201 lines
35 KiB
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201 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.119781" ID-Pensoft-Pub="1313-2970-1198-17" ID-Pensoft-UUID="9A0A0A5CD22451C7ADAE5C8460C568DC" ID-ZooBank="BDC5B2CD1BB442AE8672E57CC0FBBF6F" ModsDocID="1313-2970-1198-17" checkinTime="1713962616471" checkinUser="pensoft" docAuthor="Dyachkov, Yurii V. & Bonato, Lucio" docDate="2024" docId="C4E48964648050C1BC78CC928941CF3F" docLanguage="en" docName="ZooKeys 1198: 17-54" docOrigin="ZooKeys 1198" docPubDate="2024-04-23" docSource="http://dx.doi.org/10.3897/zookeys.1198.119781" docTitle="Escaryus retusidens Attems 1904" docType="treatment" docVersion="1" id="9A0A0A5CD22451C7ADAE5C8460C568DC" lastPageNumber="17" masterDocId="9A0A0A5CD22451C7ADAE5C8460C568DC" masterDocTitle="An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia" masterLastPageNumber="54" masterPageNumber="17" pageNumber="17" updateTime="1713962616471" updateUser="pensoft">
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<mods:title id="8158D9A09DBC7E591DF9FEB2D42DF640">An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia</mods:title>
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<mods:namePart id="3D0D5A7F03CC0D57BD59AA60168EAAE0">Dyachkov, Yurii V.</mods:namePart>
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<mods:affiliation id="7F7C92B421B59759A8B08A1FA2131086">Altai State University, Lenin Avenue, 61, 656049, Barnaul, Russia & Tomsk State University, Lenin Avenue, 36, 634050, Tomsk, Russia & Western Caspian University, Istiglaliyyat Street, 31, Baku, Azerbaijan</mods:affiliation>
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<mods:nameIdentifier id="3359604CC0AFA3D9665CFD0797A8043B" type="email">dyachkov793@mail.ru</mods:nameIdentifier>
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<mods:namePart id="B8F786F78891B942AE92EF39BF1EC183">Bonato, Lucio</mods:namePart>
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<mods:nameIdentifier id="BB75B5BA6760240D1479DDA6E036788C" type="ORCID">https://orcid.org/0000-0002-8312-7570</mods:nameIdentifier>
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<mods:affiliation id="2B39D029842C2C0C0F9EB1467035F4B8">Dipartimento di Biologia, Universita di Padova, via U. Bassi 58 b, 35131 Padova, Italy</mods:affiliation>
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<mods:typeOfResource id="1334FA4701A61D661ECFD744E2EDDD5D">text</mods:typeOfResource>
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<mods:date id="93FF0DDFE4B99A9304F79B255AA4CAF1">2024</mods:date>
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<mods:number id="EC8233ADED8241D20FC5943F581FB7B0">2024-04-23</mods:number>
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<mods:number id="D1BCD6DE8ADDFA100B5774D76CB7E084">1198</mods:number>
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<mods:start id="ECEB8ABFF1F4CF0FF06964CCBCA74948">17</mods:start>
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<mods:classification id="ADD32CD5C08FDD006507FC96305BBFBE">journal article</mods:classification>
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<mods:identifier id="F73EC44FC997BA1869F530FC61D3D332" type="DOI">http://dx.doi.org/10.3897/zookeys.1198.119781</mods:identifier>
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<treatment id="C4E48964648050C1BC78CC928941CF3F" LSID="urn:lsid:plazi:treatment:C4E48964648050C1BC78CC928941CF3F" httpUri="http://treatment.plazi.org/id/C4E48964648050C1BC78CC928941CF3F" lastPageNumber="17" pageId="0" pageNumber="17">
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<subSubSection id="8AC44F73F80D54C05A37530ED4472FD3" pageId="0" pageNumber="17" type="nomenclature">
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<paragraph id="760063AC50DFB07433EC8D8EEF556514" pageId="0" pageNumber="17">
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35.
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<taxonomicName id="914738FE35308817F8158F7BC7A70602" ID-CoL="3BGLS" LSID="C4E48964-6480-50C1-BC78-CC928941CF3F" authority="Attems, 1904" authorityName="Attems" authorityYear="1904" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">Escaryus retusidens Attems, 1904</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C275C319741FF1F4A121C31B0EA1C593" pageId="0" pageNumber="17" type="reference_group">
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<paragraph id="6818C2FF78EA37519EF20F73E2AFE238" pageId="0" pageNumber="17">
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<taxonomicName id="90878974D1839849CEB1C7A74FED37FE" authorityName="Attems" authorityYear="1904" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">Escaryus retusidens</taxonomicName>
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<bibRefCitation id="8BCC3DBCAB9F594FEA06D4E77BD65D96" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems 1904</bibRefCitation>
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: 121.
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</paragraph>
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<paragraph id="76B02A0DF680663BC14B9E1C8E868AC1" pageId="0" pageNumber="17">
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<taxonomicName id="A0EA8122382E16693BDB9FABB6D79C0D" authorityName="Attems" authorityYear="1904" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">Escaryus retusidens</taxonomicName>
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-
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<bibRefCitation id="1FEBF5D8E15207D693D4AAB346F5712B" DOI="https://doi.org/10.1515/9783111430638" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="17" pagination="1 - 388" refId="B7" refString="Attems, C, 1929. Myriapoda. 1. Geophilomorpha. Das Tierreich 52: 1 - 388, DOI: https://doi.org/10.1515/9783111430638" title="Myriapoda. 1. Geophilomorpha." url="https://doi.org/10.1515/9783111430638" volume="52" year="1929">Attems 1929</bibRefCitation>
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: 96;
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<bibRefCitation id="4DACBD0ACDD074BD75B966E6FEC19E59" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
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: 135, 116;
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<bibRefCitation id="6475149167E937F1166562A590239253" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
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: 110;
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<bibRefCitation id="74E50E0E3B9AF5284741A1327BB5D1B9" DOI="https://doi.org/10.1134/S0013873816040138" author="Volkova, YuS" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="17" pagination="669 - 678" refId="B113" refString="Volkova, YuS, 2016. An annotated catalogue of geophilomorph centipedes (Chilopoda, Geophilomorpha) from the European part of Russia. Zoologicheskii Zhurnal 95 (6): 669 - 678, DOI: https://doi.org/10.1134/S0013873816040138" title="An annotated catalogue of geophilomorph centipedes (Chilopoda, Geophilomorpha) from the European part of Russia." url="https://doi.org/10.1134/S0013873816040138" volume="95" year="2016">Volkova 2016</bibRefCitation>
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: 675;
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<bibRefCitation id="1BCB64554320F7DC9ECA673DB27EA9CD" DOI="https://doi.org/10.15298/arthsel.25.1.03" author="Zuev, RV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="23 - 38" refId="B117" refString="Zuev, RV, 2016. Centipedes (Chilopoda) from the Stavropol Territory, northern Caucasus, Russia. Arthropoda Selecta 25 (1): 23 - 38, DOI: https://doi.org/10.15298/arthsel.25.1.03" title="Centipedes (Chilopoda) from the Stavropol Territory, northern Caucasus, Russia." url="https://doi.org/10.15298/arthsel.25.1.03" volume="25" year="2016">Zuev 2016</bibRefCitation>
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: 33;
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<bibRefCitation id="D0C348F5486316DDFD66FF1060C095FD" DOI="https://doi.org/10.15298/arthsel.26.1.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="8 - 14" refId="B69" refString="Nefediev, PS, Tuf, IH, Farzalieva, GS, 2017a. Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha. Arthropoda Selecta 26 (1): 8 - 14, DOI: https://doi.org/10.15298/arthsel.26.1.02" title="Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha." url="https://doi.org/10.15298/arthsel.26.1.02" volume="26" year="2017 a">Nefediev et al. 2017a</bibRefCitation>
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: 11;
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<bibRefCitation id="8C1B39AAA27FCCAD09CFF36DD67924AC" DOI="https://doi.org/10.15298/arthsel.26.3.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="217 - 224" refId="B71" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, 2017c. A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha). Arthropoda Selecta 26 (3): 217 - 224, DOI: https://doi.org/10.15298/arthsel.26.3.02" title="A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha)." url="https://doi.org/10.15298/arthsel.26.3.02" volume="26" year="2017 c">2017c</bibRefCitation>
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: 222;
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<bibRefCitation id="9C573ED336E3AF965A09D25632B20340" DOI="https://doi.org/10.3897/zookeys.741.21936" author="Nefediev, PS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="17" pagination="219 - 254" refId="B72" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, Nedoev, KK, Niyazov, ST, 2018. Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda). ZooKeys 741: 219 - 254, DOI: https://doi.org/10.3897/zookeys.741.21936" title="Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda)." url="https://doi.org/10.3897/zookeys.741.21936" volume="741" year="2018">2018</bibRefCitation>
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: 239;
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<bibRefCitation id="671C36525AC320A8D54BAF4D3FF109D2" DOI="https://doi.org/10.15298/arthsel.27.4.04" author="Dyachkov, YuV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="293 - 299" refId="B40" refString="Dyachkov, YuV, Tuf, IH, 2018. New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan. Arthropoda Selecta 26 (4): 293 - 299, DOI: https://doi.org/10.15298/arthsel.27.4.04" title="New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan." url="https://doi.org/10.15298/arthsel.27.4.04" volume="26" year="2018">Dyachkov and Tuf 2018</bibRefCitation>
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: 295;
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<bibRefCitation id="DC4D187B4ED9FD1D5A454ACAA175170A" DOI="https://doi.org/10.25221/fee.380.4" author="Nefediev, PS" journalOrPublisher="Far Eastern Entomologist = Dal'nevostochnyi Entomolog" pageId="0" pageNumber="17" pagination="23 - 28" refId="B68" refString="Nefediev, PS, 2019. New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia. Far Eastern Entomologist = Dal'nevostochnyi Entomolog 380: 23 - 28, DOI: https://doi.org/10.25221/fee.380.4" title="New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia." url="https://doi.org/10.25221/fee.380.4" volume="380" year="2019">Nefediev 2019</bibRefCitation>
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: 27;
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<bibRefCitation id="959576ED94A759C3DCA9FADF0A054CFD" DOI="https://doi.org/10.37828/em.2022.60.11" author="Dyachkov, YuV" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="17" pagination="70 - 79" refId="B35" refString="Dyachkov, YuV, 2022. On new records of Geophilomorpha (Chilopoda) from Middle Asia. Ecologica Montenegrina 60: 70 - 79, DOI: https://doi.org/10.37828/em.2022.60.11" title="On new records of Geophilomorpha (Chilopoda) from Middle Asia." url="https://doi.org/10.37828/em.2022.60.11" volume="60" year="2022">Dyachkov 2022</bibRefCitation>
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: 25.
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</paragraph>
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</subSubSection>
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<subSubSection id="F7CCCF7A49C0D470FBEF2F5EB676C81E" pageId="0" pageNumber="17" type="type locality">
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<paragraph id="2ED1AF840AC4D3A78CA3B1F6CF463995" pageId="0" pageNumber="17">Type locality.</paragraph>
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<paragraph id="87CD0E6B9D1A20E5ACCF910AA5D8E68A" pageId="0" pageNumber="17">
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Kyrgyzstan: Issyk-Kul Region:
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<normalizedToken id="3E213C6ABEEC1DD5859BF97A71AD22C4" originalValue="“Przewalsk”">"Przewalsk"</normalizedToken>
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(
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<bibRefCitation id="810FB7F2C38CA63FC338FA47FB3A6E3E" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems 1904</bibRefCitation>
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) = Karakol.
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</paragraph>
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</subSubSection>
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<subSubSection id="E1DB19B82F56BC58388F28439F8460FB" pageId="0" pageNumber="17" type="type series">
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<paragraph id="4FE64932745061C3D0D0EC91C2332231" pageId="0" pageNumber="17">Type series.</paragraph>
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<paragraph id="2357A1B7443BE95E4A5B4E253D0CC855" pageId="0" pageNumber="17">
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<emphasis id="AD95EC92FA5E29494F9829F8FC79AD52" bold="true" italics="true" pageId="0" pageNumber="17">Syntypes</emphasis>
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: 4 specimens, including 2 males and 2 females. Deposited in NHMW (
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<bibRefCitation id="14515DCE9735D4DAC610AE7B61C98C13" author="Ilie, V" journalOrPublisher="Verlag des Naturhistorischen Museum in Vienna" pageId="0" pageNumber="17" pagination="1 - 75" refId="B55" refString="Ilie, V, Schiller, E, Stagl, V, 2009. Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien. Verlag des Naturhistorischen Museum in Vienna 22 (4): 1 - 75" title="Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien." volume="22" year="2009">Ilie et al. 2009</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="E7BB9B4C8FB7E12B05EAADD1DF409DE4" pageId="0" pageNumber="17" type="diagnosis">
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<paragraph id="A480E107DCC8041149CC718CE2295816" pageId="0" pageNumber="17">Diagnosis.</paragraph>
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<paragraph id="1BE703D83ED8D8876A01962E9BCD150E" pageId="0" pageNumber="17">
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An
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<taxonomicName id="03AA6CFE0FAB2D18A5F22E1FE184561D" authorityName="Cook & Collins" authorityYear="1891" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="ACEA4DA176DC4A2F976BA5D9190E6E8F" italics="true" pageId="0" pageNumber="17">Escaryus</emphasis>
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</taxonomicName>
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species with body length reaching ≥ 4 cm; clypeus with small plagulae; labral arc relatively deep, with denticles long and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur, and tibia with small denticles, tarsungulum with a small bulge; 49-55 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="39D2E1561ACCF250CA6D385DA95E2C49" pageId="0" pageNumber="17" type="distribution">
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<paragraph id="E8559C6344456DD8723AF55A0494706D" pageId="0" pageNumber="17">Distribution.</paragraph>
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<paragraph id="85E5A162481043C90725EB869EA42E2C" pageId="0" pageNumber="17">
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Western Siberia: Altai krai, Republic of Altai, and Kemerovo oblast (
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<bibRefCitation id="19E82800BF6D1A86E061A3AF4A3E767B" DOI="https://doi.org/10.15298/arthsel.26.1.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="8 - 14" refId="B69" refString="Nefediev, PS, Tuf, IH, Farzalieva, GS, 2017a. Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha. Arthropoda Selecta 26 (1): 8 - 14, DOI: https://doi.org/10.15298/arthsel.26.1.02" title="Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha." url="https://doi.org/10.15298/arthsel.26.1.02" volume="26" year="2017 a">Nefediev et al. 2017a</bibRefCitation>
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,
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<bibRefCitation id="F68BF3682C00298F826288BB029AB131" DOI="https://doi.org/10.15298/arthsel.26.3.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="217 - 224" refId="B71" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, 2017c. A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha). Arthropoda Selecta 26 (3): 217 - 224, DOI: https://doi.org/10.15298/arthsel.26.3.02" title="A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha)." url="https://doi.org/10.15298/arthsel.26.3.02" volume="26" year="2017 c">2017c</bibRefCitation>
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,
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<bibRefCitation id="A6EA87E30CFB97370EB452FB5716197E" DOI="https://doi.org/10.3897/zookeys.741.21936" author="Nefediev, PS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="17" pagination="219 - 254" refId="B72" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, Nedoev, KK, Niyazov, ST, 2018. Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda). ZooKeys 741: 219 - 254, DOI: https://doi.org/10.3897/zookeys.741.21936" title="Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda)." url="https://doi.org/10.3897/zookeys.741.21936" volume="741" year="2018">2018</bibRefCitation>
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;
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<bibRefCitation id="A466E4B484F466706C3C15E11503C351" DOI="https://doi.org/10.25221/fee.380.4" author="Nefediev, PS" journalOrPublisher="Far Eastern Entomologist = Dal'nevostochnyi Entomolog" pageId="0" pageNumber="17" pagination="23 - 28" refId="B68" refString="Nefediev, PS, 2019. New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia. Far Eastern Entomologist = Dal'nevostochnyi Entomolog 380: 23 - 28, DOI: https://doi.org/10.25221/fee.380.4" title="New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia." url="https://doi.org/10.25221/fee.380.4" volume="380" year="2019">Nefediev 2019</bibRefCitation>
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). Possibly Far East (
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<bibRefCitation id="7B52180C2DBF0251E8938225267F2064" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
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,
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<bibRefCitation id="22661D4164B2B950A2976C87DE73440E" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
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; see Remarks). Outside Asian Russia: westwards to Moldova (
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<bibRefCitation id="8BA397AA6F1506674577AFC31313352D" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Titova 1973</bibRefCitation>
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;
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<bibRefCitation id="E08726B953B35A807AC30E71797775B3" DOI="https://doi.org/10.1134/S0013873816040138" author="Volkova, YuS" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="17" pagination="669 - 678" refId="B113" refString="Volkova, YuS, 2016. An annotated catalogue of geophilomorph centipedes (Chilopoda, Geophilomorpha) from the European part of Russia. Zoologicheskii Zhurnal 95 (6): 669 - 678, DOI: https://doi.org/10.1134/S0013873816040138" title="An annotated catalogue of geophilomorph centipedes (Chilopoda, Geophilomorpha) from the European part of Russia." url="https://doi.org/10.1134/S0013873816040138" volume="95" year="2016">Volkova 2016</bibRefCitation>
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;
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<bibRefCitation id="6ECE423E2BCF34F9014DCFD0AE1F87AE" DOI="https://doi.org/10.15298/arthsel.25.1.03" author="Zuev, RV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="23 - 38" refId="B117" refString="Zuev, RV, 2016. Centipedes (Chilopoda) from the Stavropol Territory, northern Caucasus, Russia. Arthropoda Selecta 25 (1): 23 - 38, DOI: https://doi.org/10.15298/arthsel.25.1.03" title="Centipedes (Chilopoda) from the Stavropol Territory, northern Caucasus, Russia." url="https://doi.org/10.15298/arthsel.25.1.03" volume="25" year="2016">Zuev 2016</bibRefCitation>
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;
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<bibRefCitation id="060FC7EEFAECBE1785DE70C537959FDB" DOI="https://doi.org/10.15298/rusentj.25.4.12" author="Zuev, RV" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="17" pagination="417 - 426" refId="B118" refString="Zuev, RV, Evsyukov, AP, 2016. Centipedes (Chilopoda) from the Rostov-on-Don Region, southern Russia. Russian Entomological Journal 25 (4): 417 - 426, DOI: https://doi.org/10.15298/rusentj.25.4.12" title="Centipedes (Chilopoda) from the Rostov-on-Don Region, southern Russia." url="https://doi.org/10.15298/rusentj.25.4.12" volume="25" year="2016">Zuev and Evsyukov 2016</bibRefCitation>
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); southwards to Kazakhstan and Kyrgyzstan (
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<bibRefCitation id="6A00A597D22CC2AD3212E88D9963E614" DOI="https://doi.org/10.5962/bhl.part.18578" author="Attems, C" journalOrPublisher="Abteilung fur Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="17" pagination="113 - 130" refId="B4" refString="Attems, C, 1904. Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher. Abteilung fur Systematik, Oekologie und Geographie der Tiere 20: 113 - 130, DOI: https://doi.org/10.5962/bhl.part.18578" title="Central- und hoch-asiatische Myriopoden. Gesammelt im Jahre 1900 von Dr. von Almassy und Dr. von Stummer. Zoologische Jahrbucher." url="https://doi.org/10.5962/bhl.part.18578" volume="20" year="1904">Attems 1904</bibRefCitation>
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;
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<bibRefCitation id="883155A02135E45964E1AEDF3F1C9C34" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
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,
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<bibRefCitation id="536B35B29BE50E162F9AE789A0314430" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
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;
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<bibRefCitation id="823319B8093FC1110166D5C47EC76692" DOI="https://doi.org/10.15298/arthsel.27.4.04" author="Dyachkov, YuV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="293 - 299" refId="B40" refString="Dyachkov, YuV, Tuf, IH, 2018. New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan. Arthropoda Selecta 26 (4): 293 - 299, DOI: https://doi.org/10.15298/arthsel.27.4.04" title="New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan." url="https://doi.org/10.15298/arthsel.27.4.04" volume="26" year="2018">Dyachkov and Tuf 2018</bibRefCitation>
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;
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<bibRefCitation id="F0FADF87A55499F1628FFFC38CD6EA88" DOI="https://doi.org/10.37828/em.2022.60.11" author="Dyachkov, YuV" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="17" pagination="70 - 79" refId="B35" refString="Dyachkov, YuV, 2022. On new records of Geophilomorpha (Chilopoda) from Middle Asia. Ecologica Montenegrina 60: 70 - 79, DOI: https://doi.org/10.37828/em.2022.60.11" title="On new records of Geophilomorpha (Chilopoda) from Middle Asia." url="https://doi.org/10.37828/em.2022.60.11" volume="60" year="2022">Dyachkov 2022</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="13FF31E27D238300C020393F1436093C" pageId="0" pageNumber="17" type="remarks">
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<paragraph id="B8021BAE50A363F70FA421FD7767C627" pageId="0" pageNumber="17">Remarks.</paragraph>
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<paragraph id="4C2B8685147DC4468209839D823F7E83" pageId="0" pageNumber="17">
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A total of 19 males and 26 females collected from Trans-Ili Alatau (Almaty Region of Kazakhstan) were indicated by
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<bibRefCitation id="12E341F4221BC12693FC60248ACC9232" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Titova (1973</bibRefCitation>
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: 110) as lectotypes, but this action is not valid, as the syntypes still exist (
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<bibRefCitation id="E271BB288224ECD0893E711054076096" author="Ilie, V" journalOrPublisher="Verlag des Naturhistorischen Museum in Vienna" pageId="0" pageNumber="17" pagination="1 - 75" refId="B55" refString="Ilie, V, Schiller, E, Stagl, V, 2009. Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien. Verlag des Naturhistorischen Museum in Vienna 22 (4): 1 - 75" title="Type specimens of the Geophilomorpha (Chilopoda) in the Natural History Museum in Vienna. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museum in Wien." volume="22" year="2009">Ilie et al. 2009</bibRefCitation>
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).
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</paragraph>
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<paragraph id="919C1EC8827F2895C7C14CED7D33D448" pageId="0" pageNumber="17">
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<bibRefCitation id="C7A43938F6704FCD969767B70A0080A4" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova (1972</bibRefCitation>
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: 135;
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<bibRefCitation id="AA570B98147D95C57BF7A88538CDE49B" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
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: 116) wrote "... it is possible to distinguish territories from Cisamuria to Kuzbass and Altai, where 4 species are spread:
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<taxonomicName id="EEE10B068A3A6B82487AEB7271DF7AD5" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">
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<emphasis id="D00FB62CE316CD27D4EF5D436DA9BE39" italics="true" pageId="0" pageNumber="17">E. koreanus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="A87FA70060623CE15996BF3E49A2EBAE" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. japonicus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="2C94CB981D4A431FADAA3334BFE92E1C" italics="true" pageId="0" pageNumber="17">E. japonicus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8E94460561C2F152F4221ADC7EC2CF62" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">
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<emphasis id="8D1BBFEB4FBBA13B9E78EAB66A1F750A" italics="true" pageId="0" pageNumber="17">E. retusidens</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="93191338D8E346E569817CF578A98741" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. chadaevae" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="chadaevae">
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<emphasis id="FB46445B3748EE1C6AB3FB0E920DAA75" italics="true" pageId="0" pageNumber="17">E. chadaevae</emphasis>
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</taxonomicName>
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". However, she did not mention material from this area when she listed the studied material of
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<taxonomicName id="714C95B6CC85EB0FD76FF49287FEE06F" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">
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<emphasis id="016BE0B5CFF2A63172DCC0F99EB22344" italics="true" pageId="0" pageNumber="17">E. retusidens</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="13BB76411B123E6B465D0DABE8292DA6" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Titova 1973</bibRefCitation>
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: 110): "Kazakhstan, Trans-Ili Alatau
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<normalizedToken id="672DAE734CBFB65F03DD43D3C13D5EB3" originalValue="…">...</normalizedToken>
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Moreover,
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<taxonomicName id="7160F361ABE14E97FE9B484853E56D76" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. retusidens" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="retusidens">
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<emphasis id="521AEDB69316E19B690ECE5CF7C4B7F5" italics="true" pageId="0" pageNumber="17">E. retusidens</emphasis>
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</taxonomicName>
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were studied from the Dzhungarian Alatau, the Greater Caucasus, Crimea, Moldova, Rostov, Voronezh and Voroshilovograd oblasts".
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |