187 lines
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187 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.732.21677" ID-GBIF-Dataset="05db50e8-b4e5-422a-b78f-9676d74ac319" ID-PMC="PMC5799741" ID-Pensoft-Pub="1313-2970-732-1" ID-PubMed="29416404" ID-ZBK="9995702B61464BA1BB5323DC9BA9650F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-732-1" ModsDocOrigin="ZooKeys 732" ModsDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" checkinTime="1516831538731" checkinUser="pensoft" docAuthor="Mattia, Willy De, Neiber, Marco T. & Groh, Klaus" docDate="2018" docId="BC08DBEE57C1779909BABD1034841CDE" docLanguage="en" docName="ZooKeys 732: 1-125" docOrigin="ZooKeys 732" docSource="http://dx.doi.org/10.3897/zookeys.732.21677" docTitle="Wollastonia ripkeni De Mattia & Groh, sp. n." docType="treatment" docUuid="A2E91669-0659-447D-9EB3-2F02174D4E87" docUuidSource="ZooBank" docVersion="6" lastPageNumber="37" masterDocId="FFA3FF812B32FFA8FFBAFFCDFF9AFF92" masterDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" masterLastPageNumber="125" masterPageNumber="1" pageNumber="33" updateTime="1668165370779" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mattia, Willy De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neiber, Marco T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Groh, Klaus</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>732</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>125</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.732.21677</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.732.21677</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-732-1</mods:identifier>
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<mods:identifier type="ZBK">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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<mods:identifier type="ZooBank">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140417766" LSID="urn:lsid:zoobank.org:act:A2E91669-0659-447D-9EB3-2F02174D4E87" httpUri="http://treatment.plazi.org/id/BC08DBEE57C1779909BABD1034841CDE" lastPageId="36" lastPageNumber="37" pageId="32" pageNumber="33">
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<subSubSection pageId="32" pageNumber="33" type="nomenclature">
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<paragraph pageId="32" pageNumber="33">
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†
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<taxonomicName LSID="http://zoobank.org/A2E91669-0659-447D-9EB3-2F02174D4E87" authority="De Mattia & Groh" class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia ripkeni" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="ripkeni">Wollastonia ripkeni De Mattia & Groh</taxonomicName>
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<taxonomicNameLabel pageId="32" pageNumber="33">sp. n.</taxonomicNameLabel>
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Figs 122-124, 125
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</paragraph>
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</subSubSection>
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<subSubSection pageId="33" pageNumber="34" type="type material">
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<paragraph pageId="33" pageNumber="34">
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<pageBreakToken pageId="33" pageNumber="34" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="33" pageNumber="34">SMF 348929, holotype, from loc. typ., leg. W. De Mattia & J. Macor, May 2015, NMWC Z.2016.013.00008, 2 PT (paratypes), NHMW 112144/1 PT, CKG, 2 PT, CWDM, 11 PT, from loc. typ., leg. W. De Mattia & J. Macor, May 2015.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="locus typicus">
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<paragraph pageId="34" pageNumber="35">
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<pageBreakToken pageId="34" pageNumber="35" start="start">Locus</pageBreakToken>
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typicus.
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</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Porto Santo, excavated mud walls behind the cart speedway E of the new harbour of Porto Santo,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.06333">33°03'48"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.30611">16°18'22"W</geoCoordinate>
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, 30 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="diagnosis">
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<paragraph pageId="34" pageNumber="35">Diagnosis.</paragraph>
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<paragraph pageId="34" pageNumber="35">Shell large for the genus, solid and conical. Whorls not rounded, not vertical but somewhat slanting. Surface of the shell covered by small, and relatively densely set tubercles. Body whorl bicarinated, with upper keel weaker than the lower one. Remaining teleoconch whorls only with the upper keel visible. Umbilicus narrow. Last whorl descending toward the aperture. Aperture oval with continuous peristome.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="description">
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<paragraph pageId="34" pageNumber="35">Description of the holotype.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Shell large for the genus, with 6⅔ regularly growing whorls, the protoconch with 1.85 whorls. The form of the shell is conical, the convex teleoconch whorls showing two rounded keels, the upper one less developed than the lower one. The last whorl measures 66%, the penultimate whorl 12% of the total shell height. The lower
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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of the body whorl are beneath the lower peripheral keel, which is set off by a constriction above and below the keel; outline of body whorl below the lower constriction regularly convex in frontal view. The two keels of the body whorl are placed in the upper 1∕11 and
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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of its total height. The suture between the whorls is simple, not sunken. The aperture, which is inclined to the vertical axis of the shell in an angle of 50° and is descending in the last 5% of the last whorl in an angle of 35° to the horizontal axis, has an elliptic form; its width is 46%, of the total shell width, its height 31% of the total shell height. It has a slightly reflected lip, which is completely detached from the body whorl. The eccentric umbilicus, which measures 10% of the total shell width, is in the last whorls circular and completely closed in the earlier whorls. The protoconch is smooth, the teleoconch shows a number of oblique radial ribs, 13 in the penultimate quadrant of the body-whorl and is additionally covered by numerous rounded tubercles. The number of tubercles in the standard-quadrate of the base is 89. There are no traces of colouration (Figs 122-124).
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</paragraph>
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<caption pageId="34" pageNumber="35">
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<paragraph pageId="34" pageNumber="35">
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Figures 122-124. Shells of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia ripkeni" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="ripkeni">Wollastonia ripkeni</taxonomicName>
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sp. n. 122 holotype, SMF 348929 123, 124 paratypes from the loc. typ. Scale bar 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="variation">
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<paragraph pageId="34" pageNumber="35">Variation of the paratypes.</paragraph>
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<paragraph pageId="34" pageNumber="35">Although the shell size is slightly variable, its shape is remarkably stable in all specimens.</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="description">
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<paragraph pageId="34" pageNumber="35">Measurements.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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D 8.0
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.4 mm (range 7.2-8.6 mm); H 6.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.4 mm (range 5.4-6.8 mm); FW 4.1
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2 mm; PA 42
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<normalizedToken originalValue="±">+/-</normalizedToken>
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4.1°; DU 0.9
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.08 mm; NT 89
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<normalizedToken originalValue="±">+/-</normalizedToken>
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23; NW 6.7
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2 (n = 16). Ratio D/H 1.3; ratio FW/H 0.7.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="distribution">
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<paragraph pageId="34" pageNumber="35">Distribution.</paragraph>
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<paragraph pageId="34" pageNumber="35">Known only from the locus typicus. The species was hitherto found only along a 50 m stretch of mud deposits close to the cart speedway. Additional deposits in the vicinity could not be checked for the presence of the species because of difficult accessibility of the steep slopes and associated risks of stone fall and landslides. The currently known distribution is shown in Fig. 125.</paragraph>
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<caption pageId="34" pageNumber="35">
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<paragraph pageId="34" pageNumber="35">
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Figure 125. Distribution of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia ripkeni" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="ripkeni">Wollastonia ripkeni</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="34" pageNumber="35" type="etymology">
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<paragraph pageId="34" pageNumber="35">Etymology.</paragraph>
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<paragraph pageId="34" pageNumber="35">
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Named for the Dutch malacologist Theodor (
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<normalizedToken originalValue="“Theo”">"Theo"</normalizedToken>
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) J. E. Ripken from Delft, The Netherlands, to honour his valuable contributions to the malacofauna of the Macaronesian Islands.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="comparisons">
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<paragraph pageId="34" pageNumber="35">Comparison and comments.</paragraph>
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<paragraph lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia ripkeni" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="ripkeni">Wollastonia ripkeni</taxonomicName>
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sp. n. can be confused on first glance with other large-sized species such as
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<taxonomicName lsidName="H. echinoderma" pageId="34" pageNumber="35" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
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,
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<taxonomicName lsidName="W. falknerorum" pageId="34" pageNumber="35" rank="species" species="falknerorum">W. falknerorum</taxonomicName>
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sp. n. or
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<taxonomicName lsidName="W. vermetiformis" pageId="34" pageNumber="35" rank="species" species="vermetiformis">W. vermetiformis</taxonomicName>
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. From the first it is separated by the development of two distinct keels, from the second by a much finer sculpture, a narrower umbilicus and narrower aperture. From the quite similar
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<taxonomicName lsidName="H. vermetiformis" pageId="34" pageNumber="35" rank="species" species="vermetiformis">H. vermetiformis</taxonomicName>
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it differs by a much more solid shell, a much higher last whorl, a larger aperture, finer granulation and coarser ribbing of the
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<pageBreakToken pageId="35" pageNumber="36" start="start">body</pageBreakToken>
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whorl. From similar sized
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<taxonomicName lsidName="W. subcarinulata" pageId="35" pageNumber="36" rank="species" species="subcarinulata">W. subcarinulata</taxonomicName>
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and
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<taxonomicName lsidName="W. inexpectata" pageId="35" pageNumber="36" rank="species" species="inexpectata">W. inexpectata</taxonomicName>
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sp. n. it is separated by a coarser granulation and the presence of two keels.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="taxonomic remarks">
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<paragraph pageId="35" pageNumber="36">Taxonomic remarks.</paragraph>
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<paragraph lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia ripkeni" order="Stylommatophora" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="ripkeni">Wollastonia ripkeni</taxonomicName>
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sp. n. is included in the genus
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
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because it is similar to
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<taxonomicName lsidName="W. oxytropis" pageId="35" pageNumber="36" rank="species" species="oxytropis">W. oxytropis</taxonomicName>
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in size and surface sculpture, considerably
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<pageBreakToken pageId="36" pageNumber="37" start="start">differing</pageBreakToken>
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from
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">Hystricella</taxonomicName>
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s. str. where the tubercles are bigger and less densely set. We are well aware that, due to the lack of anatomical and molecular data, the generic placement of this subfossil species is exclusively based upon shell features, with all the taxonomic limits that this might implicate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="36" pageNumber="37" type="status and conservation">
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<paragraph pageId="36" pageNumber="37">Status and conservation.</paragraph>
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<paragraph pageId="36" pageNumber="37">
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Extinct before the
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<normalizedToken originalValue="islands’">islands'</normalizedToken>
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scientific exploration in the 19th century, possibly already before human settlement.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |