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<document id="94A8DA399A7797B5F6AE3F127D97663D" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFFD6147FF0263E4FE12FC3D" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madecorphnus falciger Lansberge 1886" docType="treatment" docVersion="16" lastPageNumber="46" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="45" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="AC697BCBBADA211D7DCA37D28DF51965">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<treatment id="5B4FBD00FFFD6147FF0263E4FE12FC3D" ID-DOI="http://doi.org/10.5281/zenodo.6057664" ID-GBIF-Taxon="125107276" ID-Zenodo-Dep="6057664" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFD6147FF0263E4FE12FC3D" httpUri="http://treatment.plazi.org/id/5B4FBD00FFFD6147FF0263E4FE12FC3D" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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<paragraph id="D3590C16FFFD6146FF0263E4FD13FE27" blockId="44.[151,665,440,499]" box="[151,665,440,466]" pageId="44" pageNumber="45">
<heading id="8811BB7AFFFD6146FF0263E4FD13FE27" bold="true" box="[151,665,440,466]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<taxonomicName id="14E67795FFFD6146FF0263E4FD13FE27" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[151,665,440,466]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFD6146FF0263E4FD13FE27" bold="true" box="[151,665,440,466]" pageId="44" pageNumber="45">
<emphasis id="E192D004FFFD6146FF0263E4FE3BFE27" bold="true" box="[151,433,440,466]" italics="true" pageId="44" pageNumber="45">Madecorphnus falciger</emphasis>
(
<bibRefCitation id="B77771E7FFFD6146FE5463E4FD1BFE27" author="Lansberge" box="[449,657,440,466]" pageId="44" pageNumber="45" refString="Lansberge, J. W. van (1886) Scarabaeides, Bupretides et Cerambycides de l'Afrique occidentale envoyes au Musee de Leyden par M. M. Veth et van der Kellen. Notes from the Leyden Museum, 8, 69 - 120." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FFFD6146FF026386FEBDFE06" blockId="44.[151,665,440,499]" box="[151,311,474,499]" pageId="44" pageNumber="45">
(
<figureCitation id="4BDD1093FFFD6146FF0A6386FF70FE06" box="[159,250,474,499]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="44" pageNumber="45">Figs. 22</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFD6146FF02607FFC2EFD83" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="D3590C16FFFD6146FF02607FFDEEFDCD" blockId="44.[151,932,546,630]" box="[151,612,546,568]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="F3F62B38FFFD6146FF02607FFDEEFDCD" box="[151,612,546,568]" pageId="44" pageNumber="45">
<taxonomicName id="14E67795FFFD6146FF02607FFDEAFDCD" authority="Lansberge, 1886: 93" authorityName="Lansberge" authorityPageNumber="93" authorityYear="1886" box="[151,608,546,568]" class="Insecta" family="Scarabaeidae" genus="Drepanognatus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFD6146FF02607FFE03FDCD" box="[151,393,546,568]" italics="true" pageId="44" pageNumber="45">Drepanognatus falciger</emphasis>
<treatmentCitation id="52472A07FFFD6146FE1A607EFDEAFDCD" author="Lansberge" box="[399,608,546,568]" page="93" pageId="44" pageNumber="45" year="1886">
<bibRefCitation id="B77771E7FFFD6146FE1A607EFDB0FDCD" author="Lansberge" box="[399,570,546,568]" pageId="44" pageNumber="45" refString="Lansberge, J. W. van (1886) Scarabaeides, Bupretides et Cerambycides de l'Afrique occidentale envoyes au Musee de Leyden par M. M. Veth et van der Kellen. Notes from the Leyden Museum, 8, 69 - 120." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
: 93
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D3590C16FFFD6146FF02601DFC2EFD83" blockId="44.[151,932,546,630]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="F3F62B38FFFD6146FF02601DFC2EFD83" pageId="44" pageNumber="45">
<taxonomicName id="14E67795FFFD6146FF02601DFE64FDA2" authority="Lansberge" authorityName="Lansberge" box="[151,494,577,599]" class="Insecta" family="Scarabaeidae" genus="Sissantobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFD6146FF02601DFEE1FDA2" box="[151,363,577,599]" italics="true" pageId="44" pageNumber="45">Sissantobius falciger</emphasis>
(Lansberge)
</taxonomicName>
:
<treatmentCitation id="52472A07FFFD6146FE6D601DFD38FDA2" author="Paulian" box="[504,690,577,599]" page="140" pageId="44" pageNumber="45" year="1937">
<bibRefCitation id="B77771E7FFFD6146FE6D601DFD0AFDA2" author="Paulian" box="[504,640,577,599]" pageId="44" pageNumber="45" refString="Paulian, R. (1937) Faune entomologique de Madagascar. Coleoptera Lamellicornia, Scarabaeidae Acanthocerini, Trogini, Aulonocnemis, Hybosorini, Orphnini et Ochodaeini. Bulletin de l'Academie malgache (nouvelle serie), 19, 129 - 143." type="journal article" year="1937">Paulian 1937</bibRefCitation>
: 140
</treatmentCitation>
, 1977: 1203.
<taxonomicName id="14E67795FFFD6146FF02603CFD8DFD83" authority="Lansberge" authorityName="Lansberge" box="[151,519,608,630]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFD6146FF02603CFE0EFD83" box="[151,388,608,630]" italics="true" pageId="44" pageNumber="45">Madecorphnus falciger</emphasis>
(Lansberge)
</taxonomicName>
:
<treatmentCitation id="52472A07FFFD6146FD84603CFD51FD83" author="Frolov" box="[529,731,608,630]" page="1107" pageId="44" pageNumber="45" year="2010">
<bibRefCitation id="B77771E7FFFD6146FD84603CFD17FD83" author="Frolov" box="[529,669,608,630]" pageId="44" pageNumber="45" refString="Frolov, A. V. (2010 c) Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera, Scarabaeidae, Orphninae). Journal of Natural History, 44, 1095 - 1111." type="journal article" year="2010" yearSuffix="c">Frolov 2010c</bibRefCitation>
: 1107
</treatmentCitation>
;
<treatmentCitation id="52472A07FFFD6146FD70603CFC2AFD83" author="Paulian" box="[741,928,608,630]" page="171" pageId="44" pageNumber="45" year="1992">
<bibRefCitation id="B77771E7FFFD6146FD70603CFCE7FD83" author="Paulian" box="[741,877,608,630]" pageId="44" pageNumber="45" refString="Paulian, R. (1992) Un nouveau genre et une nouvelle espece d'Orphnidae de Madagascar (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (N. S.), 14, 169 - 171." type="journal article" year="1992">Paulian 1992</bibRefCitation>
: 171
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="87995C9EFFFD6146FF02653CFB42F861" httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="44" pageNumber="45" targetBox="[270,1362,698,1864]" targetPageId="44">
<paragraph id="D3590C16FFFD6146FF02653CFB42F861" blockId="44.[151,1436,1888,1940]" pageId="44" pageNumber="45">
<emphasis id="E192D004FFFD6146FF02653CFEABF883" bold="true" box="[151,289,1888,1910]" pageId="44" pageNumber="45">FIGURE 22.</emphasis>
<taxonomicName id="14E67795FFFD6146FEBD653CFD9FF883" box="[296,533,1888,1910]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFD6146FEBD653CFD9FF883" box="[296,533,1888,1910]" italics="true" pageId="44" pageNumber="45">Madecorphnus falciger</emphasis>
</taxonomicName>
, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature).
</paragraph>
</caption>
<subSubSection id="9BFC5F9DFFFC6147FF0262CBFD7BFF0D" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="D3590C16FFFC6147FF0262CBFD7BFF0D" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF0262CBFE42FF45" bold="true" box="[151,456,151,176]" pageId="45" pageNumber="46">Type material examined.</emphasis>
Holotype (
<figureCitation id="4BDD1093FFFC6147FDC562CBFD3BFF45" box="[592,689,151,176]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Figs. 22</figureCitation>
AE), male, “
<taxonomicName id="14E67795FFFC6147FCC062CBFBA8FF45" authority="Rits." authorityName="Rits." box="[853,1058,151,176]" class="Insecta" family="Scarabaeidae" genus="Sissantobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FFFC6147FCC062CBFC6BFF45" box="[853,993,151,176]" italics="true" pageId="45" pageNumber="46">Sissantobius</emphasis>
Rits.
</taxonomicName>
?
<taxonomicName id="14E67795FFFC6147FBD062CBFAD3FF45" authority="Lansb. Madag." authorityName="Lansb. Madag." box="[1093,1369,151,176]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFC6147FBD062CBFB16FF45" box="[1093,1180,151,176]" italics="true" pageId="45" pageNumber="46">falciger</emphasis>
Lansb. Madag.
</taxonomicName>
”, Ex- Musaeo VAN LANSBERGE / G.J.Arrow vidit 1908 / MUSEUM PARIS 1952 COLL. R. OBERTHÜR /
<taxonomicName id="14E67795FFFC6147FF026283FE5EFF0D" authority="Lansb." authorityName="Lansb." box="[151,468,223,248]" class="Insecta" family="Scarabaeidae" genus="Sissantobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFC6147FF026283FE0BFF0D" box="[151,385,223,248]" italics="true" pageId="45" pageNumber="46">Sissantobius falciger</emphasis>
Lansb.
</taxonomicName>
HOLOTYPE” (MNHN).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF526358FC10FE91" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="D3590C16FFFC6147FF526358FC10FE91" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF526358FECBFEE8" bold="true" box="[199,321,260,285]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
This species differs from other
<taxonomicName id="14E67795FFFC6147FD506358FCE7FEE8" box="[709,877,260,285]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FFFC6147FD506358FCE7FEE8" box="[709,877,260,285]" italics="true" pageId="45" pageNumber="46">Madecorphnus</emphasis>
</taxonomicName>
species in having a characteristic shape of the parameres (
<figureCitation id="4BDD1093FFFC6147FE8C637BFEE1FEB5" box="[281,363,295,320]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Fig. 22</figureCitation>
B) and armature of the internal sac of the aedeagus consisting of 2 large, semicircular sclerites; 1 short, spur-like sclerite; and numerous small spinules (
<figureCitation id="4BDD1093FFFC6147FD6F6310FCDCFE91" box="[762,854,332,357]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Figs. 22</figureCitation>
DE).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF526333FE28FC75" pageId="45" pageNumber="46" type="description">
<paragraph id="D3590C16FFFC6147FF526333FC3BFE7D" blockId="45.[151,1437,151,968]" box="[199,945,367,392]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF526333FE29FE7D" bold="true" box="[199,419,367,392]" pageId="45" pageNumber="46">Description. Male.</emphasis>
Body length 6.0 mm. Color brown (
<figureCitation id="4BDD1093FFFC6147FCA96333FC05FE7D" box="[828,911,367,392]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Fig. 22</figureCitation>
A).
</paragraph>
<paragraph id="D3590C16FFFC6147FF5263C8FE1CFE25" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">Right mandible 2.5 longer than left, without tooth behind apex. Labrum trapezoidal, length about 1/51/6 width (in dorsal view).</paragraph>
<paragraph id="D3590C16FFFC6147FF526380FAA4FDE2" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and a number of smaller setae. Canthus and frontal suture indistinct. Frontoclypeus slightly depressed in the middle anteriorly, punctate.</paragraph>
<paragraph id="D3590C16FFFC6147FF526079FC8FFDAA" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">Pronotum 2 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, sparsely punctate.</paragraph>
<paragraph id="D3590C16FFFC6147FF526030FAF5FD39" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral humps. Elytra sparsely punctate with relatively large punctures.</paragraph>
<paragraph id="D3590C16FFFC6147FF52608BFD42FCE0" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">All legs of the holotype are broken with only the right profemur, left mesofemur, right mesofemur and mesotibia, and half of right metafemur remaining.</paragraph>
<paragraph id="D3590C16FFFC6147FF52617CFB64FCA9" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">
Parameres angulate apically (in lateral view), with small but distinct teeth laterally (
<figureCitation id="4BDD1093FFFC6147FBFF6143FB34FCCD" box="[1130,1214,799,824]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Fig. 22</figureCitation>
B). Internal sac with 2 large semicircular sclerites, 1 short spur-like sclerite and numerous small spinules (
<figureCitation id="4BDD1093FFFC6147FBDA6118FB21FCA9" box="[1103,1195,836,861]" captionStart="FIGURE 22" captionStartId="44.[151,250,1888,1910]" captionTargetBox="[270,1362,698,1864]" captionTargetId="figure@44.[218,1369,679,1866]" captionTargetPageId="44" captionText="FIGURE 22. Madecorphnus falciger, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature)." httpUri="https://zenodo.org/record/272643/files/figure.png" pageId="45" pageNumber="46">Figs. 22</figureCitation>
DE).
</paragraph>
<paragraph id="D3590C16FFFC6147FF52613BFE28FC75" blockId="45.[151,1437,151,968]" box="[199,418,871,896]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF52613BFEAFFC75" bold="true" box="[199,293,871,896]" pageId="45" pageNumber="46">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF5261D0FE12FC3D" pageId="45" pageNumber="46" type="distribution">
<paragraph id="D3590C16FFFC6147FF5261D0FE12FC3D" blockId="45.[151,1437,151,968]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF5261D0FE7FFC50" bold="true" box="[199,501,908,933]" pageId="45" pageNumber="46">Distribution and habitat.</emphasis>
The only known specimen of this species lacks any precise record of the locality where it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>