236 lines
35 KiB
XML
236 lines
35 KiB
XML
<document id="4E9AA21FBE36190BEE0168A48D92F88B" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFE8FFB1FCB47174D940F9C2" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Tetrabothrus sulawesicus Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="103" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="102" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:title id="5354B0A3D77BBC15E45ED733F8FE0A88">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="CD894C76F024789BEF9902427A1BCD93">Assing, Volker</mods:namePart>
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<mods:title id="AA3E7AB535D2164C8A6E8162D622FE09">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="8EEF97F17CD4D0E4895DF53DA98C1318">
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<mods:date id="E582B4E1003400BF36322F966FA44620">2016</mods:date>
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<mods:number id="58CA98EB9D762D302FAC275E7707670C">2016-12-20</mods:number>
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<mods:number id="4E19A8B5C43A2C5AB39C0148950072DF">66</mods:number>
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<treatment id="766F7C36FFE8FFB1FCB47174D940F9C2" ID-DOI="http://doi.org/10.5281/zenodo.5903520" ID-GBIF-Taxon="192602758" ID-Zenodo-Dep="5903520" LSID="urn:lsid:plazi:treatment:766F7C36FFE8FFB1FCB47174D940F9C2" httpUri="http://treatment.plazi.org/id/766F7C36FFE8FFB1FCB47174D940F9C2" lastPageId="90" lastPageNumber="103" pageId="89" pageNumber="102">
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<subSubSection id="B6DC9EABFFE8FFB2FCB47174DCCCFBED" box="[831,1204,1107,1134]" pageId="89" pageNumber="102" type="nomenclature">
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<paragraph id="FE79CD20FFE8FFB2FCB47174DCCCFBED" blockId="89.[831,1204,1107,1167]" box="[831,1204,1107,1134]" pageId="89" pageNumber="102">
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<heading id="A5317A4CFFE8FFB2FCB47174DCCCFBED" box="[831,1204,1107,1134]" centered="true" fontSize="11" level="2" pageId="89" pageNumber="102" reason="3">
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<taxonomicName id="39C6B6A3FFE8FFB2FCB47174DC3CFBED" ID-CoL="55L2Z" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[831,1092,1107,1134]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="sulawesicus" status="sp. nov.">
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<emphasis id="CCB21132FFE8FFB2FCB47174DC3CFBED" box="[831,1092,1107,1134]" italics="true" pageId="89" pageNumber="102">Tetrabothrus sulawesicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D781AC49FFE8FFB2FBC07174DCCCFBED" box="[1099,1204,1107,1134]" pageId="89" pageNumber="102" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6DC9EABFFE8FFB2FCB4715EDC6BFB0C" box="[831,1043,1145,1167]" pageId="89" pageNumber="102" type="description">
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<paragraph id="FE79CD20FFE8FFB2FCB4715EDC6BFB0C" blockId="89.[831,1204,1107,1167]" box="[831,1043,1145,1167]" pageId="89" pageNumber="102">
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(
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<figureCitation id="66FDD1A5FFE8FFB2FCCF715EDBC2FB0C" box="[836,954,1145,1167]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Figs 167–169</figureCitation>
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,
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<figureCitation id="66FDD1A5FFE8FFB2FC49715EDC76FB0C" box="[962,1038,1145,1167]" captionStart-0="Figs 340–358" captionStart-1="Figs 359–378" captionStart-2="Figs 379–395" captionStart-3="Figs 396–419" captionStart-4="Figs 420–441" captionStart-5="Figs 442–458" captionStartId-0="26.[189,229,1970,1992]" captionStartId-1="27.[151,191,1971,1993]" captionStartId-2="28.[189,229,1943,1965]" captionStartId-3="29.[151,191,2000,2022]" captionStartId-4="30.[189,229,1972,1994]" captionStartId-5="31.[151,191,1971,1993]" captionTargetBox-0="[193,1432,233,1911]" captionTargetBox-1="[155,1396,235,1964]" captionTargetBox-2="[193,1428,232,1921]" captionTargetBox-3="[151,1395,235,1961]" captionTargetBox-4="[189,1427,232,1968]" captionTargetBox-5="[155,1398,240,1950]" captionTargetId-0="figure-134@26.[189,1436,232,1946]" captionTargetId-1="figure-155@27.[151,1398,232,1964]" captionTargetId-2="figure-160@28.[189,1436,232,1934]" captionTargetId-3="figure-113@29.[151,1398,232,1971]" captionTargetId-4="figure-139@30.[189,1436,232,1971]" captionTargetId-5="figure-119@31.[151,1398,232,1971]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionTargetPageId-3="29" captionTargetPageId-4="30" captionText-0="Figs 340–358. Orphnebius vates (340–344), O. tortus (345–347), O. baccillatus (348–350), O. carinatus (351–353), and O. dispar (354–358): median lobe of aedeagus in lateral and in ventral view (340–341, 354–355); paramere (342, 356); tergite VIII (343, 345, 348, 351, 357); male sternite VIII (344); female sternite VIII (346, 349, 352); spermatheca (347, 350, 353, 358). Scale bars: 0.2 mm." captionText-1="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." captionText-2="Figs 379–395. Drusilla bifida (379), Rabdotodrusilla pectinata (380–383), Amaurodera arunica (384–388), A. gilvios (389–390, 395), and A. reticulata (391–394): sutural portion of left elytron (379); median lobe of aedeagus in lateral and in ventral view (380– 381, 385–386, 389–392); male tergite VIII (382); male sternite VIII (383); male pronotum (384); spermatheca (387–388, 395); subapical portion of median lobe of aedeagus in dry preparation (393); subapical portion of median lobe of aedeagus in transparent light (394). Scale bars: 384: 0.5 mm; 379–383, 385–392, 395: 0.2 mm; 393–394: 0.1 mm." captionText-3="Figs 396–419. Amaurodera dentata (396–399), A. parvoculata (400–404), A. thailandensis (405–410), A. fasciata (411–414), and A. meorum (415–419: 415–416, 419:Khao Yai Nat.Park):median lobe of aedeagus in lateral and in ventral view (396–397, 400–402, 405–409, 411–412, 415–418); spermatheca (398–399, 403–404, 410, 413–414, 419). Scale bars: 0.2 mm." captionText-4="Figs 420–441. Amaurodera meorum (420–422), A. disparicollis (423–429), A. spinans (430–431), A. varicollis (432–434), Tetrabothrus nilgiricus (435–436), T. indicus (437–439; 439: lectotype), andT. inflexus fromLaos (440–441): spermatheca (420–422, 429, 434); male pronotum (423–424); female pronotum (425–426); median lobe of aedeagus in lateral and in ventral view (427–428, 430–433, 435–437, 440); ventral process of aedeagus (438–439, 441). Scale bars: 423–426: 1.0 mm; 420–422, 427–441: 0.2 mm." captionText-5="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6421216" figureDoi-1="http://doi.org/10.5281/zenodo.6421218" figureDoi-2="http://doi.org/10.5281/zenodo.6421220" figureDoi-3="http://doi.org/10.5281/zenodo.6421222" figureDoi-4="http://doi.org/10.5281/zenodo.6421224" figureDoi-5="http://doi.org/10.5281/zenodo.6421226" httpUri-0="https://zenodo.org/record/6421216/files/figure.png" httpUri-1="https://zenodo.org/record/6421218/files/figure.png" httpUri-2="https://zenodo.org/record/6421220/files/figure.png" httpUri-3="https://zenodo.org/record/6421222/files/figure.png" httpUri-4="https://zenodo.org/record/6421224/files/figure.png" httpUri-5="https://zenodo.org/record/6421226/files/figure.png" pageId="89" pageNumber="102">342–454</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B6DC9EABFFE8FFB2FC91718EDD7EFA62" pageId="89" pageNumber="102" type="materials_examined">
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<paragraph id="FE79CD20FFE8FFB2FC91718EDBFBFAE2" blockId="89.[794,1399,1193,1505]" pageId="89" pageNumber="102">
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<emphasis id="CCB21132FFE8FFB2FC91718EDBCAFB42" bold="true" box="[794,946,1193,1217]" pageId="89" pageNumber="102">Type material</emphasis>
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:
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<materialsCitation id="4EAEC77DFFE8FFB2FC4A718EDBFBFAE2" ID-GBIF-Occurrence="3455118301" collectingDate="1985-11" collectingMethod="Flight interception trap" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." country="Indonesia" determinerName="V. Assing 2016" elevation="200" location="Sulawesi-Utara" pageId="89" pageNumber="102" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="217D7382FFE8FFB2FC4A718EDC5CFB42" box="[961,1060,1193,1217]" pageId="89" pageNumber="102" type="holotype">Holotype</typeStatus>
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<specimenCount id="E8C006A9FFE8FFB2FBA5718DDC3BFB42" box="[1070,1091,1194,1217]" pageId="89" pageNumber="102" type="male">♂</specimenCount>
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: “
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<collectingCountry id="86D18DB0FFE8FFB2FBD3718EDCBCFB42" box="[1112,1220,1193,1217]" name="Indonesia" pageId="89" pageNumber="102">Indonesia</collectingCountry>
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:
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<location id="FB199BFBFFE8FFB2FB5B718EDD0BFB42" LSID="urn:lsid:plazi:treatment:766F7C36FFE8FFB1FCB47174D940F9C2:FB199BFBFFE8FFB2FB5B718EDD0BFB42" box="[1232,1395,1193,1217]" country="Indonesia" name="Sulawesi-Utara" pageId="89" pageNumber="102">Sulawesi-Utara</location>
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,
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<location id="FB199BFBFFE8FFB2FC9771EDDB93FB62" LSID="urn:lsid:plazi:treatment:766F7C36FFE8FFB1FCB47174D940F9C2:FB199BFBFFE8FFB2FC9771EDDB93FB62" box="[796,1003,1226,1250]" country="Indonesia" name="Dumoga-Bone N. P" pageId="89" pageNumber="102">Dumoga-Bone N.P</location>
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.,
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<date id="8A78EBE0FFE8FFB2FC7071EEDCDEFB62" box="[1019,1190,1225,1249]" pageId="89" pageNumber="102" value="1985-11">
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<collectingDate id="9A3C1208FFE8FFB2FC7071EEDCDEFB62" box="[1019,1190,1225,1249]" pageId="89" pageNumber="102" value="1985-11">November 1985</collectingDate>
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</date>
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/ Plot A, ca
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<quantity id="393E60C5FFE8FFB2FABB71EEDD0BFB62" box="[1328,1395,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="102" unit="m" value="200.0">
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<elevation id="75EB2A13FFE8FFB2FABB71EEDD0BFB62" box="[1328,1395,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="102" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
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<location id="FB199BFBFFE8FFB2FC9071CEDBC7FA82" LSID="urn:lsid:plazi:treatment:766F7C36FFE8FFB1FCB47174D940F9C2:FB199BFBFFE8FFB2FC9071CEDBC7FA82" box="[795,959,1257,1281]" country="Indonesia" name="Lowland forest" pageId="89" pageNumber="102">Lowland forest</location>
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/
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<collectingMethod id="2787B537FFE8FFB2FC5771CEDCA3FA82" box="[988,1243,1257,1281]" pageId="89" pageNumber="102">Flight interception trap</collectingMethod>
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/ R. Ent. Soc. Lond., Project Wallace, B.M. 1985-17 / Holotypus ♂
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<taxonomicName id="39C6B6A3FFE8FFB2FC91700EDC50FAC2" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[794,1064,1321,1345]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="sulawesicus" status="sp. nov.">
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<emphasis id="CCB21132FFE8FFB2FC91700EDC50FAC2" box="[794,1064,1321,1345]" italics="true" pageId="89" pageNumber="102">Tetrabothrus sulawesicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D781AC49FFE8FFB2FBB0700DDC0BFAC2" box="[1083,1139,1322,1345]" pageId="89" pageNumber="102" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="0BA521E4FFE8FFB2FB4F700DDD16FAC2" box="[1220,1390,1321,1345]" pageId="89" pageNumber="102">
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V.
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<bibRefCitation id="9A57B0D1FFE8FFB2FB54700EDD16FAC2" author="ASSING, V." box="[1247,1390,1321,1345]" pageId="89" pageNumber="102" pagination="87 - 175" refId="ref76876" refString="ASSING, V. 2016: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">Assing 2016</bibRefCitation>
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</determinerName>
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” (
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<collectionCode id="98D755E5FFE8FFB2FCAA706EDB0FFAE2" box="[801,887,1353,1377]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="89" pageNumber="102">BMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="FE79CD20FFE8FFB2FC91704DDD7EFA62" blockId="89.[794,1399,1193,1505]" pageId="89" pageNumber="102">
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<materialsCitation id="4EAEC77DFFE8FFB2FC91704DDD79FA02" ID-GBIF-Occurrence="3455118404" box="[794,1281,1385,1410]" collectingDate="1985-11" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." country="Indonesia" elevation="200" location="Lowland forest" pageId="89" pageNumber="102" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="217D7382FFE8FFB2FC91704DDBFCFA01" box="[794,900,1386,1410]" pageId="89" pageNumber="102" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="E8C006A9FFE8FFB2FC04704EDBCCFA02" box="[911,948,1385,1409]" pageId="89" pageNumber="102" type="female">1 ♀</specimenCount>
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: same data as holotype (cAss)
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</materialsCitation>
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;
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<materialsCitation id="4EAEC77DFFE8FFB2FA86704EDCEFFA22" ID-GBIF-Occurrence="3455118307" collectingDate="1985-10" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." country="Indonesia" elevation="200" location="Lowland forest" pageId="89" pageNumber="102" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="E8C006A9FFE8FFB2FA86704EDD4AFA02" box="[1293,1330,1385,1409]" pageId="89" pageNumber="102" type="male">1 ♂</specimenCount>
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: same data, but “
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<date id="8A78EBE0FFE8FFB2FC0070AEDC58FA22" box="[907,1056,1417,1441]" pageId="89" pageNumber="102" value="1985-10">
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<collectingDate id="9A3C1208FFE8FFB2FC0070AEDC58FA22" box="[907,1056,1417,1441]" pageId="89" pageNumber="102" value="1985-10">October 1985</collectingDate>
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</date>
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” (
|
||
<collectionCode id="98D755E5FFE8FFB2FBB170AEDCF7FA22" box="[1082,1167,1417,1441]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="89" pageNumber="102">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EAEC77DFFE8FFB2FB2F70AEDC2EFA42" ID-GBIF-Occurrence="3455118306" collectingDate="1985-02-24" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." country="Indonesia" elevation="200" location="Lowland forest" pageId="89" pageNumber="102" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="E8C006A9FFE8FFB2FB2F70AEDCB1FA22" box="[1188,1225,1417,1441]" pageId="89" pageNumber="102" type="female">1 ♀</specimenCount>
|
||
: same data, but “
|
||
<date id="8A78EBE0FFE8FFB2FCA8708EDBA5FA42" box="[803,989,1449,1473]" pageId="89" pageNumber="102" value="1985-02-24">
|
||
<collectingDate id="9A3C1208FFE8FFB2FCA8708EDBA5FA42" box="[803,989,1449,1473]" pageId="89" pageNumber="102" value="1985-02-24">24 Februar 1985</collectingDate>
|
||
</date>
|
||
” (
|
||
<collectionCode id="98D755E5FFE8FFB2FC71708EDC37FA42" box="[1018,1103,1449,1473]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="89" pageNumber="102">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EAEC77DFFE8FFB2FBED708EDD79FA62" ID-GBIF-Occurrence="3455118555" collectingDate="1985-04" collectionCode="BMNH" collectorName="R. Ent. Soc. Lond." country="Indonesia" elevation="300" location="Lowland forest" pageId="89" pageNumber="102" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="E8C006A9FFE8FFB2FBED708EDCF3FA42" box="[1126,1163,1449,1473]" pageId="89" pageNumber="102" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="E8C006A9FFE8FFB2FB17708EDCB9FA42" box="[1180,1217,1449,1473]" pageId="89" pageNumber="102" type="female">1 ♀</specimenCount>
|
||
: same data, but “
|
||
<date id="8A78EBE0FFE8FFB2FCA970EEDBE8FA62" box="[802,912,1481,1505]" pageId="89" pageNumber="102" value="1985-04">
|
||
<collectingDate id="9A3C1208FFE8FFB2FCA970EEDBE8FA62" box="[802,912,1481,1505]" pageId="89" pageNumber="102" value="1985-04">April 1985</collectingDate>
|
||
</date>
|
||
/ Plot B, ca
|
||
<quantity id="393E60C5FFE8FFB2FB8470EEDC29FA62" box="[1039,1105,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="89" pageNumber="102" unit="m" value="300.0">
|
||
<elevation id="75EB2A13FFE8FFB2FB8470EEDC29FA62" box="[1039,1105,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="89" pageNumber="102" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
” (
|
||
<collectionCode id="98D755E5FFE8FFB2FBEC70EEDCC3FA62" box="[1127,1211,1481,1505]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="89" pageNumber="102">BMNH</collectionCode>
|
||
, cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFFE8FFB2FC91732EDBB8F9E2" pageId="89" pageNumber="102" type="discussion">
|
||
<paragraph id="FE79CD20FFE8FFB2FC91732EDBB8F9E2" blockId="89.[794,1399,1545,1633]" pageId="89" pageNumber="102">
|
||
<emphasis id="CCB21132FFE8FFB2FC91732EDBF1F9A2" bold="true" box="[794,905,1545,1569]" pageId="89" pageNumber="102">Comment</emphasis>
|
||
: The specific epithet (adjective) is derived from
|
||
<collectingRegion id="3C0203C2FFE8FFB2FCD3730EDBCDF9C2" box="[856,949,1577,1601]" country="Indonesia" name="Sulawesi Utara" pageId="89" pageNumber="102">Sulawesi</collectingRegion>
|
||
, the Indonesian island where the species was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFFE8FFB1FC9173AED9DBFAE2" lastPageId="90" lastPageNumber="103" pageId="89" pageNumber="102" type="description">
|
||
<paragraph id="FE79CD20FFE8FFB2FC9173AEDC53F822" blockId="89.[792,1401,1673,2081]" pageId="89" pageNumber="102">
|
||
<emphasis id="CCB21132FFE8FFB2FC9173AEDBE4F922" bold="true" box="[794,924,1673,1697]" pageId="89" pageNumber="102">Description</emphasis>
|
||
: Body length 4.5–6.0 mm; length of forebody
|
||
<quantity id="393E60C5FFE8FFB2FC93738EDBE3F942" box="[792,923,1705,1729]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="89" pageNumber="102" unit="mm" value="2.2" valueMax="2.5" valueMin="1.9">1.9–2.5 mm</quantity>
|
||
. Coloration: head and pronotum variable, bright-reddish to blackish-brown; elytra blackish-brown; abdomen variable, reddish to black; legs bicoloured: femora pale yellowish with the apices – especially those of the metafemora rather broadly – brown to blackishbrown, tibiae and tarsi brown; antennae black with antennomeres I–III and at least the apex of XI reddish; maxillary palpi yellowish.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFE8FFB1FC91728EDAFEFEC2" blockId="89.[792,1401,1673,2081]" lastBlockId="90.[189,794,233,1025]" lastPageId="90" lastPageNumber="103" pageId="89" pageNumber="102">
|
||
Head (
|
||
<figureCitation id="66FDD1A5FFE8FFB2FCE5728EDBBDF842" box="[878,965,1961,1985]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 167</figureCitation>
|
||
) strongly transverse, 1.25–1.30 times as broad as long; posterior angles obsolete; punctation extremely fine and moderately dense, except along the impunctate middle of the dorsal surface; pubescence long and pale. Eyes large, approximately as long as distance from posterior margin of eye to posterior constriction of head. Antenna (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FEEC740ED9C7FEC2" box="[359,447,297,321]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="90" pageNumber="103">Fig. 168</figureCitation>
|
||
)
|
||
<quantity id="393E60C5FFEBFFB1FE47740EDA33FEC2" box="[460,587,297,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="90" pageNumber="103" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFEBFFB1FF36746DDB7FFE42" blockId="90.[189,794,233,1025]" pageId="90" pageNumber="103">
|
||
Pronotum (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FEB0746ED9EBFEE2" box="[315,403,329,353]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="90" pageNumber="103">Fig. 167</figureCitation>
|
||
) weakly transverse, 1.06–1.15 times as broad as long and 1.02–1.08 times as broad as head, strongly convex in cross-section; punctation fine and moderately dense; pubescence pale, long, and suberect.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFEBFFB1FF3674EEDA41FDE2" blockId="90.[189,794,233,1025]" pageId="90" pageNumber="103">
|
||
Elytra (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FE9B74EED91FFE62" box="[272,359,457,481]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="90" pageNumber="103">Fig. 167</figureCitation>
|
||
) 0.85–0.90 times as long as pronotum; punctation denser and less fine than that of pronotum; pubescence very long and suberect. Hind wings fully developed. Metatarsomere as long as the combined length of metatarsomeres II and III.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFEBFFB1FF36774EDA18FCC2" blockId="90.[189,794,233,1025]" pageId="90" pageNumber="103">
|
||
Abdomen (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FECB774ED9E0FD02" box="[320,408,617,641]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="90" pageNumber="103">Fig. 169</figureCitation>
|
||
) narrower than elytra and with moderately deep anterior impressions on tergites III– VI; tergites III–VII each with four setiferous punctures at posterior margins; tergite VII without non-setiferous punctation, posterior margin with palisade fringe; tergite VIII with dense fine setiferous punctures near anterior margin, otherwise impunctate.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFEBFFB1FF36766DDAA7FB82" blockId="90.[189,794,233,1025]" pageId="90" pageNumber="103">
|
||
<specimenCount id="E8C006A9FFEBFFB1FF36766DD8AAFCE2" box="[189,210,842,865]" pageId="90" pageNumber="103" type="male">♂</specimenCount>
|
||
: posterior margin of sternite VIII strongly produced, convexly pointed in the middle (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FDAF764EDA06FC02" box="[548,638,873,897]" captionStart="Figs 442–458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="90" pageNumber="103">Fig. 454</figureCitation>
|
||
); median lobe of aedeagus (
|
||
<figureCitation id="66FDD1A5FFEBFFB1FEC676AED9A6FC22" box="[333,478,905,929]" captionStart="Figs 442–458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="90" pageNumber="103">Figs 452–453</figureCitation>
|
||
)
|
||
<quantity id="393E60C5FFEBFFB1FE6576AEDAFFFC22" box="[494,647,905,929]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="90" pageNumber="103" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.53–0.58 mm</quantity>
|
||
long; ventral process subapically weakly curved in lateral view and rather narrow in ventral view; paramere nearly
|
||
<quantity id="393E60C5FFEBFFB1FD4376EEDB62FC62" box="[712,794,969,993]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="90" pageNumber="103" unit="mm" value="0.6">0.6 mm</quantity>
|
||
long, its apical lobe of moderate length and slender.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FFEBFFB1FF36710ED9DBFAE2" blockId="90.[189,794,1065,1377]" pageId="90" pageNumber="103">
|
||
<emphasis id="CCB21132FFEBFFB1FF36710ED9E8FBC2" bold="true" box="[189,400,1065,1089]" pageId="90" pageNumber="103">Comparative notes</emphasis>
|
||
:
|
||
<taxonomicName id="39C6B6A3FFEBFFB1FE2B710EDADEFBC2" authorityName="Assing" authorityYear="2016" box="[416,678,1065,1089]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="103" phylum="Arthropoda" rank="species" species="sulawesicus">
|
||
<emphasis id="CCB21132FFEBFFB1FE2B710EDADEFBC2" box="[416,678,1065,1089]" italics="true" pageId="90" pageNumber="103">Tetrabothrus sulawesicus</emphasis>
|
||
</taxonomicName>
|
||
is reliably distinguished from other congeners with bicoloured legs and without non-setiferous punctation on tergite VII (and VIII) only by the shape of the median lobe of the aedeagus, from most species also by the more strongly pointed posterior margin of the male sternite VIII. It additionally differs from
|
||
<taxonomicName id="39C6B6A3FFEBFFB1FE6371CDDA2EFA82" authorityName="Bernhauer" authorityYear="1915" box="[488,598,1257,1281]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="103" phylum="Arthropoda" rank="species" species="clavatus">
|
||
<emphasis id="CCB21132FFEBFFB1FE6371CDDA2EFA82" box="[488,598,1257,1281]" italics="true" pageId="90" pageNumber="103">T. clavatus</emphasis>
|
||
</taxonomicName>
|
||
, the only other representative of the genus recorded from
|
||
<collectingRegion id="3C0203C2FFEBFFB1FD1D702EDA8BFAA2" box="[662,755,1289,1313]" country="Indonesia" name="Sulawesi Utara" pageId="90" pageNumber="103">Sulawesi</collectingRegion>
|
||
, by smaller and less bulging eyes, bicoloured legs, and darker antennomeres IV–IX.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFFEBFFB1FF3670AED940F9C2" pageId="90" pageNumber="103" type="distribution">
|
||
<paragraph id="FE79CD20FFEBFFB1FF3670AED940F9C2" blockId="90.[189,794,1417,1601]" pageId="90" pageNumber="103">
|
||
<emphasis id="CCB21132FFEBFFB1FF3670AEDA44FA22" bold="true" box="[189,572,1417,1441]" pageId="90" pageNumber="103">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="217D7382FFEBFFB1FD0270ADDAC0FA21" box="[649,696,1418,1442]" pageId="90" pageNumber="103">type</typeStatus>
|
||
locality, today Bogani Nani Wartabone National Park, is situated on Minahassa Peninsula in the north of
|
||
<collectingRegion id="3C0203C2FFEBFFB1FD3C70EEDB6EFA62" box="[695,790,1481,1505]" country="Indonesia" name="Sulawesi Utara" pageId="90" pageNumber="103">Sulawesi</collectingRegion>
|
||
,
|
||
<collectingCountry id="86D18DB0FFEBFFB1FF3670CED953F982" box="[189,299,1513,1537]" name="Indonesia" pageId="90" pageNumber="103">Indonesia</collectingCountry>
|
||
. The specimens were collected with flight interception traps in lowland forests at altitudes of 200 and
|
||
<specimenCount id="E8C006A9FFEBFFB1FF65730ED940F9C2" box="[238,312,1577,1601]" pageId="90" pageNumber="103" type="generic">
|
||
<quantity id="393E60C5FFEBFFB1FF65730ED94DF9C2" box="[238,309,1577,1601]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="90" pageNumber="103" unit="m" value="300.0">300 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |