treatments-xml/data/76/69/87/76698788FFACE317FF6CFC39415A2D7B.xml

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<mods:title id="EF7946D9DE0800B364A5D1284A3919BE">New taxonomic considerations in Zodariellum Andreeva &amp; Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera</mods:title>
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<mods:namePart id="C71604448BE107952FDCE9F83976CBC8">Zamani, Alireza</mods:namePart>
<mods:affiliation id="456DA888E5FFEB26BB835F7D5193933C">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland.</mods:affiliation>
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<mods:namePart id="F6439F48721240E5CE234321965D27AC">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="BA3775E7F40ECB288CCC3CE300C322DE">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia. &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<heading id="A53781F2FFACE314FF6CFC3942FD2CB6" bold="true" box="[151,750,982,1009]" fontSize="11" level="1" pageId="1" pageNumber="162" reason="1">
<emphasis id="CCB4EA8CFFACE314FF6CFC3942FD2CB6" bold="true" box="[151,750,982,1009]" pageId="1" pageNumber="162">
Genus 
<taxonomicName id="39C04D1DFFACE314FF11FC3942FD2CB6" ID-CoL="63WNV" ID-ENA="2987170" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" box="[234,750,982,1009]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FF11FC39416A2CB7" bold="true" box="[234,377,982,1009]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
<bibRefCitation id="9A514B6FFFACE314FE7AFC3942FD2CB6" author="Andreeva, E. M. &amp; Tyshchenko, V. P." box="[385,750,982,1009]" pageId="1" pageNumber="162" pagination="684 - 689" refId="ref7055" refString="Andreeva, E. M. &amp; Tyshchenko, V. P. (1968) Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskii Zhurnal, 47, 684 - 689. [in Russian]" type="journal article" year="1968">Andreeva &amp; Tyshchenko, 1968</bibRefCitation>
</taxonomicName>
</emphasis>
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<taxonomicName id="39C04D1DFFACE314FF6CFBF442962B75" ID-CoL="63WNV" authority="Andreeva &amp; Tystshenko, 1968: 688" authorityName="Andreeva &amp; Tystshenko" authorityPageNumber="688" authorityYear="1968" box="[151,645,1051,1075]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FF6CFBF441072B75" box="[151,276,1051,1075]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
Andreeva &amp; Tystshenko, 1968: 688
</taxonomicName>
.
</paragraph>
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<treatmentCitation id="7F61108FFFACE314FF6CFBD542402B14" author="Marusik, Y. M. &amp; Koponen, S." box="[151,595,1082,1106]" page="40" pageId="1" pageNumber="162" year="2001">
<taxonomicName id="39C04D1DFFACE314FF6CFBD542402B14" ID-CoL="63WNV" authority=": Marusik &amp; Koponen 2001: 40" authorityName=": Marusik &amp; Koponen" authorityPageNumber="40" authorityYear="2001" box="[151,595,1082,1106]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FF6CFBD541072B14" box="[151,276,1082,1106]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
: 
<bibRefCitation id="9A514B6FFFACE314FED9FBD542402B14" author="Marusik, Y. M. &amp; Koponen, S." box="[290,595,1082,1106]" pageId="1" pageNumber="162" pagination="39 - 48" refId="ref7521" refString="Marusik, Y. M. &amp; Koponen, S. (2001) Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia, 34, 39 - 48." type="journal article" year="2001">Marusik &amp; Koponen 2001: 40</bibRefCitation>
</taxonomicName>
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(removed from the synonymy of 
<taxonomicName id="39C04D1DFFACE314FC52FBD544142B14" ID-CoL="8B2B" authorityName="Walckenaer" authorityYear="1826" box="[937,1031,1082,1106]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FC52FBD544142B14" box="[937,1031,1082,1106]" italics="true" pageId="1" pageNumber="162">Zodarion</emphasis>
</taxonomicName>
).— 
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<bibRefCitation id="9A514B6FFFACE314FBD7FBD545722B14" author="Zamani, A. &amp; Marusik, Y. M." box="[1068,1377,1082,1106]" pageId="1" pageNumber="162" pagination="145 - 193" refId="ref7939" refString="Zamani, A. &amp; Marusik, Y. M. (2021) Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species. ZooKeys, 1035, 145 - 193. https: // doi. org / 10.3897 / zookeys. 1035.65767" type="journal article" year="2021">Zamani &amp; Marusik 2021: 184</bibRefCitation>
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.
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<paragraph id="FE7F369EFFACE314FF6CFB6F45792BDC" blockId="1.[151,1437,1152,2043]" box="[151,1386,1152,1178]" pageId="1" pageNumber="162">
<materialsCitation id="4EA83CC3FFACE314FF6CFB6F45752BDC" ID-GBIF-Occurrence="4435949303" box="[151,1382,1152,1178]" country="Tajikistan" location="Tajikistan" pageId="1" pageNumber="162" specimenCount="1" typeStatus="holotype">
<emphasis id="CCB4EA8CFFACE314FF6CFB6F41212BDC" bold="true" box="[151,306,1152,1178]" pageId="1" pageNumber="162">
<typeStatus id="217B883CFFACE314FF6CFB6F40C12BDC" box="[151,210,1152,1178]" pageId="1" pageNumber="162">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="39C04D1DFFACE314FEC2FB6F43B32BDC" ID-CoL="7GF4P" authority="Andreeva &amp; Tyshchenko, 1968" authorityName="Andreeva &amp; Tyshchenko" authorityYear="1968" box="[313,928,1152,1178]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis id="CCB4EA8CFFACE314FEC2FB6F422E2BDC" box="[313,573,1152,1178]" italics="true" pageId="1" pageNumber="162">Zodariellum surprisum</emphasis>
<bibRefCitation id="9A514B6FFFACE314FDB8FB6F43B32BDC" author="Andreeva, E. M. &amp; Tyshchenko, V. P." box="[579,928,1152,1178]" pageId="1" pageNumber="162" pagination="684 - 689" refId="ref7055" refString="Andreeva, E. M. &amp; Tyshchenko, V. P. (1968) Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskii Zhurnal, 47, 684 - 689. [in Russian]" type="journal article" year="1968">Andreeva &amp; Tyshchenko, 1968</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="86D7760EFFACE314FC1FFB6F44462BDC" box="[996,1109,1152,1178]" name="Tajikistan" pageId="1" pageNumber="162">Tajikistan</collectingCountry>
, by original designation
</materialsCitation>
.
</paragraph>
<paragraph id="FE7F369EFFACE314FF3CFB4B42E1290C" blockId="1.[151,1437,1152,2043]" pageId="1" pageNumber="162">
<emphasis id="CCB4EA8CFFACE314FF3CFB4B41A52BF8" bold="true" box="[199,438,1188,1214]" pageId="1" pageNumber="162">Emended diagnosis.</emphasis>
Males of 
<taxonomicName id="39C04D1DFFACE314FDD2FB4B42A02BF8" ID-CoL="63WNV" authorityName=": Marusik &amp; Koponen" authorityYear="2001" box="[553,691,1188,1214]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FDD2FB4B42A02BF8" box="[553,691,1188,1214]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
</taxonomicName>
differ from that of 
<taxonomicName id="39C04D1DFFACE314FC6AFB4B45092BF8" ID-CoL="5D6C8" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[913,1306,1188,1214]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="nitidum">
<emphasis id="CCB4EA8CFFACE314FC6AFB4B44442BF8" box="[913,1111,1188,1214]" italics="true" pageId="1" pageNumber="162">Zodarion nitidum</emphasis>
(Audouin, 1826)
</taxonomicName>
(
<figureCitation id="66FB2A1BFFACE314FAD1FB4B40BF2BA4" captionStart="FIGURE 11" captionStartId="11.[151,250,1553,1577]" captionTargetBox="[196,1390,807,1530]" captionTargetId="figure-58@11.[196,1390,807,1530]" captionTargetPageId="11" captionText="FIGURE 11. SEM images of the male palp of Zodarion nitidum. A ventral view; B retrolateral view; C dorsal view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Tg—tegular apophysis, Va—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7022271" httpUri="https://zenodo.org/record/7022271/files/figure.png" pageId="1" pageNumber="162">Fig. 11A– C</figureCitation>
), the generotype of 
<taxonomicName id="39C04D1DFFACE314FE74FB2741E42BA4" ID-CoL="8B2B" authorityName="Walckenaer" authorityYear="1826" box="[399,503,1224,1250]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FE74FB2741E42BA4" box="[399,503,1224,1250]" italics="true" pageId="1" pageNumber="162">Zodarion</emphasis>
</taxonomicName>
, by having: 1) RTA (
<emphasis id="CCB4EA8CFFACE314FD1CFB2643152BA4" box="[743,774,1225,1250]" italics="true" pageId="1" pageNumber="162">Ra</emphasis>
) longer than wide, with solid base and either bifurcated or solid tip, with branch or branches (
<emphasis id="CCB4EA8CFFACE314FDDDFB0342562A40" box="[550,581,1260,1286]" italics="true" pageId="1" pageNumber="162">Rb</emphasis>
) bent retrolaterally at more than right angle (
<figureCitation id="66FB2A1BFFACE314FBB2FB0344D92A40" box="[1097,1226,1260,1286]" captionStart="FIGURE 4" captionStartId="5.[151,250,1907,1931]" captionTargetBox="[192,1383,207,1870]" captionTargetId="figure-17@5.[175,1412,181,1884]" captionTargetPageId="5" captionText="FIGURE 4. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.2 mm. Abbreviations: Ap—anterodorsal part of retrolateral tibial apophysis, Cn—conductor, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022249" httpUri="https://zenodo.org/record/7022249/files/figure.png" pageId="1" pageNumber="162">Figs 4A–D</figureCitation>
, 
<figureCitation id="66FB2A1BFFACE314FB2DFB03450E2A41" box="[1238,1309,1260,1287]" captionStart="FIGURE 5" captionStartId="6.[151,250,1584,1608]" captionTargetBox="[177,1411,181,1560]" captionTargetId="figure-21@6.[177,1411,181,1560]" captionTargetPageId="6" captionText="FIGURE 5. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Di—diverticulum, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022251" httpUri="https://zenodo.org/record/7022251/files/figure.png" pageId="1" pageNumber="162">5A–D</figureCitation>
) (
<emphasis id="CCB4EA8CFFACE314FACDFB02455E2A40" box="[1334,1357,1261,1286]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. wider than long and divided into 3 parts basally; 
<figureCitation id="66FB2A1BFFACE314FD71FAFF42FC2A6C" box="[650,751,1296,1322]" captionStart="FIGURE 11" captionStartId="11.[151,250,1553,1577]" captionTargetBox="[196,1390,807,1530]" captionTargetId="figure-58@11.[196,1390,807,1530]" captionTargetPageId="11" captionText="FIGURE 11. SEM images of the male palp of Zodarion nitidum. A ventral view; B retrolateral view; C dorsal view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Tg—tegular apophysis, Va—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7022271" httpUri="https://zenodo.org/record/7022271/files/figure.png" pageId="1" pageNumber="162">Fig. 11B</figureCitation>
), 2) lacking ventral tibial apophysis (
<emphasis id="CCB4EA8CFFACE314FB5AFAFE44AB2A6C" box="[1185,1208,1297,1322]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. present, 
