518 lines
57 KiB
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518 lines
57 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.915.47626" ID-GBIF-Dataset="6400d265-1599-4f6b-b65d-cc0b7be75d85" ID-PMC="PMC7052023" ID-Pensoft-Pub="1313-2970-915-25" ID-Pensoft-UUID="4DC506D822E85F2F8882C245C69416FC" ID-PubMed="32148422" ID-ZooBank="EA75DB913C374037A242C6F1C144F7CC" ModsDocID="1313-2970-915-25" checkinTime="1582662221248" checkinUser="pensoft" docAuthor="Zhang, Chao & Martens, Jochen" docDate="2020" docId="936A11B763CA58278751BFC806B77B27" docLanguage="en" docName="ZooKeys 915: 25-58" docOrigin="ZooKeys 915" docSource="http://dx.doi.org/10.3897/zookeys.915.47626" docTitle="Toccolus kuryi Zhang & Martens 2020, sp. nov." docType="treatment" docUuid="F49E92F7-8667-435F-A842-438280CEE8CA" docUuidSource="ZooBank" docVersion="5" id="4DC506D822E85F2F8882C245C69416FC" lastPageNumber="25" masterDocId="4DC506D822E85F2F8882C245C69416FC" masterDocTitle="A taxonomic study on Epedanidae from Thailand including functional aspects of male genital morphology (Opiliones, Laniatores)" masterLastPageNumber="58" masterPageNumber="25" pageNumber="25" updateTime="1668168517460" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic study on Epedanidae from Thailand including functional aspects of male genital morphology (Opiliones, Laniatores)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Chao</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martens, Jochen</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>915</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>58</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.915.47626</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.915.47626</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-915-25</mods:identifier>
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<mods:identifier type="ZooBank">EA75DB913C374037A242C6F1C144F7CC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4DC506D822E85F2F8882C245C69416FC</mods:identifier>
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<treatment ID-GBIF-Taxon="162267202" LSID="urn:lsid:zoobank.org:act:F49E92F7-8667-435F-A842-438280CEE8CA" httpUri="http://treatment.plazi.org/id/936A11B763CA58278751BFC806B77B27" lastPageNumber="25" pageId="0" pageNumber="25">
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<subSubSection pageId="0" pageNumber="25" type="nomenclature">
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName LSID="936A11B7-63CA-5827-8751-BFC806B77B27" authority="Zhang & Martens, 2020" authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi" status="sp. nov.">Toccolus kuryi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="25">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">Figs 102-109</figureCitation>
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">, 110-117</figureCitation>
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<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">, 118-123</figureCitation>
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<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">, 124-129</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="25" type="type material">
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<paragraph pageId="0" pageNumber="25">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Holotype</emphasis>
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male (SMF-CJM7065): THAILAND: Loel Province: Nang Hin District, near cave Wat Tham Malohan (No.18a),
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<geoCoordinate degrees="17" direction="north" minutes="06.47" orientation="latitude" precision="9" value="17.107834">17°06.47'N</geoCoordinate>
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<geoCoordinate degrees="101" direction="east" minutes="52.69" orientation="longitude" precision="9" value="101.87817">101°52.69'E</geoCoordinate>
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, alt. 362 m, 2 November 2010, S. Dashdamirov leg.
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<emphasis bold="true" pageId="0" pageNumber="25">
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<emphasis italics="true" pageId="0" pageNumber="25">Paratype</emphasis>
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.
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</emphasis>
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One female (SMF-CJM7063): THAILAND: Nong Khai Province: Sangkhom District, Tham Tip Waterfall (No. 23),
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<geoCoordinate degrees="18" direction="north" minutes="07.57" orientation="latitude" precision="9" value="18.126167">18°07.57'N</geoCoordinate>
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<geoCoordinate degrees="102" direction="east" minutes="11.16" orientation="longitude" precision="9" value="102.186">102°11.16'E</geoCoordinate>
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, alt. 198 m, 3 November 2010, S. Dashdamirov leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="25" type="diagnosis">
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<paragraph pageId="0" pageNumber="25">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="25">
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The distal part of glans nearly triangular (ventral and dorsal views, Figs
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<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">119</figureCitation>
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,
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<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">121</figureCitation>
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) and the tip curved ventrally (lateral view, Fig.
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<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">120</figureCitation>
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). Basichelicerite dorsally with three spines (Figs
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<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">103</figureCitation>
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,
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<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">104</figureCitation>
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,
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">112</figureCitation>
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,
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">113</figureCitation>
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). Pedipalpal femur ventrally with a row of six setiferous tubercles, the longest one at the base (Figs
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<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">108</figureCitation>
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,
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">116</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="25" type="etymology">
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<paragraph pageId="0" pageNumber="25">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="25">
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The specific name is a patronym in honor of Dr. Adriano B. Kury, a well-known arachnologist from Brazil. A.
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<bibRefCitation author="Kury, AB" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="25" publicationUrl="http://www.museunacional.ufrj.br/mndi/Aracnologia/opiliones.html" refId="B4" refString="Kury, AB, 2000. Classification of Opiliones. Museu Nacional/UFRJ website. http://www.museunacional.ufrj.br/mndi/Aracnologia/opiliones.html" title="Classification of Opiliones. Museu Nacional / UFRJ website." url="http://www.museunacional.ufrj.br/mndi/Aracnologia/opiliones.html" year="2000">Kury (2000)</bibRefCitation>
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created a website of
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<taxonomicName authorityName="Roewer" authorityYear="1927" lsidName="" order="Opiliones" pageId="0" pageNumber="25" rank="order">Opiliones</taxonomicName>
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and built a platform for acquiring knowledge accessible for everyone. He redefined the family
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<taxonomicName authorityName="Sorensen" authorityYear="1886" family="Epedanidae" lsidName="" pageId="0" pageNumber="25" rank="family">Epedanidae</taxonomicName>
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to include four subfamilies (
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<bibRefCitation author="Kury, AB" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="25" refId="B3" refString="Kury, AB, 1993. Analise filogenetica de Gonyleptoidea (Arachnida, Opiliones, Laniatores). PhD Thesis, Universidade de Sao Paulo, Sao Paulo." title="Analise filogenetica de Gonyleptoidea (Arachnida, Opiliones, Laniatores). PhD Thesis, Universidade de Sao Paulo, Sao Paulo." year="1993">Kury 1993</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="25" type="description">
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<paragraph pageId="0" pageNumber="25">Description.</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<emphasis bold="true" pageId="0" pageNumber="25">Male</emphasis>
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(holotype) habitus as in Figs
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<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">102</figureCitation>
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,
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">110</figureCitation>
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,
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<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">124-126</figureCitation>
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. Coloration (Figs
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<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">124-126</figureCitation>
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): body light yellow with brown patches; median area of prosoma with brown reticulations; both lateral ridges of scutum with brown stripes; opisthosomal areae I-III each with a transverse brownish band; area IV with a series of three brown flecks; all free tergites brown; all coxae yellow with brown reticulations distally; free sternites each with brown bands, these somewhat paler on the central portion; chelicerae and pedipalps reticulated; trochanters pale yellow, femur, patella, tibia and metatarsus with black reticulations, tarsus lighter.
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</paragraph>
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<caption doi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25" start="Figures 102–109" startId="F13">
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<paragraph pageId="0" pageNumber="25">
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<emphasis bold="true" pageId="0" pageNumber="25">Figures 102-109.</emphasis>
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<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
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<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
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</taxonomicName>
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sp. nov., male holotype
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<emphasis bold="true" pageId="0" pageNumber="25">102</emphasis>
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body, lateral view
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<emphasis bold="true" pageId="0" pageNumber="25">103</emphasis>
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left chelicera, medial view
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<emphasis bold="true" pageId="0" pageNumber="25">104</emphasis>
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same, lateral view
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<emphasis bold="true" pageId="0" pageNumber="25">105</emphasis>
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left basichelicerite, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="25">106</emphasis>
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left cheliceral hand, frontal view
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<emphasis bold="true" pageId="0" pageNumber="25">107</emphasis>
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left pedipalp, medial view
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<emphasis bold="true" pageId="0" pageNumber="25">108</emphasis>
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same, lateral view
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<emphasis bold="true" pageId="0" pageNumber="25">109</emphasis>
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right pedipalp, medial view. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="25">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Dorsum</emphasis>
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(Figs
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<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">110</figureCitation>
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,
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<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">124</figureCitation>
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). Scutum trapezoid in appearance, both parts nearly parallel-sided, the widest portion of body at opisthosomal area IV, posterior end of opisthosoma bluntly rounded. Prosoma with a row of ten sharply pointed tubercles along anterior margin. Dorsal scutum smooth. Ocularium oval, removed from anterior border of prosoma scutum by 0.38 mm, armed with a conspicuous short median spine of about the height of ocularium (lateral view), four small tubercles above both eyes. The area behind ocularium raised, and the posterior margin of prosoma smoothly rounded. The anterior margin of opisthosomal area I markedly bowed out, borders of other scutal areae slightly bowed out. Opisthosomal areae I-IV with a row of seta-tipped tubercles each and a longitudinal row of similar tubercles on left and right lateral margins. Free tergites with a transverse row of seta-tipped tubercles and anal operculum with scattered granules.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Venter</emphasis>
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(Fig.
|
||
<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">126</figureCitation>
|
||
). Coxae with tubercular surface, coxa I additionally with enlarged seta-tipped tubercles. Coxae II-IV with minute tubercles. Genital operculum and free sternites with seta-tipped granules. Spiracles clearly visible.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Chelicerae</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">103-106</figureCitation>
|
||
). Basichelicerite elongate, armed with two long, curved spines, one short spine dorsally; another one distally on the ectal side; six seta-tipped tubercles scattered over the ventral side; medial side with a basal protuberance (Fig.
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">103</figureCitation>
|
||
). Cheliceral hand considerably widened, pro-dorsally with few seta-tipped tubercles, the inner ones larger than the outer ones; ectally with two seta-tipped tubercles at the subdistal portion; one enlarged seta-tipped tubercle and a smaller one on the ventral side. Fingers strong, inner edges toothed (Fig.
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">106</figureCitation>
|
||
): moveable finger with five teeth, the proximal one cut rectangularly, the other four pointed; fixed finger with five pointed teeth.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25" start="Figures 110–117" startId="F14">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figures 110-117.</emphasis>
|
||
<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male (Fig.
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">110</figureCitation>
|
||
, holotype) and female (Figs
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">111-117</figureCitation>
|
||
, paratype)
|
||
<emphasis bold="true" pageId="0" pageNumber="25">110, 111</emphasis>
|
||
body, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">112</emphasis>
|
||
right chelicera, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">113</emphasis>
|
||
same, medial view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">114</emphasis>
|
||
right basichelicerite, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">115</emphasis>
|
||
right cheliceral hand, frontal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">116</emphasis>
|
||
right pedipalp, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">117</emphasis>
|
||
same, medial view. Scale bars: 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="25">110-113, 116, 117</emphasis>
|
||
), 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="25">114, 115</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Pedipalps</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">107-109</figureCitation>
|
||
). Coxa dorsally with three seta-tipped tubercles, one strong distal, one sub-distal exterior to it, and a blunt one situated proximally; ventrally with two seta-tipped tubercles. Trochanter ventrally with two setiferous tubercles and dorsally with an enlarge one and a small one. Femur ventrally with a row of five (right pedipalp, Fig.
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">102</figureCitation>
|
||
) or six (left pedipalp, Fig.
|
||
<figureCitation captionStart="Figures 102–109" captionStartId="F13" captionText="Figures 102 - 109. Toccolus kuryi sp. nov., male holotype 102 body, lateral view 103 left chelicera, medial view 104 same, lateral view 105 left basichelicerite, dorsal view 106 left cheliceral hand, frontal view 107 left pedipalp, medial view 108 same, lateral view 109 right pedipalp, medial view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures102-109" httpUri="https://binary.pensoft.net/fig/383033" pageId="0" pageNumber="25">108</figureCitation>
|
||
) setiferous tubercles, the longest one at the base of the row; dorsally with a row of ten setiferous tubercles, distal one and proximal one inconspicuous; disto-medially side with two setiferous tubercles. Patella ventro-mesally with two and ventro-ectally with one setiferous tubercles. Tibia ventro-mesally with three and ventro-ectally with four setiferous tubercles. Tarsus with three setiferous tubercles each ventro-ectally and ventro-medially. Tarsal claw massive, curved, longer than tarsus.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
All segments unarmed, nearly smooth. Legs I-II slender, legs III-IV strong. Femora I-III not curved, almost straight, femur IV slightly curved. Distitarsi I and II with two tarsomeres. Distitarsi III and IV without scopula and with two unpectinated claws. Tarsal formula (I-IV): 7(2)/16(2)/7/7.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Penis</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">118-121</figureCitation>
|
||
). Shaft slender, nearly parallel-sided, apical part (pars distalis) distended. Ventral plate conspicuously constricted apically, and dorsally with deep indented near the glans. Glans partially sunken into dorsal depressed portion of pars distalis and extended the distal margin of the ventral plate (Fig.
