192 lines
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192 lines
20 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.20.4344" ID-GBIF-Dataset="5aeaed8f-855a-47aa-a72b-588b29d98ef8" ID-PMC="PMC3652410" ID-Pensoft-Pub="1314-2003-20-17" ID-Pensoft-UUID="FFFDFF9BFFDFDE546166FFE4D472FFD1" ID-PubMed="23717192" ID-Zenodo-Dep="576158" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1314-2003-20-17" ModsDocOrigin="PhytoKeys 20" ModsDocTitle="A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments" checkinTime="1555332075685" checkinUser="pensoft" docAuthor="Penneys, Darin S. & Judd, Walter S." docDate="2013" docId="7182BC6414E389C16B79688A73C0C9C7" docLanguage="en" docName="PhytoKeys 20: 17-32" docOrigin="PhytoKeys 20" docSource="http://dx.doi.org/10.3897/phytokeys.20.4344" docTitle="Blakea P. Browne, Civ. Nat. Hist. Jamaica 323. tab. 35. 1756" docType="treatment" docVersion="6" id="FFFDFF9BFFDFDE546166FFE4D472FFD1" lastPageNumber="21" masterDocId="FFFDFF9BFFDFDE546166FFE4D472FFD1" masterDocTitle="A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments" masterLastPageNumber="32" masterPageNumber="17" pageNumber="20" updateTime="1668140735209" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Penneys, Darin S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Judd, Walter S.</mods:namePart>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>20</mods:number>
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<mods:start>17</mods:start>
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<mods:end>32</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.20.4344</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.20.4344</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-20-17</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFFDFF9BFFDFDE546166FFE4D472FFD1</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576158</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156201568" LSID="urn:lsid:plazi:treatment:7182BC6414E389C16B79688A73C0C9C7" httpUri="http://treatment.plazi.org/id/7182BC6414E389C16B79688A73C0C9C7" lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20">
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<subSubSection pageId="3" pageNumber="20" type="nomenclature">
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName LSID="BE5379C6-38A5-5A90-8437-C3E731E027D1" authority="P. Browne, Civ. Nat. Hist. Jamaica 323. tab. 35. 1756." class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Blakea" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">Blakea P.Browne, Civ. Nat. Hist. Jamaica 323. tab. 35. 1756.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="20" type="reference_group">
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName authorityName="Aublet" authorityYear="1775" class="Magnoliopsida" family="Melastomataceae" genus="Topobea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">Topobea</taxonomicName>
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Aubl., Hist. Pl. Guiane 1: 476. 1775. Type:
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<taxonomicName authorityName="Aubl" authorityYear="1775" class="Magnoliopsida" family="Melastomataceae" genus="Topobea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="parasitica">
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<emphasis italics="true" pageId="3" pageNumber="20">Topobea parasitica</emphasis>
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</taxonomicName>
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Aubl., Hist. Pl. Guiane 1: 476. 1775. TYPE. FRENCH GUIANA. Aublet s.n. (holotype: BM!).
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</paragraph>
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName class="Arachnida" family="Daesiidae" genus="Valdesia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Solifugae" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">Valdesia</taxonomicName>
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Ruiz & Pav., Fl. Peruv. Prodr. 67. 1794. Type:
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<taxonomicName authorityName="Ruiz & Pav" authorityYear="1798" class="Arachnida" family="Daesiidae" genus="Valdesia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Solifugae" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="repens">
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<emphasis italics="true" pageId="3" pageNumber="20">Valdesia repens</emphasis>
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</taxonomicName>
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Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 121. 1798. TYPE. PERU. Ruiz & Pav. s.n. (holotype: B (destroyed), photos at MO!, US!).
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</paragraph>
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName authorityName="Naudin" authorityYear="1852" class="Magnoliopsida" family="Melastomataceae" genus="Pyxidanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">Pyxidanthus</taxonomicName>
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Naudin, Ann. Sci. Nat. Bot. ser 3 18: 150. 1852. Type:
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<taxonomicName authorityName="Naud" authorityYear="1852" class="Magnoliopsida" family="Melastomataceae" genus="Pyxidanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="schlimii">
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<emphasis italics="true" pageId="3" pageNumber="20">Pyxidanthus schlimii</emphasis>
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</taxonomicName>
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Naudin, Ann. Sci. Nat., Bot.
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<normalizedToken originalValue="sér">ser</normalizedToken>
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. 3 18: 151. 1852. TYPE. VENEZUELA. Trujillo, Funck & Schlim 738 (holotype: G; (fragment), photos at MO!, US!; isotypes: BM!, BR!, MPU!).