<emphasis id="CCB4EA8CFFACE314FAD0FAFE45542A6C" box="[1323,1351,1297,1322]" italics="true" pageId="1" pageNumber="162">Va</emphasis>
in 
<figureCitation id="66FB2A1BFFACE314FA8BFAFF40F82A08" captionStart="FIGURE 11" captionStartId="11.[151,250,1553,1577]" captionTargetBox="[196,1390,807,1530]" captionTargetId="figure-58@11.[196,1390,807,1530]" captionTargetPageId="11" captionText="FIGURE 11. SEM images of the male palp of Zodarion nitidum. A ventral view; B retrolateral view; C dorsal view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Tg—tegular apophysis, Va—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7022271" httpUri="https://zenodo.org/record/7022271/files/figure.png" pageId="1" pageNumber="162">Fig. 11A–B</figureCitation>
), 3) cymbium with short tip (i.e. as long as wide, shorter than RTA) and prominent tutaculum (
<emphasis id="CCB4EA8CFFACE314FAD9FADA452D2A08" box="[1314,1342,1333,1358]" italics="true" pageId="1" pageNumber="162">Tu</emphasis>
) (
<emphasis id="CCB4EA8CFFACE314FAA3FADA457C2A08" box="[1368,1391,1333,1358]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. tip longer than wide and longer than RTA, tutaculum absent), 4) large tegular apophysis (
<emphasis id="CCB4EA8CFFACE314FB8EFAB644802A34" box="[1141,1171,1369,1394]" italics="true" pageId="1" pageNumber="162">Tg</emphasis>
) longer that bulb, with prolateral spur (
<emphasis id="CCB4EA8CFFACE314FEB6FA93417A2AD0" box="[333,361,1404,1430]" italics="true" pageId="1" pageNumber="162">Sp</emphasis>
) located basally and directed postero-ectally, and long retrolateral arm (
<emphasis id="CCB4EA8CFFACE314FB5BFA9244AA2AD0" box="[1184,1209,1405,1430]" italics="true" pageId="1" pageNumber="162">Tr</emphasis>
) with tip resting on tutaculum (
<emphasis id="CCB4EA8CFFACE314FEE3FA4E413C2AFC" box="[280,303,1441,1466]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. tegular apophysis shorter than bulb), 5) very long conductor (
<emphasis id="CCB4EA8CFFACE314FC0CFA4F440B2AFC" box="[1015,1048,1440,1466]" italics="true" pageId="1" pageNumber="162">Cn</emphasis>
), longer than bulb, with prolateral arm (
<emphasis id="CCB4EA8CFFACE314FF28FA2B40E72A98" box="[211,244,1476,1502]" italics="true" pageId="1" pageNumber="162">Cp</emphasis>
) terminating at a 6 o’clock position or earlier (
<emphasis id="CCB4EA8CFFACE314FCF8FA2A43092A98" box="[771,794,1477,1502]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. conductor lacking prolateral arm and shorter than bulb), and 6) longer filamentous embolus originating at a 6 o’clock position or earlier (
<emphasis id="CCB4EA8CFFACE314FBDBFA0644242944" box="[1056,1079,1513,1538]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. 6:30). Females of 
<taxonomicName id="39C04D1DFFACE314FAE9FA07458F2944" ID-CoL="63WNV" authorityName=": Marusik &amp; Koponen" authorityYear="2001" box="[1298,1436,1512,1538]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FAE9FA07458F2944" box="[1298,1436,1512,1538]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="39C04D1DFFACE314FE87F9E341F72960" ID-CoL="8B2B" authorityName="Walckenaer" authorityYear="1826" box="[380,484,1548,1574]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="CCB4EA8CFFACE314FE87F9E341F72960" box="[380,484,1548,1574]" italics="true" pageId="1" pageNumber="162">Zodarion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D78757F7FFACE314FE17F9E2424F2960" box="[492,604,1548,1575]" pageId="1" pageNumber="162" sensu="lato">sensu lato</taxonomicNameLabel>
by the long, twisted (i.e. similar to a screw), converging copulatory ducts (
<figureCitation id="66FB2A1BFFACE314FF64F9DF413B290C" box="[159,296,1584,1610]" captionStart="FIGURE 10" captionStartId="11.[151,250,722,746]" captionTargetBox="[381,1206,181,698]" captionTargetId="figure-17@11.[381,1206,181,698]" captionTargetPageId="11" captionText="FIGURE 10. Epigynes of Zodariellum nenilini (A) and Z. proszynskii (B–D). A–B intact, ventral view; C–D macerated, dorsal and ventral views. Scale bars = 0.2 mm. Abbreviations: Cd—copulatory duct, Fo—fovea, Re—receptacle." figureDoi="http://doi.org/10.5281/zenodo.7022269" httpUri="https://zenodo.org/record/7022269/files/figure.png" pageId="1" pageNumber="162">Fig. 10A–D</figureCitation>
) (
<emphasis id="CCB4EA8CFFACE314FEBBF9DE4144290C" box="[320,343,1585,1610]" italics="true" pageId="1" pageNumber="162">vs</emphasis>
. not twisted or diverging if twisted).
</paragraph>
<paragraph id="FE7F369EFFACE314FF3CF9BB44B62865" blockId="1.[151,1437,1152,2043]" pageId="1" pageNumber="162">
Furthermore, males of 
<taxonomicName id="39C04D1DFFACE314FE3DF9BB42432928" ID-CoL="63WNV" authorityName=": Marusik &amp; Koponen" authorityYear="2001" box="[454,592,1620,1646]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FE3DF9BB42432928" box="[454,592,1620,1646]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
</taxonomicName>
differ from those of all other zodariid genera by having a very deep (=long) diverticulum of cymbium (
<emphasis id="CCB4EA8CFFACE314FE38F99641CC29D4" box="[451,479,1657,1682]" italics="true" pageId="1" pageNumber="162">Di</emphasis>
, 
<figureCitation id="66FB2A1BFFACE314FE17F997422E29D4" box="[492,573,1656,1682]" captionStart="FIGURE 2" captionStartId="3.[151,250,1852,1876]" captionTargetBox="[191,1400,424,1829]" captionTargetId="figure-65@3.[173,1414,412,1829]" captionTargetPageId="3" captionText="FIGURE 2. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B, F), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E). A–E ventral view; F dorsal view of cymbium, showing cymbial diverticulum. Scale bars = 0.2 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branch(es) of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, To—triangular outgrowth, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022243" httpUri="https://zenodo.org/record/7022243/files/figure.png" pageId="1" pageNumber="162">Fig. 2F</figureCitation>
) (about 0.4 time of cymbium length and 0.75 of cymbium width). The species of 
<taxonomicName id="39C04D1DFFACE314FF4EF973416A29F0" ID-CoL="M2C" authorityName="Denis" authorityYear="1966" box="[181,377,1692,1718]" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="CCB4EA8CFFACE314FF4EF973416A29F0" box="[181,377,1692,1718]" italics="true" pageId="1" pageNumber="162">Acanthinozodium</emphasis>
</taxonomicName>
also have a cymbial diverticulum, although in the form of a short conical (i.e. depth about one radius) round pit (
<bibRefCitation id="9A514B6FFFACE314FE9EF92F426F299D" author="Jocque, R. &amp; Henrard, A." box="[357,636,1728,1755]" pageId="1" pageNumber="162" pagination="1 - 23" refId="ref7351" refString="Jocque, R. &amp; Henrard, A. (2015) Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy, 114, 1 - 23. https: // doi. org / 10.5852 / ejt. 2015.114" type="journal article" year="2015">Jocqué &amp; Henrard 2015</bibRefCitation>
: fig. 9A, B; 
<bibRefCitation id="9A514B6FFFACE314FCF7F92F4422299C" author="Zamani, A. &amp; Marusik, Y. M." box="[780,1073,1728,1754]" pageId="1" pageNumber="162" pagination="145 - 193" refId="ref7939" refString="Zamani, A. &amp; Marusik, Y. M. (2021) Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species. ZooKeys, 1035, 145 - 193. https: // doi. org / 10.3897 / zookeys. 1035.65767" type="journal article" year="2021">Zamani &amp; Marusik, 2021</bibRefCitation>
: figs 13C, 14G‒H, 15C, F, 21F, 22H‒I, 23C, 24C). Furthermore, the male palps of 
<taxonomicName id="39C04D1DFFACE314FD30F90B43BE29B8" box="[715,941,1764,1790]" pageId="1" pageNumber="162">
<emphasis id="CCB4EA8CFFACE314FD30F90B434629B8" box="[715,853,1764,1790]" italics="true" pageId="1" pageNumber="162">Zodariellum</emphasis>
species
</taxonomicName>
have serrated processes of RTA and serrated spur of the tegular apophysis (
<emphasis id="CCB4EA8CFFACE314FE10F8E742142864" box="[491,519,1800,1826]" italics="true" pageId="1" pageNumber="162">Sp</emphasis>
), which have not been recorded in other 
<taxonomicName id="39C04D1DFFACE314FC28F8E7445F2864" authorityName="Thorell" authorityYear="1881" box="[979,1100,1800,1826]" class="Arachnida" family="Zodariidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="subFamily" subFamily="Zodariinae">Zodariinae</taxonomicName>
genera.
</paragraph>
<paragraph id="FE7F369EFFACE314FF3CF8C344962800" blockId="1.[151,1437,1152,2043]" box="[199,1157,1836,1863]" pageId="1" pageNumber="162">
<emphasis id="CCB4EA8CFFACE314FF3CF8C341742800" bold="true" box="[199,359,1836,1862]" pageId="1" pageNumber="162">Composition.</emphasis>
21 species (including those described here and the new combinations):
</paragraph>
<paragraph id="FE7F369EFFACE317FF3CF8BE45942FBF" blockId="1.[151,1437,1152,2043]" lastBlockId="2.[151,1436,150,573]" lastPageId="2" lastPageNumber="163" pageId="1" pageNumber="162">
<taxonomicName id="39C04D1DFFACE314FF3CF8BE422C282C" ID-CoL="94HVR" authority="(Tyshchenko, 1970)" baseAuthorityName="Tyshchenko" baseAuthorityYear="1970" box="[199,575,1872,1898]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="asiaticum">
<emphasis id="CCB4EA8CFFACE314FF3CF8BE4147282C" box="[199,340,1873,1898]" italics="true" pageId="1" pageNumber="162">Z. asiaticum</emphasis>
(Tyshchenko, 1970)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FDA9F8BF4278282C" box="[594,619,1872,1898]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FD90F8BF426F282C" box="[619,636,1872,1898]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FD6EF8BE4402282D" ID-CoL="5D694" authority="(Kroneberg, 1875)" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[661,1041,1872,1899]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis id="CCB4EA8CFFACE314FD6EF8BE42B6282C" box="[661,677,1873,1898]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FD4EF8BF4325282C" box="[693,822,1872,1898]" italics="true" pageId="1" pageNumber="162">bactrianum</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FCBDF8BF441A282D" author="Kroneberg, A." box="[838,1033,1872,1899]" pageId="1" pageNumber="162" pagination="1 - 58" refId="ref7412" refString="Kroneberg, A. (1875) Araneae. In: Fedtschenko, A. P. (Ed.), Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau, 19 (3), 1 - 58, pls. 1 - 5. [in Russian]" type="journal article" year="1875">Kroneberg, 1875</bibRefCitation>
)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FBDFF8BF442E282C" box="[1060,1085,1872,1898]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FBC6F8BF445D282C" box="[1085,1102,1872,1898]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FB9CF8BE458F282D" ID-CoL="93S87" authority="(Nenilin, 1985)" baseAuthorityName="Nenilin" baseAuthorityYear="1985" box="[1127,1436,1872,1899]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="bekuzini">
<emphasis id="CCB4EA8CFFACE314FB9CF8BE4464282C" box="[1127,1143,1873,1898]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FB7CF8BF44F7282C" box="[1159,1252,1872,1898]" italics="true" pageId="1" pageNumber="162">bekuzini</emphasis>
(Nenilin, 1985)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FF5AF89B40A928C8" box="[161,186,1908,1934]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FF41F89B40D828C8" box="[186,203,1908,1934]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FF1AF89A426428C8" ID-CoL="93S88" authority="(Zhu &amp; Zhu, 1983)" baseAuthorityName="Zhu &amp; Zhu" baseAuthorityYear="1983" box="[225,631,1908,1935]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="chaoyangense">
<emphasis id="CCB4EA8CFFACE314FF1AF89A40E228C8" box="[225,241,1909,1934]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FF05F89B418E28C8" box="[254,413,1908,1934]" italics="true" pageId="1" pageNumber="162">chaoyangense</emphasis>
(Zhu &amp; Zhu, 1983)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FD7CF89B42B328C8" box="[647,672,1908,1934]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FD5BF89B42A228C8" box="[672,689,1908,1934]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FD3CF89A44F228C8" ID-CoL="5D69Q" authority="(Andreeva &amp; Tyshchenko, 1968)" baseAuthorityName="Andreeva &amp; Tyshchenko" baseAuthorityYear="1968" box="[711,1249,1908,1935]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="continentale">
<emphasis id="CCB4EA8CFFACE314FD3CF89A42C428C8" box="[711,727,1909,1934]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FD18F89B437E28C8" box="[739,877,1908,1934]" italics="true" pageId="1" pageNumber="162">continentale</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FC80F89B44CA28C8" author="Andreeva, E. M. &amp; Tyshchenko, V. P." box="[891,1241,1908,1934]" pageId="1" pageNumber="162" pagination="684 - 689" refId="ref7055" refString="Andreeva, E. M. &amp; Tyshchenko, V. P. (1968) Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskii Zhurnal, 47, 684 - 689. [in Russian]" type="journal article" year="1968">Andreeva &amp; Tyshchenko, 1968</bibRefCitation>
)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FB0AF89B451928C8" box="[1265,1290,1908,1934]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FAF1F89B450828C8" box="[1290,1307,1908,1934]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FACAF89A410C28F4" ID-CoL="93S8C" authority="(Zhu, 1988)" baseAuthorityName="Zhu" baseAuthorityYear="1988" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="furcum">
<emphasis id="CCB4EA8CFFACE314FACAF89A455228C8" box="[1329,1345,1909,1934]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FAB5F89B458F28C8" box="[1358,1436,1908,1934]" italics="true" pageId="1" pageNumber="162">furcum</emphasis>
(Zhu, 1988)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FEC9F877415828F4" box="[306,331,1944,1970]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FEB0F877414F28F4" box="[331,348,1944,1970]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FE8EF876428B28F4" authority="(Yin, 2012)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Yin" baseAuthorityYear="2012" box="[373,664,1944,1971]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="hunanense" status="comb. nov.">
<emphasis id="CCB4EA8CFFACE314FE8EF876419628F4" box="[373,389,1945,1970]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FE6FF877421F28F4" box="[404,524,1944,1970]" italics="true" pageId="1" pageNumber="162">hunanense</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FDE6F877428228F4" author="Yin, C. M. &amp; Peng, X. J. &amp; Yan, H. M. &amp; Bao, Y. H. &amp; Xu, X. &amp; Tang, G. &amp; Zhou, Q. S. &amp; Liu, P." box="[541,657,1944,1970]" pageId="1" pageNumber="162" refId="ref7867" refString="Yin, C. M., Peng, X. J., Yan, H. M., Bao, Y. H., Xu, X., Tang, G., Zhou, Q. S. &amp; Liu, P. (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">Yin, 2012</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFACE314FD5AF877431A28F4" bold="true" box="[673,777,1944,1970]" pageId="1" pageNumber="162">
<taxonomicNameLabel id="D78757F7FFACE314FD5AF877431A28F4" box="[673,777,1944,1970]" pageId="1" pageNumber="162" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFACE314FCE6F877432328F4" box="[797,816,1944,1970]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FCB2F876452328F4" ID-CoL="93S8F" authority="Marusik &amp; Koponen, 2001" authorityName="Marusik &amp; Koponen" authorityYear="2001" box="[841,1328,1944,1971]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="mongolicum">
<emphasis id="CCB4EA8CFFACE314FCB2F876434A28F4" box="[841,857,1945,1970]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FC92F87743E028F4" box="[873,1011,1944,1970]" italics="true" pageId="1" pageNumber="162">mongolicum</emphasis>
<bibRefCitation id="9A514B6FFFACE314FC07F877452328F4" author="Marusik, Y. M. &amp; Koponen, S." box="[1020,1328,1944,1971]" pageId="1" pageNumber="162" pagination="39 - 48" refId="ref7521" refString="Marusik, Y. M. &amp; Koponen, S. (2001) Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia, 34, 39 - 48." type="journal article" year="2001">Marusik &amp; Koponen, 2001</bibRefCitation>
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FAB8F877454F28F4" box="[1347,1372,1944,1970]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FAA7F877457E28F4" box="[1372,1389,1944,1970]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FA7DF876419C2891" ID-CoL="5D6C2" authority="(Eskov, 1995)" baseAuthorityName="Eskov" baseAuthorityYear="1995" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="CCB4EA8CFFACE314FA7DF876458528F4" box="[1414,1430,1945,1970]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FF6CF85340FE2890" box="[151,237,1980,2006]" italics="true" pageId="1" pageNumber="162">nenilini</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FF00F85341942891" author="Eskov, K. Y. &amp; Marusik, Y. M." box="[251,391,1980,2007]" pageId="1" pageNumber="162" pagination="55 - 94" refId="ref7209" refString="Eskov, K. Y. &amp; Marusik, Y. M. (1995) On the spiders from Saur Mt. range, eastern Kazakhstan (Arachnida: Araneae). Beitrage zur Araneologie, 4, 55 - 94. [for 1994, published in Dec. 1995]" type="journal article" year="1995">Eskov, 1995</bibRefCitation>
