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<document id="99FBF5E54C5AF76B3A80A8FAB0015DE5" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FF906129FF0260CCFE52FCCD" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus bicavatus Fairmaire 1905" docType="treatment" docVersion="16" lastPageNumber="68" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="66" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="B0200EEB66C748252A311989EF45F016">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:name id="A06F0F2B94DE4392F7DD7C57054C0B37" type="personal">
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<mods:namePart id="6F3FD9B1B68C3AC23866764C8AD4DFFF">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="E78681EF9E6DB9796712C5B7BF7D7448">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="3D67CCB35BC1662A0FC9065A8E55D129">Akhmetova, Lilia A.</mods:namePart>
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<mods:title id="D747D2B7394D6BC6E38184BA33683D4A">Zootaxa</mods:title>
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<mods:date id="F51F6CA2E6BEBD9F7306BABFF4BA6D6A">2016</mods:date>
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<mods:identifier id="DAEC1F75E5DF89721D9674E9BF32A639" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<treatment id="5B4FBD00FF906129FF0260CCFE52FCCD" ID-DOI="http://doi.org/10.5281/zenodo.6057686" ID-GBIF-Taxon="125107302" ID-Zenodo-Dep="6057686" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD" httpUri="http://treatment.plazi.org/id/5B4FBD00FF906129FF0260CCFE52FCCD" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
<subSubSection id="9BFC5F9DFF90612BFF0260CCFEB0FD3E" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="D3590C16FF90612BFF0260CCFDFDFD5F" blockId="65.[151,631,656,715]" box="[151,631,656,682]" pageId="65" pageNumber="66">
<heading id="8811BB7AFF90612BFF0260CCFDFDFD5F" bold="true" box="[151,631,656,682]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<taxonomicName id="14E67795FF90612BFF0260CCFDFDFD5F" authority="Fairmaire, 1905" authorityName="Fairmaire" authorityYear="1905" box="[151,631,656,682]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF90612BFF0260CCFDFDFD5F" bold="true" box="[151,631,656,682]" pageId="65" pageNumber="66">
<emphasis id="E192D004FF90612BFF0260CCFE1FFD5F" bold="true" box="[151,405,656,682]" italics="true" pageId="65" pageNumber="66">Triodontus bicavatus</emphasis>
(
<bibRefCitation id="B77771E7FF90612BFE3160CCFDE5FD5F" author="Fairmaire" box="[420,623,656,682]" pageId="65" pageNumber="66" refString="Fairmaire, L. (1905) Materiaux pour la faune coleopterique de la region Malgache (19 e note). Annales de la Societe Entomologique de Belgique, 49, 114 - 138." type="journal article" year="1905">Fairmaire, 1905</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FF90612BFF0260EEFEB0FD3E" blockId="65.[151,631,656,715]" box="[151,314,690,715]" pageId="65" pageNumber="66">
(
<figureCitation id="4BDD1093FF90612BFF0A60EEFF73FD3E" box="[159,249,690,715]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="65" pageNumber="66">Figs. 33</figureCitation>
AH)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF90612BFF0260A6FD74FCBB" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="D3590C16FF90612BFF0260A6FDB2FCE5" blockId="65.[151,766,762,846]" box="[151,568,762,784]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="F3F62B38FF90612BFF0260A6FDB2FCE5" box="[151,568,762,784]" pageId="65" pageNumber="66">
<taxonomicName id="14E67795FF90612BFF0260A6FDBEFCE5" authority="Fairmaire, 1905: 118" authorityName="Fairmaire" authorityPageNumber="118" authorityYear="1905" box="[151,564,762,784]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF90612BFF0260A6FEDDFCE5" box="[151,343,762,784]" italics="true" pageId="65" pageNumber="66">Orphnus bicavatus</emphasis>
<treatmentCitation id="52472A07FF90612BFEC860A6FDBEFCE5" author="Fairmaire" box="[349,564,762,784]" page="118" pageId="65" pageNumber="66" year="1905">
<bibRefCitation id="B77771E7FF90612BFEC860A6FD88FCE5" author="Fairmaire" box="[349,514,762,784]" pageId="65" pageNumber="66" refString="Fairmaire, L. (1905) Materiaux pour la faune coleopterique de la region Malgache (19 e note). Annales de la Societe Entomologique de Belgique, 49, 114 - 138." type="journal article" year="1905">Fairmaire, 1905</bibRefCitation>
: 118
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D3590C16FF90612BFF026145FD74FCBB" blockId="65.[151,766,762,846]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="F3F62B38FF90612BFF026145FD74FCBB" pageId="65" pageNumber="66">
<taxonomicName id="14E67795FF90612BFF026145FE63FCDA" authority="Fairmaire" authorityName="Fairmaire" box="[151,489,793,815]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF90612BFF026145FEE6FCDA" box="[151,364,793,815]" italics="true" pageId="65" pageNumber="66">Triodontus bicavatus</emphasis>
(Fairmaire)
</taxonomicName>
:
<treatmentCitation id="52472A07FF90612BFE666145FD31FCDA" author="Paulian" box="[499,699,793,815]" page="1221" pageId="65" pageNumber="66" year="1977">
<bibRefCitation id="B77771E7FF90612BFE666145FDF1FCDA" author="Paulian" box="[499,635,793,815]" pageId="65" pageNumber="66" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian 1977</bibRefCitation>
: 1221
</treatmentCitation>
.
<taxonomicName id="14E67795FF90612BFF026164FE50FCBB" authority="Brancsick" authorityName="Brancsick" box="[151,474,824,846]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="E192D004FF90612BFF026164FEE1FCBB" box="[151,363,824,846]" italics="true" pageId="65" pageNumber="66">Triodontus obsoletus</emphasis>
Brancsick
</taxonomicName>
: synonymy by
<bibRefCitation id="B77771E7FF90612BFDE66164FD70FCBB" author="Paulian" box="[627,762,824,846]" pageId="65" pageNumber="66" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian 1977</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF90612BFF026126FAA5FAB6" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="D3590C16FF90612BFF026126FC88FC42" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">
<emphasis id="E192D004FF90612BFF026126FE41FC66" bold="true" box="[151,459,890,915]" pageId="65" pageNumber="66">Type material examined.</emphasis>
Holotype (
<figureCitation id="4BDD1093FF90612BFDC36126FD32FC66" box="[598,696,890,915]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="65" pageNumber="66">Figs. 33</figureCitation>
A, D), male, “MUSÉUM PARIS 1906 Coll. Leon Fairmaire /
<taxonomicName id="14E67795FF90612BFF0261C2FEE6FC42" box="[151,364,926,951]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF90612BFF0261C2FEE6FC42" box="[151,364,926,951]" italics="true" pageId="65" pageNumber="66">Orphnus bicavatus</emphasis>
</taxonomicName>
opm n. sp. XXX / TYPE” (MNHN)
</paragraph>
<paragraph id="D3590C16FF90612BFF52619EFAA5FAB6" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">
<emphasis id="E192D004FF90612BFF52619EFC6DFC2E" bold="true" box="[199,999,962,987]" pageId="65" pageNumber="66">
Additional material examined. MADAGASCAR.
<materialsCitation id="638E064BFF90612BFCDC619EFC6DFC2E" ID-GBIF-Occurrence="1416610081" box="[841,999,962,987]" collectionCode="MNHN" collectorName="Museum Paris &amp; Brouhard" latitude="-12.530556" location="Mt. d'Ambre" longLatPrecision="21" longitude="49.173336" municipality="Amber Mountains" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Antsiranana">
<collectingRegion id="1122C2F4FF90612BFCDC619EFC69FC2E" box="[841,995,962,987]" country="Madagascar" name="Antsiranana" pageId="65" pageNumber="66">Antsiranana</collectingRegion>
:
</materialsCitation>
</emphasis>
one male, “
<location id="D6395ACDFF90612BFB06619FFABBFC2E" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFB06619FFABBFC2E" box="[1171,1329,962,987]" latitude="-12.530556" longLatPrecision="21" longitude="49.173336" municipality="Amber Mountains" name="Mt. d'Ambre" pageId="65" pageNumber="66" stateProvince="Antsiranana">Mt. d'Ambre</location>
[
<collectingMunicipality id="333D966CFF90612BFADE619EFE98FC0A" pageId="65" pageNumber="66">Amber Mountains</collectingMunicipality>
,
<geoCoordinate id="B6D26AD1FF90612BFEB661BAFE2CFC0A" box="[291,422,998,1023]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="15" value="-12.530556">12°3150″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF90612BFE2261BAFDB6FC0A" box="[439,572,998,1023]" direction="east" orientation="longitude" pageId="65" pageNumber="66" precision="15" value="49.173336">49°1024″E</geoCoordinate>
] /
<collectorName id="7E1369C0FF90612BFDF061BBFC86FC0A" box="[613,780,998,1023]" pageId="65" pageNumber="66">Museum Paris</collectorName>
1934
<collectorName id="7E1369C0FF90612BFCCB61BAFC40FC0A" box="[862,970,998,1023]" pageId="65" pageNumber="66">Brouhard</collectorName>
” (
<collectionCode id="B5F794D3FF90612BFC7C61BBFBCFFC0A" box="[1001,1093,999,1023]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
);
<materialsCitation id="638E064BFF90612BFBCB61BBFBF1FBD6" ID-GBIF-Occurrence="1416610088" collectingDate="1936-12" collectionCode="MNHN" county="Ampanefena" latitude="-13.866667" location="Ampanefena" longLatPrecision="21" longitude="49.966667" municipality="Vohemar" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Antsiranana">
one male, “Mad. Vohemar,
<location id="D6395ACDFF90612BFF026656FEACFBD6" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFF026656FEACFBD6" box="[151,294,1034,1059]" county="Ampanefena" latitude="-13.866667" longLatPrecision="21" longitude="49.966667" municipality="Vohemar" name="Ampanefena" pageId="65" pageNumber="66" stateProvince="Antsiranana">Ampanefena</location>
<collectingDate id="B71CD33EFF90612BFEB86656FEFDFBD6" box="[301,375,1034,1059]" pageId="65" pageNumber="66" value="1936-12">
<date id="A7582AD6FF90612BFEB86656FEFDFBD6" box="[301,375,1034,1059]" pageId="65" pageNumber="66" value="1936-12">XII.36</date>
</collectingDate>
<location id="D6395ACDFF90612BFEE86656FE4FFBD6" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFEE86656FE4FFBD6" box="[381,453,1034,1059]" county="Ampanefena" latitude="-13.866667" longLatPrecision="21" longitude="49.966667" municipality="Vohemar" name="Vadon" pageId="65" pageNumber="66" stateProvince="Antsiranana">Vadon</location>
[
<collectingMunicipality id="333D966CFF90612BFE446656FDB1FBD6" box="[465,571,1034,1059]" pageId="65" pageNumber="66">Vohemar</collectingMunicipality>
,
<collectingCounty id="3A38749AFF90612BFDD36656FD52FBD6" box="[582,728,1034,1059]" pageId="65" pageNumber="66">Ampanefena</collectingCounty>
,
<geoCoordinate id="B6D26AD1FF90612BFD776656FCECFBD6" box="[738,870,1034,1059]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="15" value="-13.866667">13°5200″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF90612BFCE56656FC7DFBD6" box="[880,1015,1034,1059]" direction="east" orientation="longitude" pageId="65" pageNumber="66" precision="15" value="49.966667">49°5800″E</geoCoordinate>
]” (
<collectionCode id="B5F794D3FF90612BFB826657FBF9FBD6" box="[1047,1139,1035,1059]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFB106657FB9EFBB2" ID-GBIF-Occurrence="1416610093" collectionCode="MNHN" collectorName="Museum Paris &amp; Coll. A." latitude="-12.266666" location="Diego Suarez" longLatPrecision="21" longitude="49.283054" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Antsiranana">
one male, “
<location id="D6395ACDFF90612BFA966656FA16FBD6" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFA966656FA16FBD6" box="[1283,1436,1034,1059]" latitude="-12.266666" longLatPrecision="21" longitude="49.283054" name="Diego Suarez" pageId="65" pageNumber="66" stateProvince="Antsiranana">Diego Suarez</location>
[
<geoCoordinate id="B6D26AD1FF90612BFF0B6672FEA9FBB2" box="[158,291,1070,1095]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="15" value="-12.266666">12°1600″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF90612BFEBB6672FE38FBB2" box="[302,434,1070,1095]" direction="east" orientation="longitude" pageId="65" pageNumber="66" precision="15" value="49.283054">49°1659″E</geoCoordinate>
] /
<collectorName id="7E1369C0FF90612BFE5B6673FDE4FBB2" box="[462,622,1070,1095]" pageId="65" pageNumber="66">Museum Paris</collectorName>
1936
<collectorName id="7E1369C0FF90612BFD266672FC86FBB2" box="[691,780,1070,1095]" pageId="65" pageNumber="66">Coll. A.</collectorName>
Boucomont” (
<collectionCode id="B5F794D3FF90612BFC256673FB86FBB2" box="[944,1036,1071,1095]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFB8A6672FAC6FB46" ID-GBIF-Occurrence="1416610095" collectingDate="1937-12" collectingDateMax="1960-11" collectingDateMin="1937-12" collectionCode="MNHN" collectorName="Madagascar-Est &amp; Sambava R. N. &amp; Ambatosoratra &amp; P. Soga &amp; Natural Reserve &amp; Marojejy" country="Madagascar" county="Andapa" elevation="1700" latitude="-14.433056" location="Andapa" longLatPrecision="21" longitude="49.733055" municipality="Antalaha District" pageId="65" pageNumber="66" specimenCount="1">
four males, five females, “
<location id="D6395ACDFF90612BFAD66672FA17FBB2" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFAD66672FA17FBB2" box="[1347,1437,1070,1095]" country="Madagascar" county="Andapa" latitude="-14.433056" longLatPrecision="21" longitude="49.733055" municipality="Antalaha District" name="Andapa" pageId="65" pageNumber="66">Andapa</location>
/
<collectingCountry id="ABF14C86FF90612BFF3D660EFEA5FB9E" box="[168,303,1106,1131]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
Ron Antalaha
<collectingDate id="B71CD33EFF90612BFE74660FFDA0FB9E" box="[481,554,1107,1131]" pageId="65" pageNumber="66" value="1937-12">
<date id="A7582AD6FF90612BFE74660FFDA0FB9E" box="[481,554,1107,1131]" pageId="65" pageNumber="66" value="1937-12">XII.37</date>
</collectingDate>
<location id="D6395ACDFF90612BFDA1660EFDF1FB9E" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFDA1660EFDF1FB9E" box="[564,635,1106,1131]" country="Madagascar" county="Andapa" latitude="-14.433056" longLatPrecision="21" longitude="49.733055" municipality="Antalaha District" name="Vadon" pageId="65" pageNumber="66">Vadon</location>
[
<collectingMunicipality id="333D966CFF90612BFD19660EFCDFFB9E" box="[652,853,1106,1131]" pageId="65" pageNumber="66">Antalaha District</collectingMunicipality>
,
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,
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,
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” (
<collectionCode id="B5F794D3FF90612BFB69660FFAD2FB9E" box="[1276,1368,1107,1131]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
); four males, nine females, “
<collectorName id="7E1369C0FF90612BFE36662AFDD0FB7A" box="[419,602,1142,1167]" pageId="65" pageNumber="66">Madagascar-Est</collectorName>
dct.
<collectorName id="7E1369C0FF90612BFD0A662AFCC8FB7A" box="[671,834,1142,1167]" pageId="65" pageNumber="66">Sambava R.N.</collectorName>
XII-Marojejy
<collectorName id="7E1369C0FF90612BFC7B662AFB1EFB7A" box="[1006,1172,1142,1167]" pageId="65" pageNumber="66">Ambatosoratra</collectorName>
<elevation id="58CBEB25FF90612BFB36662BFB7BFB7A" box="[1187,1265,1143,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="65" pageNumber="66" unit="m" value="1700.0">
<quantity id="141EA1F3FF90612BFB36662BFB7BFB7A" box="[1187,1265,1143,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="65" pageNumber="66" unit="m" value="1700.0">1700m</quantity>
</elevation>
<collectingDate id="B71CD33EFF90612BFB6A662AFAC8FB7A" box="[1279,1346,1142,1167]" pageId="65" pageNumber="66" value="1960-11">
<date id="A7582AD6FF90612BFB6A662AFAC8FB7A" box="[1279,1346,1142,1167]" pageId="65" pageNumber="66" value="1960-11">XI-60</date>
</collectingDate>
<collectorName id="7E1369C0FF90612BFAC4662BFA16FB7A" box="[1361,1436,1143,1167]" pageId="65" pageNumber="66">P.Soga</collectorName>
[
<collectorName id="7E1369C0FF90612BFF0866C6FEC3FB46" box="[157,329,1178,1203]" pageId="65" pageNumber="66">Ambatosoratra</collectorName>
,
<collectorName id="7E1369C0FF90612BFECC66C6FD99FB47" box="[345,531,1178,1203]" pageId="65" pageNumber="66">Natural Reserve</collectorName>
of
<collectorName id="7E1369C0FF90612BFDD766C6FD23FB46" box="[578,681,1178,1203]" pageId="65" pageNumber="66">Marojejy</collectorName>
(RNI XII),
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<quantity id="141EA1F3FF90612BFCA866C7FC13FB47" box="[829,921,1179,1203]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="65" pageNumber="66" unit="m" value="1700.0">1700 m</quantity>
</elevation>
,
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,
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]” (
<collectionCode id="B5F794D3FF90612BFB7D66C7FACEFB46" box="[1256,1348,1179,1203]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFAC966C7FE2CFB0E" ID-GBIF-Occurrence="1416610101" collectingDate="1935-12" collectionCode="MNHN" country="Madagascar" county="Andapa" latitude="-14.65" location="Vadon Andapa" longLatPrecision="21" longitude="49.65" municipality="Antalaha District" pageId="65" pageNumber="66" specimenCount="10" specimenCount-female="10">
seven males,
<specimenCount id="C5E0C79FFF90612BFF7F66E3FEE1FB22" box="[234,363,1214,1239]" pageId="65" pageNumber="66" type="female">10 females</specimenCount>
, “
<collectingCountry id="ABF14C86FF90612BFE1166E2FD84FB22" box="[388,526,1214,1239]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
Ron Antalaha
<collectingDate id="B71CD33EFF90612BFD5666E2FC87FB22" box="[707,781,1214,1239]" pageId="65" pageNumber="66" value="1935-12">
<date id="A7582AD6FF90612BFD5666E2FC87FB22" box="[707,781,1214,1239]" pageId="65" pageNumber="66" value="1935-12">XII.35</date>
</collectingDate>
<location id="D6395ACDFF90612BFC8D66E2FC4BFB22" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFC8D66E2FC4BFB22" box="[792,961,1214,1239]" country="Madagascar" county="Andapa" latitude="-14.65" longLatPrecision="21" longitude="49.65" municipality="Antalaha District" name="Vadon Andapa" pageId="65" pageNumber="66">Vadon Andapa</location>
[
<collectingMunicipality id="333D966CFF90612BFC4766E2FB17FB22" box="[978,1181,1214,1239]" pageId="65" pageNumber="66">Antalaha District</collectingMunicipality>
,
<collectingCounty id="3A38749AFF90612BFB3966E2FA8CFB22" box="[1196,1286,1214,1239]" pageId="65" pageNumber="66">Andapa</collectingCounty>
,
<geoCoordinate id="B6D26AD1FF90612BFA8066E2FA1DFB22" box="[1301,1431,1214,1239]" direction="south" orientation="latitude" pageId="65" pageNumber="66" precision="15" value="-14.65">14°3900″S</geoCoordinate>
,
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]” (
<collectionCode id="B5F794D3FF90612BFED766BFFE14FB0E" box="[322,414,1251,1275]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFE2066BFFAEEFB0E" ID-GBIF-Occurrence="1416610098" box="[437,1380,1250,1275]" collectionCode="MNHN" country="Madagascar" location="Mt. d'Ambre" municipality="Amber Mountains" pageId="65" pageNumber="66" specimenCount="42" specimenCount-female="32" specimenCount-male="10">
<specimenCount id="C5E0C79FFF90612BFE2066BFFDAAFB0E" box="[437,544,1250,1275]" pageId="65" pageNumber="66" type="male">10 males</specimenCount>
,
<specimenCount id="C5E0C79FFF90612BFDBA66BFFD3BFB0E" box="[559,689,1250,1275]" pageId="65" pageNumber="66" type="female">32 females</specimenCount>
, “
<location id="D6395ACDFF90612BFD5E66BFFCEAFB0E" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFD5E66BFFCEAFB0E" box="[715,864,1250,1275]" country="Madagascar" municipality="Amber Mountains" name="Mt. d'Ambre" pageId="65" pageNumber="66">Mt. d'Ambre</location>
[
<collectingMunicipality id="333D966CFF90612BFCE466BEFBC2FB0E" box="[881,1096,1250,1275]" pageId="65" pageNumber="66">Amber Mountains</collectingMunicipality>
]
<collectingCountry id="ABF14C86FF90612BFBCF66BEFB6EFB0E" box="[1114,1252,1250,1275]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
” (
<collectionCode id="B5F794D3FF90612BFA9566BFFAD6FB0E" box="[1280,1372,1251,1275]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFAE166BFFB77FAEA" ID-GBIF-Occurrence="1416610122" collectingDate="2005-12-28" collectionCode="ABCB" collectorName="A. Ballerio" country="Madagascar" location="Madagascar" pageId="65" pageNumber="66" specimenCount="1">
one male, “
<collectingCountry id="ABF14C86FF90612BFF73675AFEFAFAEA" box="[230,368,1286,1311]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
Nord Montagne D'Ambre Foret de Sakaramy
<collectingDate id="B71CD33EFF90612BFC15675AFB80FAEA" box="[896,1034,1286,1311]" pageId="65" pageNumber="66" value="2005-12-28">
<date id="A7582AD6FF90612BFC15675AFB80FAEA" box="[896,1034,1286,1311]" pageId="65" pageNumber="66" value="2005-12-28">28.XII.2005</date>
</collectingDate>
<collectorName id="7E1369C0FF90612BFB84675AFB03FAEA" box="[1041,1161,1286,1311]" pageId="65" pageNumber="66">A.Ballerio</collectorName>
” (
<collectionCode id="B5F794D3FF90612BFB36675BFB7FFAEA" box="[1187,1269,1287,1311]" pageId="65" pageNumber="66">ABCB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF90612BFA9F675AFAA1FAB6" ID-GBIF-Occurrence="1416610115" collectingDate="1937-12" collectionCode="MNHN" collectorName="Vadon" country="Madagascar" location="Anginjanbary" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Mahajanga">
<emphasis id="E192D004FF90612BFA9F675AFA16FAEA" bold="true" box="[1290,1436,1286,1311]" pageId="65" pageNumber="66">
<collectingRegion id="1122C2F4FF90612BFA9F675AFA1DFAEA" box="[1290,1431,1286,1311]" country="Madagascar" name="Mahajanga" pageId="65" pageNumber="66">Mahajanga</collectingRegion>
:
</emphasis>
three males, “
<location id="D6395ACDFF90612BFEBA6776FE46FAB6" LSID="urn:lsid:plazi:treatment:5B4FBD00FF906129FF0260CCFE52FCCD:D6395ACDFF90612BFEBA6776FE46FAB6" box="[303,460,1322,1347]" country="Madagascar" name="Anginjanbary" pageId="65" pageNumber="66" stateProvince="Mahajanga">Anginjanbary</location>
[dubious locality] /
<collectingCountry id="ABF14C86FF90612BFD3B6776FCBFFAB6" box="[686,821,1322,1347]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
Ron Mandritsara
<collectingDate id="B71CD33EFF90612BFB946777FBC7FAB6" box="[1025,1101,1323,1347]" pageId="65" pageNumber="66" value="1937-12">
<date id="A7582AD6FF90612BFB946777FBC7FAB6" box="[1025,1101,1323,1347]" pageId="65" pageNumber="66" value="1937-12">XII-37</date>
</collectingDate>
<collectorName id="7E1369C0FF90612BFBC16776FB16FAB6" box="[1108,1180,1322,1347]" pageId="65" pageNumber="66">Vadon</collectorName>
!” (
<collectionCode id="B5F794D3FF90612BFB526777FAA9FAB6" box="[1223,1315,1323,1347]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="65" pageNumber="66">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF90612BFF526712FE73FA27" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="D3590C16FF90612BFF526712FE73FA27" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">
<emphasis id="E192D004FF90612BFF526712FECBFA92" bold="true" box="[199,321,1358,1383]" pageId="65" pageNumber="66">Diagnosis.</emphasis>
<taxonomicName id="14E67795FF90612BFEDC6712FDBDFA92" box="[329,567,1358,1383]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF90612BFEDC6712FDBDFA92" box="[329,567,1358,1383]" italics="true" pageId="65" pageNumber="66">Triodontus bicavatus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="14E67795FF90612BFC986713FC06FA92" box="[781,908,1358,1383]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="E192D004FF90612BFC986713FC06FA92" box="[781,908,1358,1383]" italics="true" pageId="65" pageNumber="66">T. nitidulus</emphasis>
</taxonomicName>
. Males of the former species can be separated by having abdominal sternite 8 rather wide, not sinuate medially, and with oval transverse concavity, and also by having the lateral lobes of the parameres with small keel-shaped processes on proximal parts. Females of these species are difficult to separate.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF906129FF526782FC7BFD5D" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66" type="description">
<paragraph id="D3590C16FF90612BFF526782FE05F9EE" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">
<emphasis id="E192D004FF90612BFF526782FE29FA02" bold="true" box="[199,419,1502,1527]" pageId="65" pageNumber="66">Description. Male.</emphasis>
Color of head, pronotum, and elytra dark brown; legs, antennae, and underside of the body brown (
<figureCitation id="4BDD1093FF90612BFF7B645EFEC0F9EE" box="[238,330,1538,1563]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="65" pageNumber="66">Figs. 33</figureCitation>
AB).
</paragraph>
<paragraph id="D3590C16FF90612BFF52647AFD8CF906" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">Frontoclypeus convex anteriorly, rounded laterally, anterior margin crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with small tubercles mediad of each eye and in some specimens with a horn in the center of the frontoclypeus. The horn, when the most developed, is about as long as the width of the head, acutely rounded apically, straight to somewhat curved caudally at the apex, slightly rugose on posterior side.</paragraph>
<paragraph id="D3590C16FF90612BFF5264A3FBFBF852" blockId="65.[151,1437,890,1995]" pageId="65" pageNumber="66">Pronotum with a bulge medially, with excavations aside the bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge has 3 small but distinct tubercles. Lateral margins with wide border appearing somewhat crenulate in dorsal view, with a row of brown setae. Anterior margin with wide, smooth border. Posterior margin with fine border, not crenulate, punctate with narrow longitudinal punctures. Surface of most pronotum smooth, anterior and posterior angles with sparse and coarse punctation.</paragraph>
<paragraph id="D3590C16FF90612BFF5265EEFC7DF83E" blockId="65.[151,1437,890,1995]" box="[199,1015,1970,1995]" pageId="65" pageNumber="66">Scutellum rounded apically, its visible part is about 1/12 length of elytra.</paragraph>
<caption id="87995C9EFF936128FF026528FE15F832" httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="66" pageNumber="67" targetBox="[176,1399,268,1883]" targetPageId="66">
<paragraph id="D3590C16FF936128FF026528FE15F832" blockId="66.[151,1435,1908,1991]" pageId="66" pageNumber="67">
<emphasis id="E192D004FF936128FF026528FEABF87F" bold="true" box="[151,289,1908,1930]" pageId="66" pageNumber="67">FIGURE 33.</emphasis>
<emphasis id="E192D004FF936128FEBD6528FD8CF87F" box="[296,518,1908,1930]" italics="true" pageId="66" pageNumber="67">
<taxonomicName id="14E67795FF936128FEBD6528FD88F87F" box="[296,514,1908,1930]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicavatus">Triodontus bicavatus</taxonomicName>
.
</emphasis>
Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H).
</paragraph>
</caption>
<paragraph id="D3590C16FF926129FF5262CBFE81FEE9" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">Elytra convex, with distinct humeral humps, their maximum width is approximately at the middle. Elytra with 10 distinct striae on disc and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria with small punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures.</paragraph>
<paragraph id="D3590C16FF926129FF52637BFB7AFEB5" blockId="67.[151,1437,151,824]" box="[199,1264,295,320]" pageId="67" pageNumber="68">
Sternite 8 medially feebly sinuate (in ventral view), with elongate transversal fovea (
<figureCitation id="4BDD1093FF926129FBEE637BFB47FEB5" box="[1147,1229,295,320]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="67" pageNumber="68">Fig. 33</figureCitation>
G).
</paragraph>
<paragraph id="D3590C16FF926129FF526311FB1BFE59" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">
Parameres (
<figureCitation id="4BDD1093FF926129FEDF6310FE10FE91" box="[330,410,332,357]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="67" pageNumber="68">Fig. 33</figureCitation>
F) with large outer lobes covering inner lobes (in lateral view). Proximal parts of outer lobes with small keel-shaped processes. Internal sac of the aedeagus with 1 strongly-sclerotized, somewhat horseshoeshaped, symmetrical sclerite and with 2 similar protuberances of the membrane (
<figureCitation id="4BDD1093FF926129FB8A63C8FBFBFE59" box="[1055,1137,404,429]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="67" pageNumber="68">Fig. 33</figureCitation>
E).
</paragraph>
<paragraph id="D3590C16FF926129FF5263EBFC66FDED" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">
<emphasis id="E192D004FF926129FF5263EBFEAFFE25" bold="true" box="[199,293,439,464]" pageId="67" pageNumber="68">Female.</emphasis>
Females (
<figureCitation id="4BDD1093FF926129FE0863EBFE7AFE25" box="[413,496,439,464]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="67" pageNumber="68">Fig. 33</figureCitation>
C) differ from males in having long apical spur on protibiae, absence of head and pronotum armature including tubercles mediad of each eye, densely punctate dorsal side of head, and longer abdominal sternites including 6th sternite, which is evenly rounded apically.
</paragraph>
<paragraph id="D3590C16FF926129FF526078FC7BFD5D" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">
<emphasis id="E192D004FF926129FF526078FEB6FDC8" bold="true" box="[199,316,548,573]" pageId="67" pageNumber="68">Variation.</emphasis>
The variation is similar to that of
<taxonomicName id="14E67795FF926129FD446079FCDAFDC8" box="[721,848,548,573]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="E192D004FF926129FD446079FCDAFDC8" box="[721,848,548,573]" italics="true" pageId="67" pageNumber="68">T. nitidulus</emphasis>
</taxonomicName>
: pronotal and head armature of males is subject to allometric variability from fully developed (as described above) to almost undeveloped (with the males having small frontoclypeal tubercles in the middle and small fossa on the pronotum anteriorly). Body length of examined specimens varied from
<quantity id="141EA1F3FF926129FE0860D3FDAEFD52" box="[413,548,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="8.2" metricValueMin="6.5" pageId="67" pageNumber="68" unit="mm" value="7.35" valueMax="8.2" valueMin="6.5">6.58.2 mm</quantity>
(males) and from
<quantity id="141EA1F3FF926129FD6760D3FCF3FD52" box="[754,889,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="67" pageNumber="68" unit="mm" value="7.0" valueMax="7.5" valueMin="6.5">6.57.5 mm</quantity>
(females).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF926129FF5260E8FB1AFD05" pageId="67" pageNumber="68" type="discussion">
<paragraph id="D3590C16FF926129FF5260E8FB1AFD05" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">
<emphasis id="E192D004FF926129FF5260E8FEB1FD38" bold="true" box="[199,315,692,717]" pageId="67" pageNumber="68">Remarks.</emphasis>
<bibRefCitation id="B77771E7FF926129FEDD60E8FE7EFD39" author="Paulian" box="[328,500,692,717]" pageId="67" pageNumber="68" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian (1977)</bibRefCitation>
considered
<taxonomicName id="14E67795FF926129FD1C60E9FC91FD38" box="[649,795,692,717]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="E192D004FF926129FD1C60E9FC91FD38" box="[649,795,692,717]" italics="true" pageId="67" pageNumber="68">O. obsoletus</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="14E67795FF926129FBB960E9FB33FD38" box="[1068,1209,692,717]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF926129FBB960E9FB33FD38" box="[1068,1209,692,717]" italics="true" pageId="67" pageNumber="68">T. bicavatus</emphasis>
</taxonomicName>
, however the
<typeStatus id="0C5DB2B4FF926129FAFE60E9FA16FD38" box="[1387,1436,693,717]" pageId="67" pageNumber="68">type</typeStatus>
specimen of the former species has all the diagnostic characters of
<taxonomicName id="14E67795FF926129FC116085FB8BFD05" box="[900,1025,727,752]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="E192D004FF926129FC116085FB8BFD05" box="[900,1025,727,752]" italics="true" pageId="67" pageNumber="68">T. nitidulus</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF926129FF5260A0FE52FCCD" pageId="67" pageNumber="68" type="distribution">
<paragraph id="D3590C16FF926129FF5260A0FE52FCCD" blockId="67.[151,1437,151,824]" pageId="67" pageNumber="68">
<emphasis id="E192D004FF926129FF5260A0FEEAFCE0" bold="true" box="[199,352,764,789]" pageId="67" pageNumber="68">Distribution.</emphasis>
<taxonomicName id="14E67795FF926129FEFD60A1FE7AFCE0" box="[360,496,764,789]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FF926129FEFD60A1FE7AFCE0" box="[360,496,764,789]" italics="true" pageId="67" pageNumber="68">T. bicavatus</emphasis>
</taxonomicName>
is known from a few localities in northern
<collectingCountry id="ABF14C86FF926129FC4960A0FBE9FCE0" box="[988,1123,764,789]" name="Madagascar" pageId="67" pageNumber="68">Madagascar</collectingCountry>
including the northernmost part of the island (
<figureCitation id="4BDD1093FF926129FEF66143FE3FFCCD" box="[355,437,799,824]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="67" pageNumber="68">Fig. 33</figureCitation>
H).
</paragraph>
</subSubSection>
</treatment>
</document>