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">120</figureCitation>
|
||
). The distal part of glans almost triangular (ventral and dorsal views, Figs
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">119</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">121</figureCitation>
|
||
), the tip curved ventrally. Spination symmetrical. One pair of setae A, B, and F. Two pairs of setae C1-2, D1-2, and E1-2 (Figs
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">120</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">121</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="female">
|
||
<paragraph pageId="0" pageNumber="25">Female</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
(SMF-CJM7063) (Figs
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">111-117</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">122</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">123</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">127-129</figureCitation>
|
||
). Body nearly oval. Granulation and spination of body similar to the male (Figs
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">111</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">127</figureCitation>
|
||
). Prosoma with a row of eight sharp pointed tubercles along anterior margin (Fig.
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">111</figureCitation>
|
||
). Surface of dorsum smooth. Ocularium with only one conspicuous tubercle near the left eye except for the long median spine. Chelicerae of normal shape, not enlarged, slight difference in inner edges of cheliceral finger (Fig.
|
||
<figureCitation captionStart="Figures 110–117" captionStartId="F14" captionText="Figures 110 - 117. Toccolus kuryi sp. nov., male (Fig. 110, holotype) and female (Figs 111 - 117, paratype) 110, 111 body, dorsal view 112 right chelicera, lateral view 113 same, medial view 114 right basichelicerite, dorsal view 115 right cheliceral hand, frontal view 116 right pedipalp, lateral view 117 same, medial view. Scale bars: 1 mm (110 - 113, 116, 117), 0.5 mm (114, 115)." figureDoi="10.3897/zookeys.915.47626.figures110-117" httpUri="https://binary.pensoft.net/fig/383034" pageId="0" pageNumber="25">115</figureCitation>
|
||
). Femora of pedipalpi ventrally with six and dorsally with a row of five setiferous tubercles, disto-medially with three setiferous tubercles. Tarsal formula (I-IV): 6(2)/15(2)/7/7.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="25">Ovipositor</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">122</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 118–123" captionStartId="F15" captionText="Figures 118 - 123. Toccolus kuryi sp. nov., genitalia of male holotype 118 penis, dorsal view 119 pars distalis of penis, dorsal view 120 same, lateral view 121 same, ventral view 122 ovipositor, ventral view 123 same, dorsal view. Abbreviations: G glans Pb pars basalis Pd pars distalis VP ventral plate. Scale bars: 0.5 mm (118), 0.25 mm (119 - 123)." figureDoi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25">123</figureCitation>
|
||
). Ventral side with four, dorsal side with six setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Measurements.</emphasis>
|
||
Male (female): body 2.27 (2.76) long, 2.08 (1.96) wide at widest portion, scutum 1.95 (1.79) long. Ocularium 0.41 (0.30) long, 0.69 (0.51) wide. Proximal chelicerae 1.24 (0.84) long, 0.47 (0.31) wide; second 2.20 (1.37) long, 0.78 (0.45) wide; distal 0.97 (0.62) long, 0.22 (0.14) wide. Pedipalp claw 0.82 (0.79) long. Penis 1.02 long. Measurements of pedipalp and legs as in Tables
|
||
<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Toccolus kuryi sp. nov. Measurements of the pedipalp and legs of the male holotype, as length / width." httpUri="http://table.plazi.org/id/45B99BDCA9980309B74D96F03ADDCE3F" pageId="0" pageNumber="25" tableUuid="45B99BDCA9980309B74D96F03ADDCE3F">7</tableCitation>
|
||
and
|
||
<tableCitation captionStart="Table 8" captionStartId="T8" captionText="Table 8. Toccolus kuryi sp. nov. Measurements of the pedipalp and legs of the female paratype, as length / width." httpUri="http://table.plazi.org/id/A412936EB67A11FF241A3AE11373F51D" pageId="0" pageNumber="25" tableUuid="A412936EB67A11FF241A3AE11373F51D">8</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="45B99BDCA9980309B74D96F03ADDCE3F" httpUri="http://table.plazi.org/id/45B99BDCA9980309B74D96F03ADDCE3F" pageId="0" pageNumber="25" start="Table 7" startId="T7">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Table 7.</emphasis>
|
||
<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Measurements of the pedipalp and legs of the male holotype, as length/width.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<table pageId="0" pageNumber="25">
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Trochanter</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Femur</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Patella</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Tibia</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Metatarsus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Tarsus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Total</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Pedipalp</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.56/0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.56/0.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.98/0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.95/0.35</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.70/0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">4.75</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg I</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.35/0.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.60/0.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.52/0.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.12/0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.94/0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.06/0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">6.59</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg II</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.34/0.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.15/0.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.59/0.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.86/0.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.74/0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.94/0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">9.62</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg III</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.42/0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.86/0.22</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.61/.032</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.30/0.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.21/0.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.03/0.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">7.43</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.48/0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.32/0.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.63/0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.76/0.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.71/0.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.15/0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">9.05</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-Table-UUID="A412936EB67A11FF241A3AE11373F51D" httpUri="http://table.plazi.org/id/A412936EB67A11FF241A3AE11373F51D" pageId="0" pageNumber="25" start="Table 8" startId="T8">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Table 8.</emphasis>
|
||
<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Measurements of the pedipalp and legs of the female paratype, as length/width.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<table pageId="0" pageNumber="25">
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Trochanter</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Femur</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Patella</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Tibia</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Metatarsus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Tarsus</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Total</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Pedipalp</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.42/0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.28/0.23</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.78/0.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.87/0.27</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.66/0.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">4.01</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg I</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.24/0.22</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.24/0.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.48/0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.93/0.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.54/0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.64/0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">5.71</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg II</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.22/0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.72/0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.52/0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.51/0.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.27/0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.46/0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">8.70</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg III</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.26/0.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.35/0.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.53/0.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.07/0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.79/0.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.76/0.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">5.76</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="25">
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Leg IV</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.32/0.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.67/0.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.56/0.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.39/0.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2.48/0.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1.13/0.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">7.55</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="habitat">
|
||
<paragraph pageId="0" pageNumber="25">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
The specimens were collected under stones (SMF-CJM7065) and under
|
||
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="0" pageNumber="25" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Acacia</emphasis>
|
||
</taxonomicName>
|
||
bark and rotten stumps (SMF-CJM7063).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="distribution">
|
||
<paragraph pageId="0" pageNumber="25">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">Thailand (Loel Province, Nong Khai Province).</paragraph>
|
||
<caption doi="10.3897/zookeys.915.47626.figures118-123" httpUri="https://binary.pensoft.net/fig/383035" pageId="0" pageNumber="25" start="Figures 118–123" startId="F15">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figures 118-123.</emphasis>
|
||
<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., genitalia of male holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="25">118</emphasis>
|
||
penis, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">119</emphasis>
|
||
pars distalis of penis, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">120</emphasis>
|
||
same, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">121</emphasis>
|
||
same, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">122</emphasis>
|
||
ovipositor, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">123</emphasis>
|
||
same, dorsal view. Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="25">G</emphasis>
|
||
glans
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Pb</emphasis>
|
||
pars basalis
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Pd</emphasis>
|
||
pars distalis
|
||
<emphasis bold="true" pageId="0" pageNumber="25">VP</emphasis>
|
||
ventral plate. Scale bars: 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="25">118</emphasis>
|
||
), 0.25 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="25">119-123</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25" start="Figures 124–129" startId="F16">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figures 124-129.</emphasis>
|
||
Photographs of male (Figs
|
||
<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">124-126</figureCitation>
|
||
, holotype) and female (Figs
|
||
<figureCitation captionStart="Figures 124–129" captionStartId="F16" captionText="Figures 124 - 129. Photographs of male (Figs 124 - 126, holotype) and female (Figs 127 - 129, paratype) of Toccolus kuryi sp. nov. 124, 127 body and parts of appendages, dorsal view 125, 128 body and parts of appendages, lateral view 126, 129 body and parts of appendages, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.915.47626.figures124-129" httpUri="https://binary.pensoft.net/fig/383036" pageId="0" pageNumber="25">127-129</figureCitation>
|
||
, paratype) of
|
||
<taxonomicName authorityName="Zhang & Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Toccolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toccolus kuryi" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="kuryi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Toccolus kuryi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="25">124, 127</emphasis>
|
||
body and parts of appendages, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">125, 128</emphasis>
|
||
body and parts of appendages, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="25">126, 129</emphasis>
|
||
body and parts of appendages, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |