,
<collectorName id="9A59534D232CFFE2FD448C21FCDECB15" box="[666,835,296,322]" pageId="15" pageNumber="366">N. Kuznetsova</collectorName>
leg. [
<collectionCode id="51BDAE5E232CFFE2FC5B8C2EFC49CB16" box="[901,980,295,321]" pageId="15" pageNumber="366">MSPU</collectionCode>
collection].
<materialsCitation id="87C43CC6232CFFE2FBB98C21FB9CCBDE" ID-GBIF-Occurrence="3053021349" collectingDate="2016-08-05" country="Japan" county="Kyushu Island" elevation="1200" latitude="32.370834" location="Shiiba Village" longLatPrecision="12" longitude="131.0855" municipality="Ohkawauchi" pageId="15" pageNumber="366" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Miyazaki" typeStatus="paratype">
<typeStatus id="E8178839232CFFE2FBB98C21FB4ACB15" box="[1127,1239,296,322]" pageId="15" pageNumber="366" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="21AAFD12232CFFE2FB3A8C2EFACECB16" box="[1252,1363,295,321]" pageId="15" pageNumber="366" type="female">
<emphasis id="05D8EA89232CFFE2FB3A8C2EFB6FCB16" bold="true" box="[1252,1266,295,321]" pageId="15" pageNumber="366">2</emphasis>
females
</specimenCount>
and
<specimenCount id="21AAFD12232CFFE2FA508C2EFF4DCB32" pageId="15" pageNumber="366" type="male">
<emphasis id="05D8EA89232CFFE2FA508C2EFA01CB16" bold="true" box="[1422,1436,295,321]" pageId="15" pageNumber="366">1</emphasis>
male
</specimenCount>
,
<collectingCounty id="DE724E17232CFFE2FF3E8C42FE14CB32" box="[224,393,331,357]" pageId="15" pageNumber="366">Kyushu Island</collectingCounty>
,
<collectingRegion id="F568F879232CFFE2FE478C42FD19CB32" box="[409,644,331,357]" country="Japan" name="Miyazaki" pageId="15" pageNumber="366">Miyazaki Prefecture</collectingRegion>
,
<collectingMunicipality id="D777ACE1232CFFE2FD4D8C42FCBECB32" box="[659,803,331,357]" pageId="15" pageNumber="366">Ohkawauchi</collectingMunicipality>
,
<location id="32736040232CFFE2FCED8C42FC44CB32" LSID="urn:lsid:plazi:treatment:BF05878D232CFFFCFF498D9EFB26C9D6:32736040232CFFE2FCED8C42FC44CB32" box="[819,985,331,357]" country="Japan" county="Kyushu Island" latitude="32.370834" longLatPrecision="12" longitude="131.0855" municipality="Ohkawauchi" name="Shiiba Village" pageId="15" pageNumber="366" stateProvince="Miyazaki">Shiiba Village</location>
, «Mt. Tsunodake»,
<location id="32736040232CFFE2FB1E8C45FA94CB31" LSID="urn:lsid:plazi:treatment:BF05878D232CFFFCFF498D9EFB26C9D6:32736040232CFFE2FB1E8C45FA94CB31" box="[1216,1289,332,358]" country="Japan" county="Kyushu Island" latitude="32.370834" longLatPrecision="12" longitude="131.0855" municipality="Ohkawauchi" name="Yatate" pageId="15" pageNumber="366" stateProvince="Miyazaki">Yatate</location>
,
<geoCoordinate id="5298505C232CFFE2FAC68C42FA01CB31" box="[1304,1436,331,358]" degrees="32" direction="north" minutes="22.25" orientation="latitude" pageId="15" pageNumber="366" precision="9" value="32.370834">32°22.25’N</geoCoordinate>
<geoCoordinate id="5298505C232CFFE2FF498C66FE87CBDD" box="[151,282,367,394]" degrees="131" direction="east" minutes="5.13" orientation="longitude" pageId="15" pageNumber="366" precision="9" value="131.0855">131°5.13’E</geoCoordinate>
, ~
<quantity id="F0549B7E232CFFE2FEEA8C66FE17CBDD" box="[308,394,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="366" unit="m" value="1200.0">
<elevation id="BC81D1A8232CFFE2FEEA8C66FE17CBDD" box="[308,394,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="366" unit="m" value="1200.0">1200 m</elevation>
</quantity>
alt., shadow beech forest, rotten wood,
<date id="4312105B232CFFE2FC958C66FB9CCBDE" box="[843,1025,367,394]" pageId="15" pageNumber="366" value="2016-08-05">
<collectingDate id="5356E9B3232CFFE2FC958C66FB9CCBDE" box="[843,1025,367,394]" pageId="15" pageNumber="366" value="2016-08-05">05 August 2016</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="21AAFD12232CFFE2FBD28C66FBEDCBDE" box="[1036,1136,367,393]" pageId="15" pageNumber="366" type="female">
<emphasis id="05D8EA89232CFFE2FBD28C66FB87CBDE" bold="true" box="[1036,1050,367,393]" pageId="15" pageNumber="366">1</emphasis>
female
</specimenCount>
, same area, but
<taxonomicName id="F0AC4D18232CFFE2FAFA8C79FF25CBFA" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="05D8EA89232CFFE2FAFA8C79FF25CBFA" italics="true" pageId="15" pageNumber="366">Cryptomeria</emphasis>
</taxonomicName>
plantation;
<materialsCitation id="87C43CC6232CFFE2FEE08C9AFB51CB86" ID-GBIF-Occurrence="3053021309" collectingDate="2016-08-04" collectionCode="T, MSPU, TRPM" collectorName="All &amp; Nakamori, S &amp; Saitoh &amp; Potapov &amp; N. Kuznetsova" country="Japan" location="Maruju" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="21AAFD12232CFFE2FEE08C9AFE10CBFA" box="[318,397,403,429]" pageId="15" pageNumber="366" type="male">
<emphasis id="05D8EA89232CFFE2FEE08C9AFED1CBFA" bold="true" box="[318,332,403,429]" pageId="15" pageNumber="366">1</emphasis>
male
</specimenCount>
, same area, but «
<location id="32736040232CFFE2FD938C9AFD3BCBFA" LSID="urn:lsid:plazi:treatment:BF05878D232CFFFCFF498D9EFB26C9D6:32736040232CFFE2FD938C9AFD3BCBFA" box="[589,678,403,429]" country="Japan" name="Maruju" pageId="15" pageNumber="366">Maruju</location>
», mixed forest on slope, litter,
<date id="4312105B232CFFE2FBDF8C9AFB2ACBFA" box="[1025,1207,403,429]" pageId="15" pageNumber="366" value="2016-08-04">
<collectingDate id="5356E9B3232CFFE2FBDF8C9AFB2ACBFA" box="[1025,1207,403,429]" pageId="15" pageNumber="366" value="2016-08-04">04 August 2016</collectingDate>
</date>
.
<collectorName id="9A59534D232CFFE2FB618C9AFB7ECBFA" box="[1215,1251,403,429]" pageId="15" pageNumber="366">All</collectorName>
<collectionCode id="51BDAE5E232CFFE2FB348C9DFB61CBF9" box="[1258,1276,404,430]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="366">T</collectionCode>
.
<collectorName id="9A59534D232CFFE2FAD98C9AFA05CBFA" box="[1287,1432,403,429]" pageId="15" pageNumber="366">Nakamori, S</collectorName>
.
<collectorName id="9A59534D232CFFE2FF498CBEFF7CCB86" box="[151,225,439,465]" pageId="15" pageNumber="366">Saitoh</collectorName>
, M.
<collectorName id="9A59534D232CFFE2FECA8CB1FEEDCB85" box="[276,368,440,466]" pageId="15" pageNumber="366">Potapov</collectorName>
&amp;
<collectorName id="9A59534D232CFFE2FE4D8CB1FDA4CB85" box="[403,569,440,466]" pageId="15" pageNumber="366">N. Kuznetsova</collectorName>
leg. [3 ind.––
<collectionCode id="51BDAE5E232CFFE2FD0A8CBEFCBFCB86" box="[724,802,439,465]" pageId="15" pageNumber="366">MSPU</collectionCode>
collection, 2 ind.––
<collectionCode id="51BDAE5E232CFFE2FC218CBEFBD3CB86" box="[1023,1102,439,465]" country="Japan" lsid="urn:lsid:biocol.org:col:15811" name="Tottori Prefectural Museum" pageId="15" pageNumber="366" type="Herbarium">TRPM</collectionCode>
collection]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232CFFE2FF198CD2FC2EC8FE" pageId="15" pageNumber="366" type="diagnosis">
<paragraph id="3713369B232CFFE2FF198CD2FC2EC8FE" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
<emphasis id="05D8EA89232CFFE2FF198CD2FEDFCBA2" bold="true" box="[199,322,475,501]" pageId="15" pageNumber="366">Diagnosis.</emphasis>
Medium sized species. Ant. IV with clear «ventral file». PAO with complex lobes divided into several branches. Main part of labium with 3 proximal setae, seta L and two organites, both its basomedial and basolateral parts with 4 setae. Mandibles with four strong apical teeth. Dorsal chaetotaxy with short ordinary setae and usual number of long differentiated sensilla, head with both a0 and d0, Th. I with 4+4 setae, Th.II–III with 4 ordinary setae (a3, a4, m4 and p4) in dorso-external group additionally to S, Abd. I–III with 4+4 p-setae between sensilla. Mucro with basal «swelling». Each anal valve with 3 hr-setae.
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232CFFE2FF198FBAFCAACCB6" pageId="15" pageNumber="366" type="description">
<paragraph id="3713369B232CFFE2FF198FBAFE75C941" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
<emphasis id="05D8EA89232CFFE2FF198FBAFEC4C89A" bold="true" box="[199,345,691,717]" pageId="15" pageNumber="366">Description.</emphasis>
Habitus typical of genus, rather wide and somewhat flattened (
<figureCitation id="AF972A1E232CFFE2FBC78FBAFBF0C89A" box="[1049,1133,691,717]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 47</figureCitation>
). Length of adults (without antennae) from
<quantity id="F0549B7E232CFFE2FE968FDEFD9FC8A5" box="[328,514,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="1.115" metricValueMax="1.38" metricValueMin="0.85" pageId="15" pageNumber="366" unit="mm" value="1.115" valueMax="1.38" valueMin="0.85">0.85 to 1.38 mm</quantity>
. Colour unknown (only cleared specimens available). Tegument granulation rather coarse, especially on Abd. VI.
</paragraph>
<paragraph id="3713369B232CFFE2FF198E16FEB9C9BA" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">Antennae slightly shorter than head, Ant. III–IV fused dorsally, ventral separation well marked. Ant. IV with trilobed apical vesicle, external ms, subapical or and seta i present; sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV clearly differentiated, S1 and S2 somewhat slender, ventral side with about 30 truncate or slightly clavate sensilliform setae around longer one. Antennal organ of Ant. III typical, inner sensilla small, inclined to one side and located in cuticular fold, sgv only slightly longer than sgd, ventral ms present. Ant. I–II with 7 and 12–13 setae, respectively.</paragraph>
<paragraph id="3713369B232CFFE2FF198EFEFC17CE91" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
Head with 8+8 subequal ocelli. PAO elliptic, consisting of 10–14 compound vesicles, divided into 2–6 branches, ratio of its longer diameter to ocellus B as 1.4–1.6: 1 (
<figureCitation id="AF972A1E232CFFE2FCEF8912FC28CE61" box="[817,949,1051,1078]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Figs 51–52</figureCitation>
). Buccal cone elongated (
<figureCitation id="AF972A1E232CFFE2FB058912FAADCE62" box="[1243,1328,1051,1077]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 48</figureCitation>
). Maxilla styliform, with several tiny apical teeth (
<figureCitation id="AF972A1E232CFFE2FDBA8936FD25CE0D" box="[612,696,1087,1114]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 50</figureCitation>
), number of lamellae uncertain. Mandible triangular at apex with four strong teeth (
<figureCitation id="AF972A1E232CFFE2FEBC896AFE2ACE2A" box="[354,439,1123,1149]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 49</figureCitation>
). Distal edge of labrum truncate, number of labral setae as: 4/2334. Main part of labium with three proximal ordinary setae, seta L and two small labial organites (
<figureCitation id="AF972A1E232CFFE2FC01898EFBA8CEF6" box="[991,1077,1159,1185]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 48</figureCitation>
); submentum and mentum with usual set of four setae each, i.e. 4+4. Perilabial area with 5+5 setae.
</paragraph>
<paragraph id="3713369B232CFFE2FF1989C6FDF1CF96" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
Dorsal setae short and smooth, sensilla clearly longer, about as 2.5–3.5 times longer than ordinary setae, their number as usual: 22/1111. Chaetotaxy stable, asymmetrical abnormalities not frequent (
<figureCitation id="AF972A1E232CFFE2FBB189FAFB59CF5A" box="[1135,1220,1267,1293]" captionStart="FIGURES 47–52" captionStartId="16.[151,264,1959,1983]" captionTargetBox="[234,1358,192,1920]" captionTargetId="figure-29@16.[216,1372,181,1935]" captionTargetPageId="16" captionText="FIGURES 47–52. Pseudachorutes morulifer sp. nov.: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="15" pageNumber="366">Fig. 47</figureCitation>
). Main characteristics: head with both a0 and d0 present, Th. I with 4+4 setae, Th. II with a2-seta and lateral ms present, dorso-external group with three setae (a3–a4, m4) in front of p3–p4. Th. III identical but without a2 and ms. Abd. I–III with 1+1 extra setae in p-row between sensilla p4, i.e. 4+4 p-setae, totally, and 3 setae (a3, m3–m4) in front of p3–p3’–p
<quantity id="F0549B7E232CFFE2FAAF8856FA01CF2E" box="[1393,1436,1375,1401]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="366" unit="in" value="4.0">4 in</quantity>
dorso-external group. Abd. IV with more setae in a-row, some m-setae laterally, extra p-seta between sensilla present or absent. Abd. V without p2 as usual.
</paragraph>
<paragraph id="3713369B232CFFE2FF1988C2FBBACC22" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, Abd. II with 5(6)+5 sternal setae, Abd. III with 10–13 such setae on each side. Tenaculum with 3+3 teeth as usual. Furca complete, manubrium with 10+10 setae on main part, 4 setae on each basolateral lobe and 2 basal setae in line. Dorsal side of dens with six setae, hyaline field on its ventral side about as large as mucro length. Mucro curved, lateral lamella high but short forming basal «swelling». Each anal valve with three tiny hr-setae.</paragraph>
<paragraph id="3713369B232CFFE2FF198B89FCAACCB6" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">Legs I–III with typical number of setae: 1, 2, 2 setae on upper subcoxae, 0, 3, 2–3 setae on lower subcoxae, 3, 7–8, 7–8 setae on coxae, 6, 6, 6 on trochanters, 13, 12, 11 setae on femora and 19, 19, 18 setae on tibiotarsi. Unguis with strong basal tooth on inner edge, lateral teeth invisible.</paragraph>
</subSubSection>
<subSubSection id="7FB66510232CFFFCFF198BE2FBFFC90A" lastPageId="17" lastPageNumber="368" pageId="15" pageNumber="366" type="etymology">
<paragraph id="3713369B232CFFE2FF198BE2FF49CD7E" blockId="15.[151,1437,259,2014]" pageId="15" pageNumber="366">
<emphasis id="05D8EA89232CFFE2FF198BE2FED3CD52" bold="true" box="[199,334,1771,1797]" pageId="15" pageNumber="366">Etymology.</emphasis>
The name of the new species reflects its most characteristic feature—the morula-like shape of its PAO.
</paragraph>
<paragraph id="3713369B232CFFFCFF198A3AFCF5CB32" blockId="15.[151,1437,259,2014]" lastBlockId="17.[151,1437,151,897]" lastPageId="17" lastPageNumber="368" pageId="15" pageNumber="366">
<emphasis id="05D8EA89232CFFE2FF198A3AFEA0CD1A" bold="true" box="[199,317,1843,1869]" pageId="15" pageNumber="366">Affinities.</emphasis>
The new species possess two unique characters, which are very rare for the genus, namely, compound vesicles in the PAO and 4+4 setae of the p-row between sensilla on Abd. I–III. The only other known congener shared both these characters is
<taxonomicName id="F0AC4D18232CFFE2FDC08A75FC3ACDC1" authority="Weiner &amp; Najt, 1985" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[542,935,1915,1942]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA89232CFFE2FDC08A75FD28CDC2" box="[542,693,1915,1941]" italics="true" pageId="15" pageNumber="366">P. prosimplex</emphasis>
<bibRefCitation id="533D4B6A232CFFE2FD628A72FC3ACDC1" author="Weiner, W. M. &amp; Najt, J." box="[700,935,1915,1942]" pageId="15" pageNumber="366" pagination="245 - 276" refId="ref31847" refString="Weiner, W. M. &amp; Najt, J. (1985) Collemboles de Coree du Nord. IV. Brachystomellinae, Frieseinae et Pseudachorutinae ad partem. Acta Zoologica Cracoviensia, 28 (6), 245 - 276." type="journal article" year="1985">Weiner &amp; Najt, 1985</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="4FBB760B232CFFE2FB808A72FB72CDC1" box="[1118,1263,1915,1942]" name="North Korea" pageId="15" pageNumber="366">North Korea</collectingCountry>
. The difference of these two forms is limited by the shape of the mandibles:
<taxonomicName id="F0AC4D18232CFFE2FCE08AA9FC59CDEE" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[830,964,1951,1977]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA89232CFFE2FCE08AA9FC59CDEE" box="[830,964,1951,1977]" italics="true" pageId="15" pageNumber="366">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232CFFE2FC148A96FBB9CDEE" bold="true" box="[970,1060,1951,1977]" pageId="15" pageNumber="366">
<taxonomicNameLabel id="1EEB57F2232CFFE2FC148A96FBB9CDEE" box="[970,1060,1951,1977]" pageId="15" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has 4, but not 2 mandibular teeth as described for Korean type of
<taxonomicName id="F0AC4D18232CFFE2FE218ACDFD0BCD8A" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[511,662,1987,2013]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA89232CFFE2FE218ACDFD0BCD8A" box="[511,662,1987,2013]" italics="true" pageId="15" pageNumber="366">P. prosimplex</emphasis>
</taxonomicName>
. Initially we believed that our Japanese specimens are conspecific to the true
<taxonomicName id="F0AC4D182332FFFCFF2C8D9EFE14CAE6" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[242,393,151,177]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFF2C8D9EFE14CAE6" box="[242,393,151,177]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
and two additional teeth on mandibles have been missed during the original description. Fortunately, the
<typeStatus id="E81788392332FFFCFEF68DB2FE17CA82" box="[296,394,187,213]" pageId="17" pageNumber="368" type="holotype">holotype</typeStatus>
of
<taxonomicName id="F0AC4D182332FFFCFE6E8DB2FDDACA82" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[432,583,187,213]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFE6E8DB2FDDACA82" box="[432,583,187,213]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
is still kept in the ISEA collection (
<collectingCountry id="4FBB760B2332FFFCFC058DB2FBB1CA81" box="[987,1068,187,214]" name="Poland" pageId="17" pageNumber="368">Poland</collectingCountry>
) and was studied again by W.M. Weiner and G. Paśnik at our request. This study unambiguously confirmed the correctness of the initial description of the mandibular shape in
<taxonomicName id="F0AC4D182332FFFCFE128C0AFDFECB4A" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[460,611,259,285]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFE128C0AFDFECB4A" box="[460,611,259,285]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
(W.M. Weiner, person. comm.). Thus, despite the obvious close relationship between
<taxonomicName id="F0AC4D182332FFFCFEF18C2EFE59CB16" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[303,452,295,321]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFEF18C2EFE59CB16" box="[303,452,295,321]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
and
<taxonomicName id="F0AC4D182332FFFCFE278C2EFDE0CB16" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[505,637,295,321]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFE278C2EFDE0CB16" box="[505,637,295,321]" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFD5D8C2EFD46CB16" bold="true" box="[643,731,295,321]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFD5D8C2EFD46CB16" box="[643,731,295,321]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, they may well be considered independent species, at least until the variability of this usually very stable structure is established.
</paragraph>
<caption id="63D366132333FFFDFF498AAEFD59CDB5" ID-DOI="http://doi.org/10.5281/zenodo.4574833" ID-Zenodo-Dep="4574833" httpUri="https://zenodo.org/record/4574833/files/figure.png" pageId="16" pageNumber="367" startId="16.[151,264,1959,1983]" targetBox="[234,1358,192,1920]" targetPageId="16">
<paragraph id="3713369B2333FFFDFF498AAEFD59CDB5" blockId="16.[151,1436,1959,2019]" pageId="16" pageNumber="367">
<emphasis id="05D8EA892333FFFDFF498AAEFECBCDE8" bold="true" box="[151,342,1959,1983]" pageId="16" pageNumber="367">FIGURES 47–52.</emphasis>
<taxonomicName id="F0AC4D182333FFFDFE838AAEFDFBCDE8" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[349,614,1959,1983]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892333FFFDFE838AAEFDFBCDE8" box="[349,614,1959,1983]" italics="true" pageId="16" pageNumber="367">Pseudachorutes morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892333FFFDFDB08AAEFD5DCDE8" bold="true" box="[622,704,1959,1983]" pageId="16" pageNumber="367">
<taxonomicNameLabel id="1EEB57F22333FFFDFDB08AAEFD5DCDE8" box="[622,704,1959,1983]" pageId="16" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 47, dorsal chaetotaxy; 48, labium; 49, mandible; 50, maxillae; 51–52, PAO. Scales: Figs 47–48 = 0.1 mm, 49–52 = 0.01 mm.
</paragraph>
</caption>
<paragraph id="3713369B2332FFFCFF198C66FC1CCBA2" blockId="17.[151,1437,151,897]" pageId="17" pageNumber="368">
It should also be noted that this study revealed additional similarities between these two species. In particular, chaetotaxy of Abd. IV in
<taxonomicName id="F0AC4D182332FFFCFE648C9AFDCFCBFA" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[442,594,403,429]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFE648C9AFDCFCBFA" box="[442,594,403,429]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
appeared to be more usual and devoid of setae m1 (as in
<taxonomicName id="F0AC4D182332FFFCFB368C9AFAEECBFA" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[1256,1395,403,429]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFB368C9AFB64CBFB" box="[1256,1273,403,428]" italics="true" pageId="17" pageNumber="368">P</emphasis>
.
<emphasis id="05D8EA892332FFFCFAD78C9AFAEECBFA" box="[1289,1395,403,429]" italics="true" pageId="17" pageNumber="368">morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFAA58C9AFF55CB86" bold="true" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFAA58C9AFF55CB86" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and contrary to fig 15A in
<bibRefCitation id="533D4B6A2332FFFCFE288CBEFD46CB86" author="Weiner, W. M. &amp; Najt, J." box="[502,731,438,465]" pageId="17" pageNumber="368" pagination="245 - 276" refId="ref31847" refString="Weiner, W. M. &amp; Najt, J. (1985) Collemboles de Coree du Nord. IV. Brachystomellinae, Frieseinae et Pseudachorutinae ad partem. Acta Zoologica Cracoviensia, 28 (6), 245 - 276." type="journal article" year="1985">Weiner &amp; Najt 1985</bibRefCitation>
), sensilla S1 and S2 on Ant. IV in both species are slender than S3 and S4 and both species have three hr-setae on each anal valve.
</paragraph>
<paragraph id="3713369B2332FFFCFF198CF6FBFFC90A" blockId="17.[151,1437,151,897]" pageId="17" pageNumber="368">
There is another species in the fauna of
<collectingCountry id="4FBB760B2332FFFCFD4F8CF6FD49C84E" box="[657,724,511,537]" name="Japan" pageId="17" pageNumber="368">Japan</collectingCountry>
, namely
<taxonomicName id="F0AC4D182332FFFCFCE58CF6FBDBC84E" authority="Yosii, 1965" authorityName="Yosii" authorityYear="1965" box="[827,1094,510,537]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="05D8EA892332FFFCFCE58CF6FC27C84E" box="[827,954,511,537]" italics="true" pageId="17" pageNumber="368">P. insularis</emphasis>
<bibRefCitation id="533D4B6A2332FFFCFC1C8CF6FBDBC84E" author="Yosii, R." box="[962,1094,510,537]" pageId="17" pageNumber="368" pagination="1 - 71" refId="ref31944" refString="Yosii, R. (1965) On some Collembola of Japan and adjacent countries. Contributions from the Biological Laboratory, Kyoto University, 19, 1 - 71." type="journal article" year="1965">Yosii, 1965</bibRefCitation>
</taxonomicName>
, whose PAO is comparable to those of
<taxonomicName id="F0AC4D182332FFFCFF278F2AFE0DC86A" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[249,400,547,573]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFF278F2AFE0DC86A" box="[249,400,547,573]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
and
<taxonomicName id="F0AC4D182332FFFCFE168F2AFDD2C86A" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[456,591,547,573]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFE168F2AFDD2C86A" box="[456,591,547,573]" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFD888F2AFD2DC86A" bold="true" box="[598,688,547,573]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFD888F2AFD2DC86A" box="[598,688,547,573]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It has been described as being
<emphasis id="05D8EA892332FFFCFBCC8F2AFEE9C836" italics="true" pageId="17" pageNumber="368">composed of ca. 16 peripheral and 20 central elements</emphasis>
, and, judging from the fig.5 G &amp; H in
<bibRefCitation id="533D4B6A2332FFFCFCF38F4EFC21C836" author="Yosii, R." box="[813,956,582,609]" pageId="17" pageNumber="368" pagination="1 - 71" refId="ref31944" refString="Yosii, R. (1965) On some Collembola of Japan and adjacent countries. Contributions from the Biological Laboratory, Kyoto University, 19, 1 - 71." type="journal article" year="1965">Yosii (1965)</bibRefCitation>
, is similar in general appearance to that of
<taxonomicName id="F0AC4D182332FFFCFF498F62FE80C8D2" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[151,285,619,645]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFF498F62FE80C8D2" box="[151,285,619,645]" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFEFA8F62FEE0C8D2" bold="true" box="[292,381,619,645]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFEFA8F62FEE0C8D2" box="[292,381,619,645]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
According to existed descriptions, there are many formal differences between
<taxonomicName id="F0AC4D182332FFFCFB318F62FAF0C8D2" authorityName="Yosii" authorityYear="1965" box="[1263,1389,619,645]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="05D8EA892332FFFCFB318F62FAF0C8D2" box="[1263,1389,619,645]" italics="true" pageId="17" pageNumber="368">P. insularis</emphasis>
</taxonomicName>
and
<taxonomicName id="F0AC4D182332FFFCFF498F86FE80C8FE" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[151,285,655,681]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFF498F86FE80C8FE" box="[151,285,655,681]" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFEFA8F86FEE0C8FE" bold="true" box="[292,381,655,681]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFEFA8F86FEE0C8FE" box="[292,381,655,681]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(and
<taxonomicName id="F0AC4D182332FFFCFE638F86FDC9C8FE" authorityName="Weiner &amp; Najt" authorityYear="1985" box="[445,596,655,681]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFE638F86FDC9C8FE" box="[445,596,655,681]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
</taxonomicName>
as well).
<taxonomicName id="F0AC4D182332FFFCFD1F8F86FC40C8FE" authorityName="Yosii" authorityYear="1965" box="[705,989,655,681]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="05D8EA892332FFFCFD1F8F86FC40C8FE" box="[705,989,655,681]" italics="true" pageId="17" pageNumber="368">Pseudachorutes insularis</emphasis>
</taxonomicName>
is smaller (
<quantity id="F0549B7E2332FFFCFB818F86FB2AC8FE" box="[1119,1207,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="368" unit="mm" value="0.6">0.6 mm</quantity>
vs
<quantity id="F0549B7E2332FFFCFB038F86FAF9C8FE" box="[1245,1380,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="17" pageNumber="368" unit="mm" value="1.15" valueMax="1.4" valueMin="0.9">0.9–1.4 mm</quantity>
in
<taxonomicName id="F0AC4D182332FFFCFA568F86FE9CC89A" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFA568F86FE9CC89A" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFED98FBAFEFDC89A" bold="true" box="[263,352,691,717]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFED98FBAFEFDC89A" box="[263,352,691,717]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), appears to have no ventral file on Ant. IV, its mandibles have only two teeth as in
<taxonomicName id="F0AC4D182332FFFCFB218FBAFF21C8A6" authority=", VT" authorityName="VT" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="prosimplex">
<emphasis id="05D8EA892332FFFCFB218FBAFA08C89A" box="[1279,1429,691,717]" italics="true" pageId="17" pageNumber="368">P. prosimplex</emphasis>
, VT
</taxonomicName>
with 2+2 setae, only 3+3 setae on Th. I present, some other peculiarities in chaetotaxy also exist. However, most of these differences can be related to the size/age of the
<typeStatus id="E81788392332FFFCFCDC8FF2FCA0C942" box="[770,829,763,789]" pageId="17" pageNumber="368">types</typeStatus>
or be a result of misinterpretation. In fact,
<taxonomicName id="F0AC4D182332FFFCFAC68FF2FA01C942" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[1304,1436,763,789]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="morulifer" status="sp. nov.">
<emphasis id="05D8EA892332FFFCFAC68FF2FA01C942" box="[1304,1436,763,789]" italics="true" pageId="17" pageNumber="368">P. morulifer</emphasis>
</taxonomicName>
<emphasis id="05D8EA892332FFFCFF498E16FF6EC96E" bold="true" box="[151,243,799,825]" pageId="17" pageNumber="368">
<taxonomicNameLabel id="1EEB57F22332FFFCFF498E16FF6EC96E" box="[151,243,799,825]" pageId="17" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
may probably be a junior synonym for
<taxonomicName id="F0AC4D182332FFFCFD618E16FCA2C96E" authorityName="Yosii" authorityYear="1965" box="[703,831,799,825]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="05D8EA892332FFFCFD618E16FCA2C96E" box="[703,831,799,825]" italics="true" pageId="17" pageNumber="368">P. insularis</emphasis>
</taxonomicName>
, but this assumption is difficult to prove or disprove without studying the
<typeStatus id="E81788392332FFFCFE5A8E4AFE22C90A" box="[388,447,835,861]" pageId="17" pageNumber="368">types</typeStatus>
of the latter species. Unfortunately, we could not find them.
</paragraph>
</subSubSection>
<subSubSection id="7FB665102332FFFCFF198E6FFB26C9D6" box="[199,1211,870,897]" pageId="17" pageNumber="368" type="distribution">
<paragraph id="3713369B2332FFFCFF198E6FFB26C9D6" blockId="17.[151,1437,151,897]" box="[199,1211,870,897]" pageId="17" pageNumber="368">
<emphasis id="05D8EA892332FFFCFF198E6FFEFCC9D7" bold="true" box="[199,353,870,896]" pageId="17" pageNumber="368">Distribution.</emphasis>
The species is known only from two Japanese Islands: Honshu and Kyushu.
</paragraph>
</subSubSection>
</treatment>
</document>
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