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<document id="6844791DE9A6782BF4417A8287262263" ID-DOI="10.5281/zenodo.7630433" ID-GBIF-Dataset="180b8c6d-242a-4fff-a1a3-e88d2e1e51ed" ID-ISBN="978-2-8277-0088-2" ID-ISSN="0373-2975" ID-Zenodo-Dep="7630433" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1637929385876" checkinUser="carolina" docAuthor="AL-Shehbaz, Ihsan A. &amp; Barriera, Gabrielle" docDate="2019" docId="AC0E341DFFFA830AFF4C5AB6FAD6FD36" docLanguage="en" docName="Boissiera.72.1-192.pdf.imf" docOrigin="Boissiera 72" docStyle="DocumentStyle:724D25757667A47A191C06980913D161.2:Boissiera.2019.journal_volume" docStyleId="724D25757667A47A191C06980913D161" docStyleName="Boissiera.2019.journal_volume" docStyleVersion="2" docTitle="Hesperis leucoclada Boiss." docType="treatment" docVersion="10" lastPageNumber="71" masterDocId="50374C65FFBD834DFFC45860FF90FFFB" masterDocTitle="Typification of Edmond Boissiers Cruciferae (Brassicaceae) names enumerated in Flora Orientalis" masterLastPageNumber="193" masterPageNumber="1" pageNumber="71" updateTime="1728913889019" updateUser="juliana">
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<mods:title id="4D0BD1D31F3375203D5CC3586A7BA552">Typification of Edmond Boissiers Cruciferae (Brassicaceae) names enumerated in Flora Orientalis</mods:title>
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<mods:namePart id="838853182DB4F78E44ECDD203812729A">AL-Shehbaz, Ihsan A.</mods:namePart>
<mods:affiliation id="C43A41A6CED0015CAB82C1D9F06AAB19">Missouri Botanical Garden 4344 Shaw Boulevard St. Louis, Missouri 63110, USA</mods:affiliation>
<mods:nameIdentifier id="9D1924FDAABF4EA08A0D2EB0370D1D08" type="email">ihsan.al-shehbaz@mobot.org</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="FED5A08E9E35511E6348397B83033D89">Barriera, Gabrielle</mods:namePart>
<mods:affiliation id="3BED5988753E4FCA6AB248B98711AEBD">Conservatoire et Jardin botaniques de la Ville de Genève Case postale 71 1292 Chambésy, Switzerland</mods:affiliation>
<mods:nameIdentifier id="D6AE2B165E1F9D666A5CA54464E193A2" type="email">gabrielle.barriera@ville-ge.ch</mods:nameIdentifier>
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<mods:titleInfo id="7301B0BB4D7512288586B80179C63033">
<mods:title id="C90CC921222FC71395A5830C17038267">Boissiera</mods:title>
</mods:titleInfo>
<mods:part id="AD54EB3617438A745DC4803F66231740">
<mods:date id="01EE153128C33EA3299581B80124C318">2019</mods:date>
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<mods:number id="DE3C316AB87F157664F892221A640C14">2019-12-23</mods:number>
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<mods:number id="3CD34B34684598EBED6868A809DAFABB">72</mods:number>
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<mods:identifier id="6EB73FD33D7441D36BE0D1DE8C6F2997" type="DOI">10.5281/zenodo.7630433</mods:identifier>
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<mods:identifier id="4D2B1C935D4F4E8A52381B229E0DFFFA" type="ISBN">978-2-8277-0088-2</mods:identifier>
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<treatment id="AC0E341DFFFA830AFF4C5AB6FAD6FD36" ID-DOI="http://doi.org/10.5281/zenodo.7637050" ID-GBIF-Taxon="209296007" ID-Zenodo-Dep="7637050" LSID="urn:lsid:plazi:treatment:AC0E341DFFFA830AFF4C5AB6FAD6FD36" httpUri="http://treatment.plazi.org/id/AC0E341DFFFA830AFF4C5AB6FAD6FD36" lastPageNumber="71" pageId="71" pageNumber="71">
<subSubSection id="6CBDD680FFFA830AFF4C5AB6FF50FCF6" pageId="71" pageNumber="71" type="nomenclature">
<paragraph id="2418850BFFFA830AFF4C5AB6FF50FCF6" blockId="71.[136,776,725,781]" pageId="71" pageNumber="71">
<treatmentCitation id="A506A31AFFFA830AFF4C5AB6FF2CFCF6" author="Boiss." journal="Ann. Sci. Nat., Bot." page="69" pageId="71" pageNumber="71" score="32.7652104432798" volume="17" year="1842">
<taxonomicName id="E3A7FE88FFFA830AFF4C5AB6FE1BFD16" ID-CoL="88ZKD" authority="Boiss." authorityName="Boiss." box="[136,395,725,749]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFF4C5AB6FED6FD16" box="[136,326,725,749]" italics="true" pageId="71" pageNumber="71">Hesperis leucoclada</emphasis>
Boiss.
</taxonomicName>
in Ann. Sci. Nat., Bot. ser. 2, 17: 69. 1842
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6CBDD680FFFA830AFF675B49FDF3FCA6" pageId="71" pageNumber="71" type="reference_group">
<paragraph id="2418850BFFFA830AFF675B49FDF3FCA6" blockId="71.[163,723,805,861]" pageId="71" pageNumber="71">
<treatmentCitationGroup id="04B7A225FFFA830AFF675B49FDF3FCA6" pageId="71" pageNumber="71">
<treatmentCitation id="A506A31AFFFA830AFF795B45FDCFFCA6" author="D. A. German &amp; Al-Shehbaz" httpUri="http://treatment.plazi.org/id/0671720F8730FFA5FF6AFBB6FE83DE5E" journal="Phytotaxa" page="96" pageId="71" pageNumber="71" score="9.729594737742866" volume="334" year="2018">
<taxonomicName id="E3A7FE88FFFA830AFF795B45FEAEFCA6" ID-CoL="8GMWY" authority="(Boiss.) D. A. German &amp; Al-Shehbaz" authorityName="D. A. German &amp; Al-Shehbaz" baseAuthorityName="Boiss." class="Magnoliopsida" family="Brassicaceae" genus="Sisymbrium" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucocladum">
<emphasis id="16D35919FFFA830AFF795B45FE52FCC6" bold="true" box="[189,450,805,829]" italics="true" pageId="71" pageNumber="71">Sisymbrium leucocladum</emphasis>
(Boiss.) D.A. German &amp; Al-Shehbaz
</taxonomicName>
in
<bibCitation id="F18D8D67FFFA830AFEA55B25FDCFFCA6" box="[353,607,837,861]" journal="Phytotaxa" page="96" pageId="71" pageNumber="71" score="9.729594737742866" volume="334" year="2018">Phytotaxa 334: 96. 2018</bibCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6CBDD680FFFA830AFF675B16FECFFBA7" pageId="71" pageNumber="71" type="materials_examined">
<paragraph id="2418850BFFFA830AFF675B16FDC3FC76" blockId="71.[163,595,885,910]" box="[163,595,885,910]" pageId="71" pageNumber="71">
Type: “[Aucher-Eloy]
<collectionCode id="42B61DCEFFFA830AFE575B16FE3AFC76" box="[403,426,886,909]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="71" pageNumber="71" type="Herbarium">N</collectionCode>
. 4123,
<collectingRegion id="E6634BE9FFFA830AFE2B5B15FDD5FC76" box="[495,581,885,909]" country="Iran" name="Esfahan" pageId="71" pageNumber="71">Ispahan</collectingRegion>
”.
</paragraph>
<paragraph id="2418850BFFFA830AFF675BC5FECFFBA7" blockId="71.[163,786,932,1116]" pageId="71" pageNumber="71">
<materialsCitation id="94CF8F56FFFA830AFF675BC5FD0AFC26" ID-GBIF-Occurrence="4030734626" collectionCode="G-BOIS" collectorName="Aucher-Eloy" country="Iran" location="Ispahan" pageId="71" pageNumber="71" specimenCode="G00332220" specimenCount="1" stateProvince="Ispahan" typeStatus="lectotype">
<emphasis id="16D35919FFFA830AFF675BC5FEB8FC46" bold="true" box="[163,296,933,957]" pageId="71" pageNumber="71">
<typeStatus id="FB1C3BA9FFFA830AFF675BC5FEB8FC46" box="[163,296,933,957]" pageId="71" pageNumber="71" type="lectotype">Lectotypus</typeStatus>
</emphasis>
(designated here):
<emphasis id="16D35919FFFA830AFDD25BC4FDC7FC46" bold="true" box="[534,599,932,957]" pageId="71" pageNumber="71">
<collectingCountry id="5CB0C59BFFFA830AFDD25BC4FDDFFC47" box="[534,591,932,956]" name="Iran" pageId="71" pageNumber="71">IRAN</collectingCountry>
:
</emphasis>
<collectingRegion id="E6634BE9FFFA830AFDAA5BC4FD59FC47" box="[622,713,932,956]" country="Iran" name="Esfahan" pageId="71" pageNumber="71">Ispahan</collectingRegion>
”, s.d.,
<emphasis id="16D35919FFFA830AFF675BA4FEFCFC26" box="[163,364,964,989]" italics="true" pageId="71" pageNumber="71">
<collectorName id="8952E0DDFFFA830AFF675BA4FEBAFC27" box="[163,298,964,988]" pageId="71" pageNumber="71">Aucher-Eloy</collectorName>
4123
</emphasis>
(
<collectionCode id="42B61DCEFFFA830AFE475BA5FE7AFC26" box="[387,490,965,989]" pageId="71" pageNumber="71">G-BOIS</collectionCode>
[
<specimenCode id="74012D70FFFA830AFE3B5BA5FD1FFC26" box="[511,655,965,989]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="71" pageNumber="71" type="Herbarium">G00332220</specimenCode>
];
</materialsCitation>
<materialsCitation id="94CF8F56FFFA830AFD6C5BA4FECBFBA7" ID-GBIF-Occurrence="4030734394" collectionCode="BM, G, K, KW, LE, P, W" country="Iran" location="Aucher-Eloy" pageId="71" pageNumber="71" specimenCode="BM000522181, BM000522182, G00446088, G00446089, K000693872, KW000127972, LE00013085, P02272565, P02272566, P02272567, W0050812" specimenCount="1" stateProvince="Esfahan" typeStatus="isolectotype">
isolecto-:
<collectionCode id="42B61DCEFFFA830AFF675B85FF40FC07" box="[163,208,997,1020]" country="United Kingdom" name="Bristol Museum" pageId="71" pageNumber="71">BM</collectionCode>
[
<specimenCode id="74012D70FFFA830AFF255B84FE1AFC07" box="[225,394,996,1020]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="71" pageNumber="71">BM000522181</specimenCode>
,
<specimenCode id="74012D70FFFA830AFE5E5B84FDD6FC07" box="[410,582,996,1020]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="71" pageNumber="71">BM000522182</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFD9F5B85FDE1FC06" box="[603,625,997,1021]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="71" pageNumber="71" type="Herbarium">G</collectionCode>
[
<specimenCode id="74012D70FFFA830AFD475B84FC9DFC07" box="[643,781,996,1020]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="71" pageNumber="71" type="Herbarium">G00446088</specimenCode>
,
<specimenCode id="74012D70FFFA830AFF675C64FEBEFBE7" box="[163,302,1028,1052]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="71" pageNumber="71" type="Herbarium">G00446089</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFE805C65FEC7FBE7" box="[324,343,1029,1052]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="71" pageNumber="71" type="Herbarium">K</collectionCode>
[
<specimenCode id="74012D70FFFA830AFEAD5C64FD90FBE7" box="[361,512,1028,1052]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="71" pageNumber="71" type="Herbarium">K000693872</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFDD25C65FDD6FBE7" box="[534,582,1029,1052]" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="71" pageNumber="71" type="Herbarium">KW</collectionCode>
[
<specimenCode id="74012D70FFFA830AFD935C65FC98FBE6" box="[599,776,1029,1053]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="71" pageNumber="71" type="Herbarium">KW000127972</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFF675C45FF56FBC7" box="[163,198,1061,1084]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="71" pageNumber="71">LE</collectionCode>
[
<specimenCode id="74012D70FFFA830AFF165C44FEF4FBC7" box="[210,356,1060,1084]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="71" pageNumber="71">LE00013085</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFEB05C45FE14FBC7" box="[372,388,1061,1084]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="71" pageNumber="71" type="Herbarium">P</collectionCode>
[
<specimenCode id="74012D70FFFA830AFE545C44FD97FBC7" box="[400,519,1060,1084]" pageId="71" pageNumber="71">P02272565</specimenCode>
,
<specimenCode id="74012D70FFFA830AFDD65C44FD17FBC7" box="[530,647,1060,1084]" pageId="71" pageNumber="71">P02272566</specimenCode>
,
<specimenCode id="74012D70FFFA830AFD555C44FC98FBC7" box="[657,776,1060,1084]" pageId="71" pageNumber="71">P02272567</specimenCode>
],
<collectionCode id="42B61DCEFFFA830AFF675C25FF2FFBA7" box="[163,191,1093,1116]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="71" pageNumber="71" type="Herbarium">W</collectionCode>
[
<specimenCode id="74012D70FFFA830AFF085C24FEDDFBA7" box="[204,333,1092,1116]" collectionCode="W" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="71" pageNumber="71" type="Herbarium">W0050812</specimenCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6CBDD680FFFA830AFF675CE5FAD6FD36" pageId="71" pageNumber="71" type="discussion">
<paragraph id="2418850BFFFA830AFF675CE5FD7DFAC7" blockId="71.[135,788,1156,1948]" pageId="71" pageNumber="71">
<emphasis id="16D35919FFFA830AFF675CE5FF72FB67" box="[163,226,1157,1180]" italics="true" pageId="71" pageNumber="71">Notes.</emphasis>
In addition to the specimen in G-BOIS,
<bibRefCitation id="4036F8FAFFFA830AFD745CE5FF4CFB46" author="BOISSIER, E." pageId="71" pageNumber="71" pagination="45 - 90" refId="ref136077" refString="BOISSIER, E. (1842 a). Plantae Aucherianae orientales enumeratae, cum novarum specierum descriptione. Ann. Sci. Nat., Bot. ser. 2, 17: 45 - 90." type="journal article" year="1842">BOISSIER (1842a)</bibRefCitation>
based the species description on the duplicate P
<date id="5019A3CBFFFA830AFF525CA4FF6BFB27" box="[150,251,1220,1244]" pageId="71" pageNumber="71" value="2565-02-27">02272565</date>
that he annotated. Therefore, the lectotypification of
<bibRefCitation id="4036F8FAFFFA830AFF675CBFFED6FB07" author="DVORAK, F." box="[163,326,1247,1277]" pageId="71" pageNumber="71" pagination="105 - 112" refId="ref138006" refString="DVORAK, F. (1968 a). Problems of the taxon Hesperis leucoclada Boiss. Feddes Repert. 77: 105 - 112." type="journal article" year="1968">DVOŘÁK (1968a</bibRefCitation>
,
<bibRefCitation id="4036F8FAFFFA830AFE8B5C84FE1FFB07" author="DVORAK, F." box="[335,399,1252,1276]" pageId="71" pageNumber="71" pagination="266 - 274" refId="ref138032" refString="DVORAK, F. (1968 b). Hesperis. In: RECHINGER, K. H. (ed.), Fl. Iranica 57: 266 - 274." type="journal article" year="1968">1968b</bibRefCitation>
), which was based on W0050812 that Boissier never examined and which was recently maintained by
<bibRefCitation id="4036F8FAFFFA830AFF6C5D45FD9DFAC6" author="AL-SHEHBAZ, I. A. &amp; D. A. GERMAN" box="[168,525,1317,1341]" pageId="71" pageNumber="71" pagination="212" refId="ref135166" refString="AL-SHEHBAZ, I. A. &amp; D. A. GERMAN (2018). (2585) Proposal to conserve the name Hesperis pendula DC. against H. pendula Murr. (Cruciferae). Taxon 67: 212." type="journal article" year="2018">GERMAN &amp; AL-SHEHBAZ (2018)</bibRefCitation>
, cannot be accepted.
</paragraph>
<paragraph id="2418850BFFFA830AFF675D24FD40F947" blockId="71.[135,788,1156,1948]" pageId="71" pageNumber="71">
The generic placement of
<taxonomicName id="E3A7FE88FFFA830AFE7C5D25FDE6FAA7" box="[440,630,1348,1372]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFE7C5D25FDE6FAA7" box="[440,630,1348,1372]" italics="true" pageId="71" pageNumber="71">Hesperis leucoclada</emphasis>
</taxonomicName>
has been controversial, and
<bibRefCitation id="4036F8FAFFFA830AFEE65D3FFE42FA87" author="DVORAK, F." box="[290,466,1375,1404]" pageId="71" pageNumber="71" pagination="105 - 112" refId="ref138006" refString="DVORAK, F. (1968 a). Problems of the taxon Hesperis leucoclada Boiss. Feddes Repert. 77: 105 - 112." type="journal article" year="1968">DVOŘÁK (1968a)</bibRefCitation>
compared it more with members of the tribe
<taxonomicName id="E3A7FE88FFFA830AFEF95DE5FE32FA67" box="[317,418,1413,1436]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="tribe" tribe="Brassiceae">
<emphasis id="16D35919FFFA830AFEF95DE5FE32FA67" box="[317,418,1413,1436]" italics="true" pageId="71" pageNumber="71">Brassiceae</emphasis>
</taxonomicName>
largely because he misinterpreted the incumbent cotyledons with slightly depressed inner one as somewhat conduplicate and eventually placed the species in monospecific
<taxonomicName id="E3A7FE88FFFA830AFE805D84FE6FFA07" authority="Gilli" authorityName="Gilli" authorityYear="1955" box="[324,511,1508,1532]" class="Magnoliopsida" family="Brassicaceae" genus="Gynophorea" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFE805D84FE52FA07" box="[324,450,1508,1532]" italics="true" pageId="71" pageNumber="71">Gynophorea</emphasis>
Gilli
</taxonomicName>
almost solely on having stipitate vs sessile fruit. About a week after the above publication,
<bibRefCitation id="4036F8FAFFFA830AFF215E7FFE09F9C7" author="DVORAK, F." box="[229,409,1567,1596]" pageId="71" pageNumber="71" pagination="266 - 274" refId="ref138032" refString="DVORAK, F. (1968 b). Hesperis. In: RECHINGER, K. H. (ed.), Fl. Iranica 57: 266 - 274." type="journal article" year="1968">DVOŘÁK (1968b)</bibRefCitation>
retained the species in
<emphasis id="16D35919FFFA830AFD5D5E45FD66F9C7" box="[665,758,1573,1596]" italics="true" pageId="71" pageNumber="71">
<taxonomicName id="E3A7FE88FFFA830AFD5D5E45FD61F9C7" authorityName="C.Linnaeus" authorityYear="1753" box="[665,753,1573,1596]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">Hesperis</taxonomicName>
,
</emphasis>
as was originally placed by
<bibRefCitation id="4036F8FAFFFA830AFE4E5E25FDD1F9A6" author="BOISSIER, E." box="[394,577,1605,1629]" pageId="71" pageNumber="71" pagination="45 - 90" refId="ref136077" refString="BOISSIER, E. (1842 a). Plantae Aucherianae orientales enumeratae, cum novarum specierum descriptione. Ann. Sci. Nat., Bot. ser. 2, 17: 45 - 90." type="journal article" year="1842">BOISSIER (1842a)</bibRefCitation>
. The species is quite anomalous in
<taxonomicName id="E3A7FE88FFFA830AFEE25E05FEEEF987" authorityName="C.Linnaeus" authorityYear="1753" box="[294,382,1637,1660]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFEE25E05FEEEF987" box="[294,382,1637,1660]" italics="true" pageId="71" pageNumber="71">Hesperis</emphasis>
</taxonomicName>
because it does not have the decurrent stigmas, branched trichomes, and uniseriate multicellular glands characteristic of almost all species of the genus.
</paragraph>
<paragraph id="2418850BFFFA830AFF675EA4FA8DFDF6" blockId="71.[135,788,1156,1948]" lastBlockId="71.[824,1476,277,717]" pageId="71" pageNumber="71">
Another more realistic placement of
<taxonomicName id="E3A7FE88FFFA830AFDEF5EA5FD3FF927" box="[555,687,1732,1756]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFDEF5EA5FD3FF927" box="[555,687,1732,1756]" italics="true" pageId="71" pageNumber="71">H. leucoclada</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation id="4036F8FAFFFA830AFF295E85FE39F906" author="KHOSRAVI, A. R." box="[237,425,1765,1789]" pageId="71" pageNumber="71" pagination="15 - 23" refId="ref140323" refString="KHOSRAVI, A. R. (2003). A reconsideration of Hesperis leucoclada Boiss. (Cruciferae). Iran. J. Bot. 10: 15 - 23." type="journal article" year="2003">KHOSRAVI (2003)</bibRefCitation>
based on molecular studies and a closer look at morphology. He transferred the species to the Iranian-endemic
<taxonomicName id="E3A7FE88FFFA830AFE825F44FDADF8C7" authority="Hedge" authorityName="Hedge" box="[326,573,1828,1852]" class="Magnoliopsida" family="Brassicaceae" genus="Pseudofortuynia" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFE825F44FE7CF8C7" box="[326,492,1828,1852]" italics="true" pageId="71" pageNumber="71">Pseudofortuynia</emphasis>
Hedge
</taxonomicName>
, but neither he nor
<bibRefCitation id="4036F8FAFFFA830AFF4D5F25FEB2F8A7" author="HEDGE, I. C." box="[137,290,1860,1885]" pageId="71" pageNumber="71" pagination="33 - 342" refId="ref139580" refString="HEDGE, I. C. (1968). Brassiceae, Lepidieae, Drabeae, Arabideae, Sisymbrieae. In: RECHINGER, K. H. (ed.), Fl. Iranica 57: 33 - 342." type="journal article" year="1968">HEDGE (1968)</bibRefCitation>
realized that they were dealing with the same species that they assigned to the tribe
<taxonomicName id="E3A7FE88FFFA830AFDE95F05FD03F887" box="[557,659,1893,1916]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="tribe" tribe="Brassiceae">
<emphasis id="16D35919FFFA830AFDE95F05FD03F887" box="[557,659,1893,1916]" italics="true" pageId="71" pageNumber="71">Brassiceae</emphasis>
</taxonomicName>
. Except for their purple instead of yellow flowers and stipitate vs sessile fruits, a critical closer examination of the
<typeStatus id="FB1C3BA9FFFA830AFB3B5976FABDFED5" box="[1279,1325,278,302]" pageId="71" pageNumber="71">type</typeStatus>
collections at G and W of
<taxonomicName id="E3A7FE88FFFA830AFC075956FB11FEB6" box="[963,1153,309,333]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFC075956FB11FEB6" box="[963,1153,309,333]" italics="true" pageId="71" pageNumber="71">Hesperis leucoclada</emphasis>
</taxonomicName>
and
<taxonomicName id="E3A7FE88FFFA830AFB715955FCEFFE96" authority="Hedge" authorityName="Hedge" class="Magnoliopsida" family="Brassicaceae" genus="Pseudofortuynia" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="esfandiari">
<emphasis id="16D35919FFFA830AFB715955FA52FEB6" box="[1205,1474,309,333]" italics="true" pageId="71" pageNumber="71">Pseudofortuynia esfandiari</emphasis>
Hedge
</taxonomicName>
reveals that, they are indistinguishable from
<taxonomicName id="E3A7FE88FFFA830AFA8F5935FA52FE96" box="[1355,1474,341,365]" class="Magnoliopsida" family="Brassicaceae" genus="Sisymbrium" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFA8F5935FA52FE96" box="[1355,1474,341,365]" italics="true" pageId="71" pageNumber="71">Sisymbrium</emphasis>
</taxonomicName>
in every morphological character and therefore should be unit-ed with it. It is interesting to note that
<bibRefCitation id="4036F8FAFFFA830AFB2859F5FCEBFE36" author="BOISSIER, E. &amp; F. BUHSE" pageId="71" pageNumber="71" pagination="13 - 31" refId="ref136409" refString="BOISSIER, E. &amp; F. BUHSE (1860). Aufzaehlung der auf einer Reise durch Transkaukasien und Persien gesammelten Pflanzen. Nouv. Mem. Soc. Imp. Naturalistes Moscou 12: 13 - 31." type="journal article" year="1860">BOISSIER &amp; BUHSE (1860)</bibRefCitation>
described
<taxonomicName id="E3A7FE88FFFA830AFC2B59D6FB34FE36" authorityName="Boiss. &amp; Buhse" authorityYear="1860" box="[1007,1188,437,461]" class="Magnoliopsida" family="Brassicaceae" genus="Sisymbrium" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="hesperidiflorum">
<emphasis id="16D35919FFFA830AFC2B59D6FB34FE36" box="[1007,1188,437,461]" italics="true" pageId="71" pageNumber="71">S. hesperidiflorum</emphasis>
</taxonomicName>
(see below) and compared it with
<taxonomicName id="E3A7FE88FFFA830AFC4059B6FBFAFE15" authority="Bunge" authorityName="Bunge" box="[900,1130,469,494]" class="Magnoliopsida" family="Brassicaceae" genus="Sisymbrium" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="subspinescens">
<emphasis id="16D35919FFFA830AFC4059B6FB8EFE16" box="[900,1054,469,493]" italics="true" pageId="71" pageNumber="71">S. subspinescens</emphasis>
Bunge
</taxonomicName>
, then Boissier in
<emphasis id="16D35919FFFA830AFAE459B5FA52FE16" box="[1312,1474,469,493]" italics="true" pageId="71" pageNumber="71">Flora Orientalis</emphasis>
reduced it to synonymy of
<taxonomicName id="E3A7FE88FFFA830AFB9C5996FA86FDF6" box="[1112,1302,501,525]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFB9C5996FA86FDF6" box="[1112,1302,501,525]" italics="true" pageId="71" pageNumber="71">Hesperis leucoclada</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2418850BFFFA830AFC975A76FAD6FD36" blockId="71.[824,1476,277,717]" pageId="71" pageNumber="71">
In conclusion,
<taxonomicName id="E3A7FE88FFFA830AFC325A76FBEFFDD6" box="[1014,1151,533,557]" class="Magnoliopsida" family="Brassicaceae" genus="Hesperis" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="species" species="leucoclada">
<emphasis id="16D35919FFFA830AFC325A76FBEFFDD6" box="[1014,1151,533,557]" italics="true" pageId="71" pageNumber="71">H. leucoclada</emphasis>
</taxonomicName>
was recognized as a member of the tribe
<taxonomicName id="E3A7FE88FFFA830AFC7A5A55FB6EFDB6" authority="(DVORAK, 1968 b)" baseAuthorityName="DVORAK" baseAuthorityYear="1968" box="[958,1278,560,589]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="tribe" tribe="Hesperideae">
<emphasis id="16D35919FFFA830AFC7A5A55FBA9FDB6" box="[958,1081,565,589]" italics="true" pageId="71" pageNumber="71">Hesperideae</emphasis>
(
<bibRefCitation id="4036F8FAFFFA830AFB8D5A50FB66FDB6" author="DVORAK, F." box="[1097,1270,560,589]" pageId="71" pageNumber="71" pagination="266 - 274" refId="ref138032" refString="DVORAK, F. (1968 b). Hesperis. In: RECHINGER, K. H. (ed.), Fl. Iranica 57: 266 - 274." type="journal article" year="1968">DVOŘÁK, 1968b</bibRefCitation>
)
</taxonomicName>
, a member of the tribe
<taxonomicName id="E3A7FE88FFFA830AFCB15A36FC4DFD96" box="[885,989,598,621]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="tribe" tribe="Brassiceae">
<emphasis id="16D35919FFFA830AFCB15A36FC4DFD96" box="[885,989,598,621]" italics="true" pageId="71" pageNumber="71">Brassiceae</emphasis>
</taxonomicName>
(
<bibRefCitation id="4036F8FAFFFA830AFC2B5A30FB0CFD96" author="DVORAK, F." box="[1007,1180,592,621]" pageId="71" pageNumber="71" pagination="105 - 112" refId="ref138006" refString="DVORAK, F. (1968 a). Problems of the taxon Hesperis leucoclada Boiss. Feddes Repert. 77: 105 - 112." type="journal article" year="1968">DVOŘÁK, 1968a</bibRefCitation>
;
<bibRefCitation id="4036F8FAFFFA830AFB6D5A35FAAEFD96" author="HEDGE, I. C." box="[1193,1342,597,621]" pageId="71" pageNumber="71" pagination="33 - 342" refId="ref139580" refString="HEDGE, I. C. (1968). Brassiceae, Lepidieae, Drabeae, Arabideae, Sisymbrieae. In: RECHINGER, K. H. (ed.), Fl. Iranica 57: 33 - 342." type="journal article" year="1968">HEDGE, 1968</bibRefCitation>
;
<bibRefCitation id="4036F8FAFFFA830AFA8F5A35FCFFFD76" author="KHOSRAVI, A. R." pageId="71" pageNumber="71" pagination="15 - 23" refId="ref140323" refString="KHOSRAVI, A. R. (2003). A reconsideration of Hesperis leucoclada Boiss. (Cruciferae). Iran. J. Bot. 10: 15 - 23." type="journal article" year="2003">KHOSRAVI, 2003</bibRefCitation>
), and
<taxonomicName id="E3A7FE88FFFA830AFC745A15FBB8FD76" box="[944,1064,629,653]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="tribe" tribe="Sisymbrieae">
<emphasis id="16D35919FFFA830AFC745A15FBB8FD76" box="[944,1064,629,653]" italics="true" pageId="71" pageNumber="71">Sisymbrieae</emphasis>
</taxonomicName>
(
<bibRefCitation id="4036F8FAFFFA830AFBFC5A15FA00FD76" author="AL-SHEHBAZ, I. A. &amp; D. A. GERMAN" box="[1080,1424,629,653]" pageId="71" pageNumber="71" pagination="212" refId="ref135166" refString="AL-SHEHBAZ, I. A. &amp; D. A. GERMAN (2018). (2585) Proposal to conserve the name Hesperis pendula DC. against H. pendula Murr. (Cruciferae). Taxon 67: 212." type="journal article" year="2018">GERMAN &amp; AL-SHEHBAZ, 2018</bibRefCitation>
, and herein). Therefore, the recent synonymy by the last authors of
<taxonomicName id="E3A7FE88FFFA830AFCFC5AD5FC4BFD36" box="[824,987,693,717]" class="Magnoliopsida" family="Brassicaceae" genus="Pseudofortuynia" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFCFC5AD5FC4BFD36" box="[824,987,693,717]" italics="true" pageId="71" pageNumber="71">Pseudofortuynia</emphasis>
</taxonomicName>
with
<taxonomicName id="E3A7FE88FFFA830AFBDE5AD5FB03FD36" box="[1050,1171,693,717]" class="Magnoliopsida" family="Brassicaceae" genus="Sisymbrium" kingdom="Plantae" order="Brassicales" pageId="71" pageNumber="71" phylum="Tracheophyta" rank="genus">
<emphasis id="16D35919FFFA830AFBDE5AD5FB03FD36" box="[1050,1171,693,717]" italics="true" pageId="71" pageNumber="71">Sisymbrium</emphasis>
</taxonomicName>
is fully justified.
</paragraph>
</subSubSection>
</treatment>
</document>