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</paragraph>
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName authorityName="Linden ex Masters" authorityYear="1871" class="Magnoliopsida" family="Melastomataceae" genus="Amaraboya" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">Amaraboya</taxonomicName>
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Linden, Illustr. Hort. 34: 15. 1887. Type:
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Amaraboya" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="princeps">
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<emphasis italics="true" pageId="3" pageNumber="20">Amaraboya princeps</emphasis>
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</taxonomicName>
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Linden, Illustr. Hort. 34: 15. 1887. (lectotype, designated here:
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<emphasis italics="true" pageId="3" pageNumber="20">pl</emphasis>
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. IV of the protologue).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="20" type="type">
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<paragraph pageId="3" pageNumber="20">Type.</paragraph>
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<paragraph pageId="3" pageNumber="20">
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="trinervia">
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<emphasis italics="true" pageId="3" pageNumber="20">Blakea trinervia</emphasis>
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</taxonomicName>
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L., Syst. Nat. ed. 10: 1044. 1759. TYPE. JAMAICA. P.Browne s.n. (holotype: LINN-612.1!).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20" type="remarks">
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<paragraph pageId="3" pageNumber="20">Remarks.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20">
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="20">Blakea</emphasis>
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</taxonomicName>
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is characterized by solitary or fasciculate axillary inflorescences, and flowers subtended by two (rarely three) pairs of decussate, (usually) expanded, subtending bracts. Additionally,
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="20">Blakea</emphasis>
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</taxonomicName>
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may usually be distinguished from
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
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</taxonomicName>
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by the species often being hemiepiphytic or epiphytic, mostly with rotate corollas, anthers that are usually laterally flattened and often connate and/or deflexed, with the connective appendages generally better developed and of diverse morphology, and many
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="20">Blakea</emphasis>
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</taxonomicName>
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species have ovaries that are not entirely inferior, often possessing stylar collars. Cladistic analyses (Penneys et al. 2004,
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<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" refId="B18" refString="Penneys, DS, 2007. Phylogeny and character evolution in the Blakeeae (Melastomataceae). PhD Thesis. University of Florida, USA." title="Phylogeny and character evolution in the Blakeeae (Melastomataceae). PhD Thesis. University of Florida, USA." year="2007">Penneys 2007</bibRefCitation>
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,
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<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" publicationUrl="10.1086/657284" refId="B20" refString="Penneys, DS, Judd, WS, 2011. Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78-106. http://www.jstor.org/stable/10.1086/657284 ,., 10.1086/657284" title="Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78 - 106. http: // www. jstor. org / stable / 10.1086 / 657284," url="10.1086/657284" year="2011">Penneys and Judd 2011</bibRefCitation>
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, in press) have
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<pageBreakToken pageId="4" pageNumber="21" start="start">provided</pageBreakToken>
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conclusive evidence that the recognition
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<taxonomicName authorityName="Aublet" authorityYear="1775" class="Magnoliopsida" family="Melastomataceae" genus="Topobea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Topobea</emphasis>
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</taxonomicName>
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renders both that genus and
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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polyphyletic, thus
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<taxonomicName authorityName="Aublet" authorityYear="1775" class="Magnoliopsida" family="Melastomataceae" genus="Topobea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Topobea</emphasis>
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</taxonomicName>
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is here relegated to synonymy under
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="4" pageNumber="21">
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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comprises approximately 180 species distributed from Mexico (Chiapas) to Bolivia and Brazil, with two species in Jamaica and one in the Lesser Antilles.
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</paragraph>
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<paragraph pageId="4" pageNumber="21">
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Formal nomenclatural transfers to
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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are made below for 62 species of
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<taxonomicName authorityName="Aublet" authorityYear="1775" class="Magnoliopsida" family="Melastomataceae" genus="Topobea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Topobea</emphasis>
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</taxonomicName>
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that were not treated by earlier workers who made similar new combinations (e.g., Don 1823, Macbride 1941). Three of these include new specific epithets for names already occupied in
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="4" pageNumber="21">
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Important references concerning the systematics of
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Blakea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="4" pageNumber="21" phylum="Magnoliophyta" rank="genus">
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<emphasis italics="true" pageId="4" pageNumber="21">Blakea</emphasis>
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</taxonomicName>
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include (
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<bibRefCitation author="Almeda, F" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="12" pageNumber="29" pagination="299 - 326" refId="B1" refString="Almeda, F, 1990. New species and new combinations in Blakea and Topobea (Melastomataceae), with an historical perspective on generic limits in the tribe Blakeeae. Proceedings of the California Academy of Sciences 46: 299 - 326" title="New species and new combinations in Blakea and Topobea (Melastomataceae), with an historical perspective on generic limits in the tribe Blakeeae." volume="46" year="1990">Almeda 1990</bibRefCitation>
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,
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<bibRefCitation author="Almeda, F" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="12" pageNumber="29" publicationUrl="10.2307/3392974" refId="B2" refString="Almeda, F, 2000a. A synopsis of the genus Blakea (Melastomataceae) in Mexico and Central America. Novon10: 299-319. http://www.jstor.org/stable/10.2307/3392974 ,., 10.2307/3392974" title="A synopsis of the genus Blakea (Melastomataceae) in Mexico and Central America. Novon 10: 299 - 319. http: // www. jstor. org / stable / 10.2307 / 3392974," url="10.2307/3392974" year="2000 a">2000a</bibRefCitation>
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,
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<bibRefCitation author="Almeda, F" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="12" pageNumber="29" pagination="97 - 109" refId="B3" refString="Almeda, F, 2000b. The hexandrous species of Topobea (Melastomataceae). Proceedings of the California Academy of Sciences 52: 97 - 109" title="The hexandrous species of Topobea (Melastomataceae)." volume="52" year="2000 b">2000b</bibRefCitation>
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,
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<bibRefCitation author="Almeda, F" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="12" pageNumber="29" refId="B4" refString="Almeda, F, 2001a. The octandrous and dodecandrous species of Topobea (Melastomataceae) in Mexico and Central America. Proceedings of the California Academy of Sciences52: 511-548." title="The octandrous and dodecandrous species of Topobea (Melastomataceae) in Mexico and Central America. Proceedings of the California Academy of Sciences 52: 511 - 548." year="2001 a">2001a</bibRefCitation>
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,
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<bibRefCitation author="Almeda, F" editor="Stevens, WD" journalOrPublisher="Monographs in Systematic Botany, Missouri Botanical Garden" pageId="12" pageNumber="29" pagination="1339 - 1419" refId="B5" refString="Almeda, F, 2001b. Melastomataceae. In: Stevens, WD, Ulloa, Ulloa C, Pool, A, Montiel, OM, Eds., Flora de Nicaragua. Monographs in Systematic Botany, Missouri Botanical Garden 85: 1339 - 1419" title="Melastomataceae." volume="85" volumeTitle="Flora de Nicaragua." year="2001 b">2001b</bibRefCitation>
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,
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<bibRefCitation author="Almeda, F" editor="Davidse, G" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico City, Mexico" pageId="12" pageNumber="29" pagination="164 - 338" refId="B6" refString="Almeda, F, 2009. Melastomataceae. In: Davidse, G, Sousa-Sanchez, M, Knapp, S, Chiang, F, Eds., Flora Mesoamericana 4. Universidad Nacional Autonoma de Mexico, Mexico City, Mexico: 164 - 338" title="Melastomataceae." volumeTitle="Flora Mesoamericana 4." year="2009">2009</bibRefCitation>
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,
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<bibRefCitation author="Gleason, HA" journalOrPublisher="Universidad Nacional Autonoma de Mexico, Mexico City, Mexico" pageId="12" pageNumber="29" publicationUrl="10.2307/2481538" refId="B8" refString="Gleason, HA, 1945. On Blakea and Topobea. Bulletin of the Torrey Botanical Club 72: 385-393. http://www.jstor.org/stable/10.2307/2481538 ,., 10.2307/2481538" title="On Blakea and Topobea. Bulletin of the Torrey Botanical Club 72: 385 - 393. http: // www. jstor. org / stable / 10.2307 / 2481538," url="10.2307/2481538" year="1945">Gleason 1945</bibRefCitation>
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,
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<bibRefCitation author="Wurdack, JJ" editor="Lasser, T" journalOrPublisher="Instituto Botanico, Ministerio de Agricultura y Cria, Venezuela" pageId="13" pageNumber="30" pagination="1 - 819" refId="B26" refString="Wurdack, JJ, 1973. Melastomataceae. In: Lasser, T, Ed., Flora de Venezuela 8. Instituto Botanico, Ministerio de Agricultura y Cria, Venezuela: 1 - 819" title="Melastomataceae." volumeTitle="Flora de Venezuela 8." year="1973">Wurdack 1973</bibRefCitation>
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,
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<bibRefCitation author="Wurdack, JJ" editor="Harling, G" journalOrPublisher="University of Goeteborg and Riksmuseum, Sweden" pageId="13" pageNumber="30" pagination="1 - 406" refId="B27" refString="Wurdack, JJ, 1980. Melastomataceae. In: Harling, G, Sparre, B, Eds., Flora of Ecuador 13. University of Goeteborg and Riksmuseum, Sweden: 1 - 406" title="Melastomataceae." volumeTitle="Flora of Ecuador 13." year="1980">1980</bibRefCitation>
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,
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<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" refId="B18" refString="Penneys, DS, 2007. Phylogeny and character evolution in the Blakeeae (Melastomataceae). PhD Thesis. University of Florida, USA." title="Phylogeny and character evolution in the Blakeeae (Melastomataceae). PhD Thesis. University of Florida, USA." year="2007">Penneys 2007</bibRefCitation>
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,
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<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" publicationUrl="10.1086/657284" refId="B20" refString="Penneys, DS, Judd, WS, 2011. Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78-106. http://www.jstor.org/stable/10.1086/657284 ,., 10.1086/657284" title="Phylogenetics and morphology in the Blakeeae (Melastomataceae). International Journal of Plant Sciences 172: 78 - 106. http: // www. jstor. org / stable / 10.1086 / 657284," url="10.1086/657284" year="2011">Penneys and Judd 2011</bibRefCitation>
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,
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<bibRefCitation author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" refId="B21" refString="Penneys, DS, Judd, WS, in press. Combined molecular and morphological phylogenetic analyses of the Blakeeae (Melastomataceae). International Journal of Plant Sciences." title="Combined molecular and morphological phylogenetic analyses of the Blakeeae (Melastomataceae). International Journal of Plant Sciences." year="in press">in press</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
|
|
</treatment>
|
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</document> |