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<document id="79D81815804E6B8CCC6AFF85FB68F064" ID-CLB-Dataset="7681" ID-DOI="10.11646/zootaxa.4942.1.1" ID-GBIF-Dataset="114135c4-23cb-444b-8cff-75d9f43aad6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4596025" ID-ZooBank="CDF398FE-B0F6-40E6-967D-FB857C1565BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615447261711" checkinUser="plazi" docAuthor="Yasunaga, Tomohide, Duwal, Ram Keshari &amp; Nakatani, Yukinobu" docDate="2021" docId="A16ADB37EB030C79FF47C56279DEFE05" docLanguage="en" docName="zootaxa.4942.1.1.pdf" docOrigin="Zootaxa 4942 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pilophorus choii Josifov 1978" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="5D53A34FEB050C7EFFD0C03C7D09FF8C" masterDocTitle="Reclassification of the plant bug genus Pilophorus in Japan and key to the genera and species of Japanese Pilophorini (Hemiptera: Heteroptera: Miridae Phylinae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="7" updateTime="1698933027681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2382C40C96143C6ADCEFA5347EFA1798" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="766724EC51C1FE58C1BD2AB5C889A7F8">
<mods:title id="123F1373589955D0BC4A0B0F64EE8008">Reclassification of the plant bug genus Pilophorus in Japan and key to the genera and species of Japanese Pilophorini (Hemiptera: Heteroptera: Miridae Phylinae)</mods:title>
</mods:titleInfo>
<mods:name id="AB3EBDF27A085B56629CB05B1945B726" type="personal">
<mods:role id="13EBA6EBC8FA6DCBDFC078C60D9DECC9">
<mods:roleTerm id="23BA369D5A289A305D32C4D8F68631B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="978438C3F46AC8FA38D77A6A02296A60">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="E026C607C97E1147B67CB1DF6AF6FDF3">Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, Central Park at 79 th Street, New York, NY 10024, c / o Nameshi 2 - 33 - 2, Nagasaki 852 - 8061, Japan.</mods:affiliation>
</mods:name>
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<mods:namePart id="B4988F0BA5DE1EDFB26519977E409DB9">Duwal, Ram Keshari</mods:namePart>
<mods:affiliation id="C7167FCF298EF17A66AA8F7F938DF7C6">Visiting Researcher, Agriculture &amp; Agri-Food Canada Environmental Health, K. W. Neatby: Bldg # 20, 960 Carling Avenue, Central Experimental Farm, Ottawa Ontario, Canada K 1 A 0 C 6.</mods:affiliation>
</mods:name>
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<mods:role id="39B258AA26EC44493AC01ABE0E7F91C9">
<mods:roleTerm id="A19DF6099EE2ACF4B48FB238A9E60CBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2D17850106789953971E319ADF2972B2">Nakatani, Yukinobu</mods:namePart>
<mods:affiliation id="835EFCA9B20EDDBBEEB9982255FE0ABF">Division of Informatics and Inventory, Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba, Ibaraki 305 - 8604, Japan.</mods:affiliation>
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<mods:typeOfResource id="913CC85AD78D310BC38388038A5DC421">text</mods:typeOfResource>
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<mods:title id="2A3D69705B073CF04DA85B09FD0F4F44">Zootaxa</mods:title>
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<mods:part id="A60A4F452213C8ADE576BB431AE813B3">
<mods:date id="40E2594C07317ADB75EFE73906A57775">2021</mods:date>
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<mods:number id="F8560E4942608CE5E85014E73614EA5F">2021-03-11</mods:number>
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<mods:number id="8D2A5E1D742E510ADF73D4DC823F6299">4942</mods:number>
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<treatment id="A16ADB37EB030C79FF47C56279DEFE05" ID-DOI="http://doi.org/10.5281/zenodo.4596031" ID-GBIF-Taxon="180588583" ID-Zenodo-Dep="4596031" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05" httpUri="http://treatment.plazi.org/id/A16ADB37EB030C79FF47C56279DEFE05" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="61D939AAEB030C78FF47C5627F01FAF4" box="[151,520,1374,1401]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="297C6A21EB030C78FF47C5627F01FAF4" blockId="6.[151,520,1374,1437]" box="[151,520,1374,1401]" pageId="6" pageNumber="7">
<heading id="7234DD4DEB030C78FF47C5627F01FAF4" bold="true" box="[151,520,1374,1401]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="EEC311A2EB030C78FF47C5627F01FAF4" ID-CoL="77K5M" authority="Josifov, 1978" authorityName="Josifov" authorityYear="1978" box="[151,520,1374,1401]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB030C78FF47C5627F01FAF4" bold="true" box="[151,520,1374,1401]" pageId="6" pageNumber="7">
<emphasis id="1BB7B633EB030C78FF47C5627C69FAF5" bold="true" box="[151,352,1374,1401]" italics="true" pageId="6" pageNumber="7">Pilophorus choii</emphasis>
<bibRefCitation id="4D5217D0EB030C78FEB8C5627F01FAF4" author="Josifov, M." box="[360,520,1374,1400]" pageId="6" pageNumber="7" pagination="279 - 287" refId="ref24518" refString="Josifov, M. (1978) Neue Miridenarten aus Nord-Korea (Heteroptera). Acta Entomologica Musei Nationalis Pragae, 39 (1977), 279 - 287." type="journal article" year="1978">Josifov, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="61D939AAEB030C78FF47C5BF7CA8FA11" box="[151,417,1411,1437]" pageId="6" pageNumber="7" type="description">
<paragraph id="297C6A21EB030C78FF47C5BF7CA8FA11" blockId="6.[151,520,1374,1437]" box="[151,417,1411,1437]" pageId="6" pageNumber="7">
(
<figureCitation id="B1F876A4EB030C78FF4FC5BF7DF5FA11" box="[159,252,1411,1437]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="6" pageNumber="7">Figs 1A</figureCitation>
,
<figureCitation id="B1F876A4EB030C78FED8C5BF7C30FA11" box="[264,313,1411,1437]" captionStart="FIGURE 11" captionStartId="23.[151,250,1788,1812]" captionTargetBox="[173,1401,286,1750]" captionTargetId="figure-61@23.[164,1423,273,1765]" captionTargetPageId="23" captionText="FIGURE 11. Male endosoma (vesica) and endosoma of Japanese Pilophorus spp. A. P. choii (from Otaru). B. P. erraticus (Nagasaki). C. P. pseudoperplexus (Kanagawa). D. P.okamotoi (Tsushima Is.). E. P.satoyamanus n. sp. (Kochi).F. P.setulosellus n. sp. (Hiroshima). G. Same (Nara). H. P. setulosus (Sapporo, type locality, Salix). I. Same, (Shiretoko, Artemisia)." figureDoi="http://doi.org/10.5281/zenodo.4596059" httpUri="https://zenodo.org/record/4596059/files/figure.png" pageId="6" pageNumber="7">11A</figureCitation>
,
<figureCitation id="B1F876A4EB030C78FE95C5BF7C56FA11" box="[325,351,1411,1437]" captionStart="FIGURE 14" captionStartId="28.[151,250,1888,1912]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-27@28.[151,1436,181,1864]" captionTargetPageId="28" captionText="FIGURE 14. Scanning electron micrographs for Japanese Pilophorus spp., males (except for E, L, P: ♀). A–D. P. choii. E–K. P. erraticus. L–N. P. pseudoperplexus. O–Q. P. miyamotoi. A, E. Left-dorsolateral habitus. B, I, N. Apex of pygophore, left lateral view. C–D, J–K, Q. Endosoma (vesica).A, E. Dorsal habitus. F. Thoracic pleura, left lateral view. H, L, M, O, P. Lateral habitus. G. Pygophore, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4596067" httpUri="https://zenodo.org/record/4596067/files/figure.png" pageId="6" pageNumber="7">14</figureCitation>
A–D)
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB030C78FF47C5FB7ED5FA72" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="297C6A21EB030C78FF47C5FB7ED5FA72" blockId="6.[151,1436,1479,1534]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="09D34D0FEB030C78FF47C5FB7ED5FA72" pageId="6" pageNumber="7">
<treatmentCitation id="A8624C30EB030C78FF47C5FB7F0FFA53" author="Josifov, M." box="[151,518,1479,1503]" page="285" pageId="6" pageNumber="7" year="1978">
<taxonomicName id="EEC311A2EB030C78FF47C5FB7F0FFA53" ID-CoL="77K5M" authority="Josifov, 1978: 285" authorityName="Josifov" authorityPageNumber="285" authorityYear="1978" box="[151,518,1479,1503]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB030C78FF47C5FB7C4BFA53" box="[151,322,1479,1503]" italics="true" pageId="6" pageNumber="7">Pilophorus choii</emphasis>
<bibRefCitation id="4D5217D0EB030C78FE99C5FB7F0FFA53" author="Josifov, M." box="[329,518,1479,1503]" pageId="6" pageNumber="7" pagination="279 - 287" refId="ref24518" refString="Josifov, M. (1978) Neue Miridenarten aus Nord-Korea (Heteroptera). Acta Entomologica Musei Nationalis Pragae, 39 (1977), 279 - 287." type="journal article" year="1978">Josifov, 1978: 285</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(n. sp., desc.);
<bibRefCitation id="4D5217D0EB030C78FD75C5FB7E2EFA53" author="Schuh, R. T." box="[677,807,1479,1503]" pageId="6" pageNumber="7" refId="ref25329" refString="Schuh, R. T. (2002 - 2013) On-line Systematic Catalog of Plant Bugs (Insecta: Heteroptera: Miridae). Available from: http: // research. amnh. org / pbi / catalog / (accessed 5 December 2020)" type="url" year="2002">Schuh, 2002</bibRefCitation>
–2013, online catalog;
<treatmentCitation id="A8624C30EB030C78FBC1C5FB780AFA53" author="Duwal, R. K. &amp; Jung, S. H. &amp; Lee, S. H." box="[1041,1283,1479,1503]" page="261" pageId="6" pageNumber="7" year="2014">
<bibRefCitation id="4D5217D0EB030C78FBC1C5FB780AFA53" author="Duwal, R. K. &amp; Jung, S. H. &amp; Lee, S. H." box="[1041,1283,1479,1503]" pageId="6" pageNumber="7" pagination="257 - 271" refId="ref24159" refString="Duwal, R. K., Jung, S. H. &amp; Lee, S. H. (2014) A taxonomic review of the plant bug tribe Pilophorini (Hemiptera: Miridae: Phylinae) from the Korean Peninsula. Journal of Asia Pacific Entomology, 17, 257 - 271. https: // doi. org / 10.1016 / j. aspen. 2014.01.003" type="journal article" year="2014">Duwal et al., 2014: 261</bibRefCitation>
</treatmentCitation>
(redesc., key);
<bibRefCitation id="4D5217D0EB030C78FF17C5DA7C56FA72" author="Aukema, B." box="[199,351,1510,1534]" pageId="6" pageNumber="7" refId="ref24010" refString="Aukema, B. (2018) Catalogue of the Palaearctic Heteroptera (searchable database). Available from: https: // catpalhet. linnaeus. naturalis. nl / (accessed 5 December 2020)" type="url" year="2018">Aukema, 2018</bibRefCitation>
, online catalog;
<treatmentCitation id="A8624C30EB030C78FDD4C5DA7E50FA72" author="Yamamoto, A. &amp; Yasunaga, T." box="[516,857,1510,1534]" page="75" pageId="6" pageNumber="7" year="2020">
<bibRefCitation id="4D5217D0EB030C78FDD4C5DA7E50FA72" author="Yamamoto, A. &amp; Yasunaga, T." box="[516,857,1510,1534]" pageId="6" pageNumber="7" pagination="75 - 83" refId="ref25574" refString="Yamamoto, A. &amp; Yasunaga, T. (2020) New distributional records of a microphysid and 10 mirid species from Hokkaido, Japan. Rostria, 65, 75 - 83. [in Japanese with English summary]" type="journal article" year="2020">Yamamoto &amp; Yasunaga, 2020: 75</bibRefCitation>
</treatmentCitation>
(faunal list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB030C78FF47C6107FFFF806" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="297C6A21EB030C78FF47C6107F85F95A" blockId="6.[151,1437,1580,2038]" pageId="6" pageNumber="7">
<emphasis id="1BB7B633EB030C78FF47C6107C88F9CA" bold="true" box="[151,385,1580,1606]" pageId="6" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="99AB607CEB030C78FE58C6107FBDF9E6" ID-GBIF-Occurrence="3053984335" collectingDate="2007-08-18" collectionCode="OMO" collectorName="A. Yamamoto" country="Japan" latitude="43.165" location="Estuary of Shinkawa River" longLatPrecision="68" longitude="141.227" municipality="Otaru City" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Hokkaido">
<emphasis id="1BB7B633EB030C78FE58C6107CEBF9CA" bold="true" box="[392,482,1580,1606]" pageId="6" pageNumber="7">
<collectingCountry id="51D42AB1EB030C78FE58C6107CEBF9CA" box="[392,482,1580,1606]" name="Japan" pageId="6" pageNumber="7">JAPAN</collectingCountry>
</emphasis>
:
<collectingRegion id="EB07A4C3EB030C78FE21C6107F6DF9CA" box="[497,612,1580,1606]" country="Japan" name="Hokkaido" pageId="6" pageNumber="7">Hokkaido</collectingRegion>
:
<collectingMunicipality id="C918F05BEB030C78FDBFC6107FEEF9CA" box="[623,743,1580,1606]" pageId="6" pageNumber="7">Otaru City</collectingMunicipality>
,
<location id="2C1C3CFAEB030C78FD23C611792CF9CA" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05:2C1C3CFAEB030C78FD23C611792CF9CA" box="[755,1061,1580,1607]" country="Japan" latitude="43.165" longLatPrecision="68" longitude="141.227" municipality="Otaru City" name="Estuary of Shinkawa River" pageId="6" pageNumber="7" stateProvince="Hokkaido">Estuary of Shinkawa River</location>
,
<geoCoordinate id="4CF70CE6EB030C78FBE1C610799DF9CB" box="[1073,1172,1580,1607]" degrees="43.165" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="55" value="43.165">N43.165</geoCoordinate>
,
<geoCoordinate id="4CF70CE6EB030C78FB70C6107806F9CA" box="[1184,1295,1580,1606]" degrees="141.227" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="141.227">E141.227</geoCoordinate>
,
<collectedFrom id="9ADF29DCEB030C78FACBC6117DE2F9E6" pageId="6" pageNumber="7">
on
<taxonomicName id="EEC311A2EB030C78FAEEC6107DE2F9E6" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="dentata">
<emphasis id="1BB7B633EB030C78FAEEC6107DE2F9E6" italics="true" pageId="6" pageNumber="7">Quercus dentata</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="5D7D4CE1EB030C78FF27C66C7C8EF9E6" box="[247,391,1616,1642]" pageId="6" pageNumber="7" value="2007-08-18">
<collectingDate id="4D39B509EB030C78FF27C66C7C8EF9E6" box="[247,391,1616,1642]" pageId="6" pageNumber="7" value="2007-08-18">18 Aug 2007</collectingDate>
</date>
,
<collectorName id="84360FF7EB030C78FE5FC66C7F22F9E7" box="[399,555,1616,1643]" pageId="6" pageNumber="7">A.Yamamoto</collectorName>
,
<specimenCount id="3FC5A1A8EB030C78FDE4C66C7F5EF9E6" box="[564,599,1616,1642]" pageId="6" pageNumber="7" type="female">1♀</specimenCount>
(
<collectionCode id="4FD2F2E4EB030C78FDB5C66C7FA5F9E6" box="[613,684,1616,1642]" pageId="6" pageNumber="7">OMO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="99AB607CEB030C78FD6DC66C7F43F902" ID-GBIF-Occurrence="3053984318" collectingDate="2009-08-06" collectionCode="OMO" collectorName="A. Yamamoto" country="Japan" latitude="43.186" location="Otaru City" longLatPrecision="68" longitude="141.269" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Hokkaido">
<location id="2C1C3CFAEB030C78FD6DC66C7E3AF9E6" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05:2C1C3CFAEB030C78FD6DC66C7E3AF9E6" box="[701,819,1616,1642]" country="Japan" latitude="43.186" longLatPrecision="68" longitude="141.269" name="Otaru City" pageId="6" pageNumber="7" stateProvince="Hokkaido">Otaru City</location>
, Tarukawa-9-sen,
<geoCoordinate id="4CF70CE6EB030C78FC2DC66C7944F9E6" box="[1021,1101,1616,1642]" degrees="43.186" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="55" value="43.186">43.186</geoCoordinate>
,
<geoCoordinate id="4CF70CE6EB030C78FB86C66C79BDF9E6" box="[1110,1204,1616,1642]" degrees="141.269" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="141.269">141.269</geoCoordinate>
,
<collectedFrom id="9ADF29DCEB030C78FB6DC66D789CF9E6" box="[1213,1429,1616,1643]" pageId="6" pageNumber="7">
on
<taxonomicName id="EEC311A2EB030C78FB0EC66C789CF9E6" box="[1246,1429,1616,1642]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="dentata">
<emphasis id="1BB7B633EB030C78FB0EC66C789CF9E6" box="[1246,1429,1616,1642]" italics="true" pageId="6" pageNumber="7">Quercus dentata</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="5D7D4CE1EB030C78FF47C6487C12F902" box="[151,283,1652,1678]" pageId="6" pageNumber="7" value="2009-08-06">
<collectingDate id="4D39B509EB030C78FF47C6487C12F902" box="[151,283,1652,1678]" pageId="6" pageNumber="7" value="2009-08-06">6 Aug 2009</collectingDate>
</date>
,
<collectorName id="84360FF7EB030C78FEF3C6487CC9F903" box="[291,448,1652,1679]" pageId="6" pageNumber="7">A. Yamamoto</collectorName>
, 1♂ (
<collectionCode id="4FD2F2E4EB030C78FE2BC6487F4BF902" box="[507,578,1652,1678]" pageId="6" pageNumber="7">OMO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="99AB607CEB030C78FD84C6487FB9F902" ID-GBIF-Occurrence="3053984343" box="[596,688,1652,1678]" country="Japan" location="Kyushu" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Hokkaido">
<location id="2C1C3CFAEB030C78FD84C6487FB9F902" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05:2C1C3CFAEB030C78FD84C6487FB9F902" box="[596,688,1652,1678]" country="Japan" name="Kyushu" pageId="6" pageNumber="7" stateProvince="Hokkaido">Kyushu</location>
</materialsCitation>
:
<materialsCitation id="99AB607CEB030C78FD69C6487F41F93F" ID-GBIF-Occurrence="3053984338" collectingDate="1995-07-27" collectionCode="NIAES, AMNH" collectorName="Y. Nakatani" country="Japan" location="Handa Plateau" pageId="6" pageNumber="7" specimenCode="PBI 00380663" specimenCount="1" stateProvince="Oita">
<collectingRegion id="EB07A4C3EB030C78FD69C6487E21F902" box="[697,808,1652,1678]" country="Japan" name="Oita" pageId="6" pageNumber="7">Oita Pref.</collectingRegion>
,
<location id="2C1C3CFAEB030C78FCE2C6487EDFF902" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05:2C1C3CFAEB030C78FCE2C6487EDFF902" box="[818,982,1652,1678]" country="Japan" name="Handa Plateau" pageId="6" pageNumber="7" stateProvince="Oita">Handa Plateau</location>
, 33.13, 131.22,
<date id="5D7D4CE1EB030C78FB56C6487803F903" box="[1158,1290,1652,1679]" pageId="6" pageNumber="7" value="1995-07-27">
<collectingDate id="4D39B509EB030C78FB56C6487803F903" box="[1158,1290,1652,1679]" pageId="6" pageNumber="7" value="1995-07-27">27 Jul 1995</collectingDate>
</date>
,
<collectorName id="84360FF7EB030C78FAC2C6497891F902" box="[1298,1432,1652,1679]" pageId="6" pageNumber="7">Y. Nakatani</collectorName>
, 1♂ (
<collectionCode id="4FD2F2E4EB030C78FF18C6A47C17F93E" box="[200,286,1688,1714]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="6" pageNumber="7">NIAES</collectionCode>
) (
<collectionCode id="4FD2F2E4EB030C78FEE3C6A47C83F93E" box="[307,394,1688,1714]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="7">AMNH</collectionCode>
_
<specimenCode id="7965C25AEB030C78FE49C6A47F49F93F" box="[409,576,1688,1715]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="6" pageNumber="7">PBI 00380663</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="99AB607CEB030C78FD81C6A47F81F95A" ID-GBIF-Occurrence="3053984314" collectingDate="2009-07-16" collectionCode="SNU" collectorName="R. K. Duwal &amp; S. Jung" country="South Korea" location="Mt. Youngmun" municipality="Yangpyeonggun" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Gyeonggi">
<emphasis id="1BB7B633EB030C78FD81C6A47FB1F93E" bold="true" box="[593,696,1688,1714]" pageId="6" pageNumber="7">
<collectingCountry id="51D42AB1EB030C78FD81C6A47FB1F93E" box="[593,696,1688,1714]" name="South Korea" pageId="6" pageNumber="7">KOREA</collectingCountry>
</emphasis>
:
<collectingRegion id="EB07A4C3EB030C78FD16C6A47E54F93E" box="[710,861,1688,1714]" country="South Korea" name="Gyeonggi" pageId="6" pageNumber="7">Gyeonggi-do</collectingRegion>
,
<collectingMunicipality id="C918F05BEB030C78FCB6C6A57916F93F" box="[870,1055,1689,1715]" pageId="6" pageNumber="7">Yangpyeonggun</collectingMunicipality>
,
<location id="2C1C3CFAEB030C78FBFAC6A579DCF93F" LSID="urn:lsid:plazi:treatment:A16ADB37EB030C79FF47C56279DEFE05:2C1C3CFAEB030C78FBFAC6A579DCF93F" box="[1066,1237,1689,1715]" country="South Korea" municipality="Yangpyeonggun" name="Mt. Youngmun" pageId="6" pageNumber="7" stateProvince="Gyeonggi">Mt. Youngmun</location>
,
<collectingMethod id="F0821236EB030C78FB0FC6A5787CF93F" box="[1247,1397,1688,1715]" pageId="6" pageNumber="7">UV light trap</collectingMethod>
,
<date id="5D7D4CE1EB030C78FA50C6A47DF3F95A" pageId="6" pageNumber="7" value="2009-07-16">
<collectingDate id="4D39B509EB030C78FA50C6A47DF3F95A" pageId="6" pageNumber="7" value="2009-07-16">16 Jul 2009</collectingDate>
</date>
,
<collectorName id="84360FF7EB030C78FED5C6817C85F95A" box="[261,396,1724,1751]" pageId="6" pageNumber="7">R.K. Duwal</collectorName>
&amp;
<collectorName id="84360FF7EB030C78FE7FC6807F0CF95B" box="[431,517,1724,1751]" pageId="6" pageNumber="7">S. Jung</collectorName>
, 1♂ (
<collectionCode id="4FD2F2E4EB030C78FD92C6807F89F95A" box="[578,640,1724,1750]" pageId="6" pageNumber="7">SNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="297C6A21EB030C78FF17C6DC7FFFF806" blockId="6.[151,1437,1580,2038]" pageId="6" pageNumber="7">
<emphasis id="1BB7B633EB030C78FF17C6DC7C57F976" bold="true" box="[199,350,1760,1786]" pageId="6" pageNumber="7">Rediagnosis.</emphasis>
Recognized by its moderate size; short antennal segment that is shorter than basal width of pronotum(see
<tableCitation id="64415F9AEB030C78FEEBC7387C98F892" box="[315,401,1796,1822]" captionStart="TABLE 1" captionStartId="8.[151,239,152,176]" captionText="TABLE 1. Measurements for Pilophorus hyotan n. sp. and P. typicus." pageId="6" pageNumber="7">Table 1</tableCitation>
for measurements of male); polished, shiny pronotum; partly interrupted, narrow median stripe of scale-like setae that are easily rubbed off (
<figureCitation id="B1F876A4EB030C78FD33C7147E37F8CE" box="[739,830,1832,1858]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
); narrowly clustered scale-like setae at anterior inner corner of cuneus; and C-shaped, small-sized male endosoma (
<figureCitation id="B1F876A4EB030C78FC98C7707EA6F8EA" box="[840,943,1868,1894]" captionStart="FIGURE 11" captionStartId="23.[151,250,1788,1812]" captionTargetBox="[173,1401,286,1750]" captionTargetId="figure-61@23.[164,1423,273,1765]" captionTargetPageId="23" captionText="FIGURE 11. Male endosoma (vesica) and endosoma of Japanese Pilophorus spp. A. P. choii (from Otaru). B. P. erraticus (Nagasaki). C. P. pseudoperplexus (Kanagawa). D. P.okamotoi (Tsushima Is.). E. P.satoyamanus n. sp. (Kochi).F. P.setulosellus n. sp. (Hiroshima). G. Same (Nara). H. P. setulosus (Sapporo, type locality, Salix). I. Same, (Shiretoko, Artemisia)." figureDoi="http://doi.org/10.5281/zenodo.4596059" httpUri="https://zenodo.org/record/4596059/files/figure.png" pageId="6" pageNumber="7">Fig. 11A</figureCitation>
). Dorsal habitus images available on https:// www.discoverlife.org/20/q?search=
<taxonomicName id="EEC311A2EB030C78FDFDC74C7FA3F806" authorityName="Hahn" authorityYear="1826" box="[557,682,1904,1930]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Pilophorus</taxonomicName>
+choii.
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB030C78FF17C7A87C98F87A" pageId="6" pageNumber="7" type="biology_ecology">
<paragraph id="297C6A21EB030C78FF17C7A87C98F87A" blockId="6.[151,1437,1580,2038]" pageId="6" pageNumber="7">
<emphasis id="1BB7B633EB030C78FF17C7A87C2EF822" bold="true" box="[199,295,1940,1966]" pageId="6" pageNumber="7">Biology.</emphasis>
The Japanese population were found to inhabit
<taxonomicName id="EEC311A2EB030C78FC94C7A87EF4F822" box="[836,1021,1940,1966]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="dentata">
<emphasis id="1BB7B633EB030C78FC94C7A87EF4F822" box="[836,1021,1940,1966]" italics="true" pageId="6" pageNumber="7">Quercus dentata</emphasis>
</taxonomicName>
, as reported for
<collectingCountry id="51D42AB1EB030C78FB67C7A979F2F823" box="[1207,1275,1941,1967]" name="South Korea" pageId="6" pageNumber="7">Korea</collectingCountry>
(
<bibRefCitation id="4D5217D0EB030C78FADBC7A87DD8F85E" author="Duwal, R. K. &amp; Jung, S. H. &amp; Lee, S. H." pageId="6" pageNumber="7" pagination="257 - 271" refId="ref24159" refString="Duwal, R. K., Jung, S. H. &amp; Lee, S. H. (2014) A taxonomic review of the plant bug tribe Pilophorini (Hemiptera: Miridae: Phylinae) from the Korean Peninsula. Journal of Asia Pacific Entomology, 17, 257 - 271. https: // doi. org / 10.1016 / j. aspen. 2014.01.003" type="journal article" year="2014">Duwal et al., 2014</bibRefCitation>
) and Russian Primorsky Territory (
<bibRefCitation id="4D5217D0EB030C78FD86C7847E01F85E" author="Kerzhner, I. M." box="[598,776,1976,2002]" pageId="6" pageNumber="7" pagination="21" refId="ref24575" refString="Kerzhner, I. M. (1988) Infraorder Cimicomorpha. 21. Family Miridae (Capsidae). In: Ler, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR [Keys to the identification of insects of the Soviet Far East]. Vol. 2. Homoptera and Heteroptera. Nauka, Leningrad, pp. 778 - 857. [in Russian]" type="book chapter" year="1988">Kerzhner, 1988</bibRefCitation>
). A univoltine life cycle is assumed for
<taxonomicName id="EEC311A2EB030C78FB6AC7857804F85E" authorityName="Josifov" authorityYear="1978" box="[1210,1293,1976,2002]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB030C78FB6AC7857804F85E" box="[1210,1293,1976,2002]" italics="true" pageId="6" pageNumber="7">P. choii</emphasis>
</taxonomicName>
and the eggs presumably hibernate.
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB020C79FF17C0AA79DEFE05" pageId="7" pageNumber="8" type="discussion">
<paragraph id="297C6A21EB020C79FF17C0AA7932FF75" blockId="7.[151,1437,150,393]" pageId="7" pageNumber="8">
<emphasis id="1BB7B633EB020C79FF17C0AA7C45FF3C" bold="true" box="[199,332,150,176]" pageId="7" pageNumber="8">Discussion.</emphasis>
This species is most similar in external appearance to
<taxonomicName id="EEC311A2EB020C79FC30C0AB7958FF3D" authorityName="Linnavuori" authorityYear="1962" box="[992,1105,151,177]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lucidus">
<emphasis id="1BB7B633EB020C79FC30C0AB7958FF3D" box="[992,1105,151,177]" italics="true" pageId="7" pageNumber="8">P. lucidus</emphasis>
</taxonomicName>
as both are associated with deciduous
<taxonomicName id="EEC311A2EB020C79FEDEC0877C65FF59" box="[270,364,187,213]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="1BB7B633EB020C79FEDEC0877C65FF59" box="[270,364,187,213]" italics="true" pageId="7" pageNumber="8">Quercus</emphasis>
</taxonomicName>
trees and have the similar size, short antennal segment II and shiny polished pronotum; however, the characters provided in the above key (couplet 25) will readily distinguish them.
</paragraph>
<paragraph id="297C6A21EB020C79FF17C13F79DEFE05" blockId="7.[151,1437,150,393]" pageId="7" pageNumber="8">
Since only a few specimens of
<taxonomicName id="EEC311A2EB020C79FDFBC13F7F89FE91" authorityName="Josifov" authorityYear="1978" box="[555,640,259,285]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB020C79FDFBC13F7F89FE91" box="[555,640,259,285]" italics="true" pageId="7" pageNumber="8">P. choii</emphasis>
</taxonomicName>
are currently available from
<collectingCountry id="51D42AB1EB020C79FC1CC13F7906FE91" box="[972,1039,259,285]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
, its food preference or predation is yet to be tested.
<bibRefCitation id="4D5217D0EB020C79FE9EC11B7F38FECD" author="Fukuda, K. &amp; Kataoka, K. &amp; Ozaki, M. &amp; Sasano, H. &amp; Yasunaga, T. &amp; Asanabe, H." box="[334,561,295,321]" pageId="7" pageNumber="8" pagination="2" refId="ref24229" refString="Fukuda, K., Kataoka, K., Ozaki, M., Sasano, H., Yasunaga, T. &amp; Asanabe, H. (2020) A preliminary assessment of the nymphal characters for ant-mimetic plant bugs of the genus Pilophorus (Hemiptera: Heteroptera: Miridae: Phylinae). Heteropterus Revista de Entomologia, 20 (2). [in press]" type="journal article" year="2020">Fukuda et al. (2020)</bibRefCitation>
and
<bibRefCitation id="4D5217D0EB020C79FDB7C11B7E69FECD" author="Miyazaki, A. &amp; Nishida, M. &amp; Uesugi, R. &amp; Yamada, U. &amp; Yasunaga, T. &amp; Serrano Leon, S. &amp; Kawashita, S. &amp; Nagashima, T." box="[615,864,295,321]" pageId="7" pageNumber="8" pagination="63 - 69" refId="ref24809" refString="Miyazaki, A., Nishida, M., Uesugi, R., Yamada, U., Yasunaga, T., Serrano Leon, S., Kawashita, S. &amp; Nagashima, T. (2020) Utilizing a synthetic diet containing a fermented milk beverage for rearing terrestrial heteropterans, with new distributional records of four species in Nagasaki Prefecture, Japan. Rostria, 64, 63 - 69. [in Japanese with English summary]" type="journal article" year="2020">
<collectingRegion id="EB07A4C3EB020C79FDB7C11B7FD8FECD" box="[615,721,295,321]" country="Japan" name="Miyazaki" pageId="7" pageNumber="8">Miyazaki</collectingRegion>
et al. (2020)
</bibRefCitation>
documented that the immature forms of
<taxonomicName id="EEC311A2EB020C79FAF9C11B789CFECD" authorityName="Linnavuori" authorityYear="1962" box="[1321,1429,295,321]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lucidus">
<emphasis id="1BB7B633EB020C79FAF9C11B789CFECD" box="[1321,1429,295,321]" italics="true" pageId="7" pageNumber="8">P. lucidus</emphasis>
</taxonomicName>
, assumed to be one of the closest relative of
<taxonomicName id="EEC311A2EB020C79FDAAC1777FC5FEE9" authorityName="Josifov" authorityYear="1978" box="[634,716,331,357]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB020C79FDAAC1777FC5FEE9" box="[634,716,331,357]" italics="true" pageId="7" pageNumber="8">P. choii</emphasis>
</taxonomicName>
, could develop into adults when feeding on dried red worms and brine-shrimp eggs as a diet. Therefore,
<taxonomicName id="EEC311A2EB020C79FD9FC1537FABFE05" authorityName="Josifov" authorityYear="1978" box="[591,674,367,393]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="choii">
<emphasis id="1BB7B633EB020C79FD9FC1537FABFE05" box="[591,674,367,393]" italics="true" pageId="7" pageNumber="8">P. choii</emphasis>
</taxonomicName>
is also supposed to have a carnivorous food habit.
</paragraph>
</subSubSection>
</treatment>
</document>
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