)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FE5BF85341AA2890" box="[416,441,1980,2006]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FE42F85341D92890" box="[441,458,1980,2006]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FE1BF85243712890" authority="(Zhang &amp; Zhang, 2019)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Zhang &amp; Zhang" baseAuthorityYear="2019" box="[480,866,1980,2007]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="planum" status="comb. nov.">
<emphasis id="CCB4EA8CFFACE314FE1BF85241E32890" box="[480,496,1981,2006]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FE06F85342422890" box="[509,593,1980,2006]" italics="true" pageId="1" pageNumber="162">planum</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FDA4F85343492890" author="Zhang, B. S. &amp; Zhang, F." box="[607,858,1980,2006]" pageId="1" pageNumber="162" pagination="67 - 87" refId="ref7994" refString="Zhang, B. S. &amp; Zhang, F. (2019) Three new species of the genus Zodarion (Araneae, Zodariidae) from China. ZooKeys, 813, 67 - 87. https: // doi. org / 10.3897 / zookeys. 813.29683" type="journal article" year="2019">Zhang &amp; Zhang, 2019</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFACE314FC93F85343DD2890" bold="true" box="[872,974,1980,2006]" pageId="1" pageNumber="162">
<taxonomicNameLabel id="D78757F7FFACE314FC93F85343DD2890" box="[872,974,1980,2006]" pageId="1" pageNumber="162" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFACE314FC26F85343E72890" box="[989,1012,1980,2006]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FBF2F852458F2891" ID-CoL="93S8H" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" box="[1033,1436,1980,2007]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="proszynskii">
<emphasis id="CCB4EA8CFFACE314FBF2F852440A2890" box="[1033,1049,1981,2006]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FBDDF85344B02890" box="[1062,1187,1980,2006]" italics="true" pageId="1" pageNumber="162">proszynskii</emphasis>
(Nenilin &amp; Fet, 1985)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FF5AF80F40A928BC" box="[161,186,2016,2042]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FF41F80F40D828BC" box="[186,203,2016,2042]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FF1FF80E42B128BC" ID-CoL="93S8K" authority="Marusik &amp; Koponen, 2001" authorityName="Marusik &amp; Koponen" authorityYear="2001" box="[228,674,2016,2043]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="schmidti">
<emphasis id="CCB4EA8CFFACE314FF1FF80E40E728BC" box="[228,244,2017,2042]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FEFFF80F417028BC" box="[260,355,2016,2042]" italics="true" pageId="1" pageNumber="162">schmidti</emphasis>
<bibRefCitation id="9A514B6FFFACE314FE96F80F42B128BC" author="Marusik, Y. M. &amp; Koponen, S." box="[365,674,2016,2043]" pageId="1" pageNumber="162" pagination="39 - 48" refId="ref7521" refString="Marusik, Y. M. &amp; Koponen, S. (2001) Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia, 34, 39 - 48." type="journal article" year="2001">Marusik &amp; Koponen, 2001</bibRefCitation>
</taxonomicName>
(
<specimenCount id="E8C6FD17FFACE314FD4DF80F42DC28BC" box="[694,719,2016,2042]" pageId="1" pageNumber="162" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFACE314FD35F80F42CC28BC" box="[718,735,2016,2042]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE314FD02F80E444428BC" authority="(Charitonov, 1946)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[761,1111,2016,2043]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="spasskyi" status="comb. nov.">
<emphasis id="CCB4EA8CFFACE314FD02F80E431A28BC" box="[761,777,2017,2042]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FCE2F80F436528BC" box="[793,886,2016,2042]" italics="true" pageId="1" pageNumber="162">spasskyi</emphasis>
(
<bibRefCitation id="9A514B6FFFACE314FC7CF80F445C28BC" author="Charitonov, D. E." box="[903,1103,2016,2042]" pageId="1" pageNumber="162" pagination="19 - 32" refId="ref7129" refString="Charitonov, D. E. (1946) New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo, 12, 19 - 32. [in Russian]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFACE314FB9BF80F44DA28BC" bold="true" box="[1120,1225,2016,2042]" pageId="1" pageNumber="162">
<taxonomicNameLabel id="D78757F7FFACE314FB9BF80F44DA28BC" box="[1120,1225,2016,2042]" pageId="1" pageNumber="162" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFACE314FB26F80F44E328BC" box="[1245,1264,2016,2042]" pageId="1" pageNumber="162" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFACE317FAF0F80E421C2FF7" ID-CoL="7GF4P" authority="(Andreeva &amp; Tyshchenko, 1968)" baseAuthorityName="Andreeva &amp; Tyshchenko" baseAuthorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" lastPageId="2" lastPageNumber="163" order="Araneae" pageId="1" pageNumber="162" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis id="CCB4EA8CFFACE314FAF0F80E450828BC" box="[1291,1307,2017,2042]" italics="true" pageId="1" pageNumber="162">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFACE314FAD0F80E458E28BC" box="[1323,1437,2017,2042]" italics="true" pageId="1" pageNumber="162">surprisum</emphasis>
(
<bibRefCitation id="9A514B6FFFAFE317FF64FF7842142FF7" author="Andreeva, E. M. &amp; Tyshchenko, V. P." box="[159,519,151,177]" pageId="2" pageNumber="163" pagination="684 - 689" refId="ref7055" refString="Andreeva, E. M. &amp; Tyshchenko, V. P. (1968) Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskii Zhurnal, 47, 684 - 689. [in Russian]" type="journal article" year="1968">Andreeva &amp; Tyshchenko, 1968</bibRefCitation>
)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFAFE317FDD9FF78422A2FF7" box="[546,569,151,177]" pageId="2" pageNumber="163" type="male">♂</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FDAAFF7844012FF7" ID-CoL="93S8P" authority="(Nenilin &amp; Fet, 1985)" baseAuthorityName="Nenilin &amp; Fet" baseAuthorityYear="1985" box="[593,1042,150,177]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="sytchevskajae">
<emphasis id="CCB4EA8CFFAFE317FDAAFF7842722FF6" box="[593,609,151,176]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FD89FF78431F2FF7" box="[626,780,151,177]" italics="true" pageId="2" pageNumber="163">sytchevskajae</emphasis>
(Nenilin &amp; Fet, 1985)
</taxonomicName>
(
<specimenCount id="E8C6FD17FFAFE317FBDDFF78442C2FF7" box="[1062,1087,151,177]" pageId="2" pageNumber="163" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFAFE317FBC4FF7844432FF7" box="[1087,1104,151,177]" pageId="2" pageNumber="163" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FB92FF78417D2F93" authority="(Andreeva &amp; Tyshchenko, 1968)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Andreeva &amp; Tyshchenko" baseAuthorityYear="1968" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="tadzhikum" status="comb. nov.">
<emphasis id="CCB4EA8CFFAFE317FB92FF78446A2FF6" box="[1129,1145,151,176]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FB71FF7844EE2FF7" box="[1162,1277,151,177]" italics="true" pageId="2" pageNumber="163">tadzhikum</emphasis>
(
<bibRefCitation id="9A514B6FFFAFE317FAF5FF7841752F93" author="Andreeva, E. M. &amp; Tyshchenko, V. P." pageId="2" pageNumber="163" pagination="684 - 689" refId="ref7055" refString="Andreeva, E. M. &amp; Tyshchenko, V. P. (1968) Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskii Zhurnal, 47, 684 - 689. [in Russian]" type="journal article" year="1968">Andreeva &amp; Tyshchenko, 1968</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FE8CFF5441CD2F93" bold="true" box="[375,478,187,213]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FE8CFF5441CD2F93" box="[375,478,187,213]" pageId="2" pageNumber="163" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFAFE317FE0AFF5442172F93" box="[497,516,187,213]" pageId="2" pageNumber="163" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FDE7FF5443AC2F93" authority="(Spassky, 1941)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Spassky" baseAuthorityYear="1941" box="[540,959,187,213]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="testaceofasciatum" status="comb. nov.">
<emphasis id="CCB4EA8CFFAFE317FDE7FF54423F2F92" box="[540,556,187,212]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FDC0FF5443172F93" box="[571,772,187,213]" italics="true" pageId="2" pageNumber="163">testaceofasciatum</emphasis>
(
<bibRefCitation id="9A514B6FFFAFE317FCE8FF5443A42F93" author="Spassky, S." box="[787,951,187,213]" pageId="2" pageNumber="163" pagination="12 - 26" refId="ref7760" refString="Spassky, S. (1941) Araneae palaearcticae novae VI. Folia Zoologica et Hydrobiologica, Riga, 11, 12 - 26." type="journal article" year="1941">Spassky, 1941</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FC3CFF54443C2F93" bold="true" box="[967,1071,187,213]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FC3CFF54443C2F93" box="[967,1071,187,213]" pageId="2" pageNumber="163" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFAFE317FBBAFF5444472F93" box="[1089,1108,187,213]" pageId="2" pageNumber="163" type="female">♀</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FB96FF5444EC2F92" authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[1133,1279,187,213]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="turanicum" status="sp. nov.">
<emphasis id="CCB4EA8CFFAFE317FB96FF54446E2F92" box="[1133,1149,187,212]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FB77FF5444EC2F92" box="[1164,1279,187,212]" italics="true" pageId="2" pageNumber="163">turanicum</emphasis>
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FAFCFF5445542F93" bold="true" box="[1287,1351,187,213]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FAFCFF5445542F93" box="[1287,1351,187,213]" pageId="2" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFAFE317FAA3FF54457C2F93" box="[1368,1391,187,213]" pageId="2" pageNumber="163" type="male">♂</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FA7DFF54413A2FBF" authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="turkmenicum" status="sp. nov.">
<emphasis id="CCB4EA8CFFAFE317FA7DFF5445852F92" box="[1414,1430,187,212]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FF6CFF30413A2FBF" box="[151,297,223,249]" italics="true" pageId="2" pageNumber="163">turkmenicum</emphasis>
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FECBFF3041632FBF" bold="true" box="[304,368,223,249]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FECBFF3041632FBF" box="[304,368,223,249]" pageId="2" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFAFE317FE7BFF3041842FBF" box="[384,407,223,249]" pageId="2" pageNumber="163" type="male">♂</specimenCount>
), 
<taxonomicName id="39C04D1DFFAFE317FE57FF3043202FBF" ID-CoL="93S8R" authority="Ponomarev, 2007" authorityName="Ponomarev" authorityYear="2007" box="[428,819,223,249]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="volgouralense">
<emphasis id="CCB4EA8CFFAFE317FE57FF3041AF2FBE" box="[428,444,223,248]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FE31FF30427A2FBF" box="[458,617,223,249]" italics="true" pageId="2" pageNumber="163">volgouralense</emphasis>
<bibRefCitation id="9A514B6FFFAFE317FD94FF3043202FBF" author="Ponomarev, A. V." box="[623,819,223,249]" pageId="2" pageNumber="163" pagination="87 - 95" refId="ref7703" refString="Ponomarev, A. V. (2007) New taxa of spiders (Aranei) from the south of Russia and western Kazakhstan. Caucasian Entomological Bulletin, 3, 87 - 95. https: // doi. org / 10.23885 / 1814 - 3326 - 2007 - 3 - 2 - 87 - 95" type="journal article" year="2007">Ponomarev, 2007</bibRefCitation>
</taxonomicName>
(
<specimenCount id="E8C6FD17FFAFE317FCBFFF30434F2FBF" box="[836,860,223,249]" pageId="2" pageNumber="163" type="male">♂</specimenCount>
<specimenCount id="E8C6FD17FFAFE317FCA7FF30437E2FBF" box="[860,877,223,249]" pageId="2" pageNumber="163" type="female">♀</specimenCount>
) and 
<taxonomicName id="39C04D1DFFAFE317FC56FF3044F62FBF" authority="(Charitonov, 1946)" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[941,1253,223,249]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="zebra" status="comb. nov.">
<emphasis id="CCB4EA8CFFAFE317FC56FF3043AE2FBE" box="[941,957,223,248]" italics="true" pageId="2" pageNumber="163">Z</emphasis>
. 
<emphasis id="CCB4EA8CFFAFE317FC30FF30441A2FBF" box="[971,1033,223,249]" italics="true" pageId="2" pageNumber="163">zebra</emphasis>
(
<bibRefCitation id="9A514B6FFFAFE317FBECFF3044CF2FBF" author="Charitonov, D. E." box="[1047,1244,223,249]" pageId="2" pageNumber="163" pagination="19 - 32" refId="ref7129" refString="Charitonov, D. E. (1946) New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo, 12, 19 - 32. [in Russian]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FB17FF3045412FBF" bold="true" box="[1260,1362,223,249]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FB17FF3045412FBF" box="[1260,1362,223,249]" pageId="2" pageNumber="163" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
(
<specimenCount id="E8C6FD17FFAFE317FA9FFF3045642FBF" box="[1380,1399,223,249]" pageId="2" pageNumber="163" type="female">♀</specimenCount>
).
</paragraph>
<paragraph id="FE7F369EFFAFE317FF3CFEEC44602E5B" blockId="2.[151,1436,150,573]" box="[199,1139,259,285]" pageId="2" pageNumber="163">According to the shape of the RTA, the genus can be split into three species-groups:</paragraph>
<paragraph id="FE7F369EFFAFE317FF3CFEC8439D2E23" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="163">
1) 
<taxonomicName id="39C04D1DFFAFE317FF0CFEC841562E07" ID-CoL="93S8C" baseAuthorityName="Zhu" baseAuthorityYear="1988" box="[247,325,295,321]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="furcum">
<emphasis id="CCB4EA8CFFAFE317FF0CFEC841562E07" box="[247,325,295,321]" italics="true" pageId="2" pageNumber="163">furcum</emphasis>
</taxonomicName>
species-group, comprising 
<taxonomicName id="39C04D1DFFAFE317FD8CFEC842F12E07" ID-CoL="93S8C" baseAuthorityName="Zhu" baseAuthorityYear="1988" box="[631,738,295,321]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="furcum">
<emphasis id="CCB4EA8CFFAFE317FD8CFEC842F12E07" box="[631,738,295,321]" italics="true" pageId="2" pageNumber="163">Z. furcum</emphasis>
</taxonomicName>
and 
<taxonomicName id="39C04D1DFFAFE317FCEDFEC843952E07" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Zhang &amp; Zhang" baseAuthorityYear="2019" box="[790,902,295,321]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="planum">
<emphasis id="CCB4EA8CFFAFE317FCEDFEC843952E07" box="[790,902,295,321]" italics="true" pageId="2" pageNumber="163">Z. planum</emphasis>
</taxonomicName>
; in both species, RTA has a process arising from its mid part, and a pointed tip (
<bibRefCitation id="9A514B6FFFAFE317FE14FEA442F52E23" author="Zhang, B. S. &amp; Zhang, F." box="[495,742,331,357]" pageId="2" pageNumber="163" pagination="67 - 87" refId="ref7994" refString="Zhang, B. S. &amp; Zhang, F. (2019) Three new species of the genus Zodarion (Araneae, Zodariidae) from China. ZooKeys, 813, 67 - 87. https: // doi. org / 10.3897 / zookeys. 813.29683" type="journal article" year="2019">Zhang &amp; Zhang 2019</bibRefCitation>
: figs 4E, 12C).
</paragraph>
<paragraph id="FE7F369EFFAFE317FF3CFE80437A2EEB" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="163">
2) 
<taxonomicName id="39C04D1DFFAFE317FF0CFE80416B2ECF" ID-CoL="5D694" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[247,376,367,393]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis id="CCB4EA8CFFAFE317FF0CFE80416B2ECF" box="[247,376,367,393]" italics="true" pageId="2" pageNumber="163">bactrianum</emphasis>
</taxonomicName>
species-group, comprising 
<taxonomicName id="39C04D1DFFAFE317FD51FE80435B2ECF" ID-CoL="5D694" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[682,840,367,393]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis id="CCB4EA8CFFAFE317FD51FE80435B2ECF" box="[682,840,367,393]" italics="true" pageId="2" pageNumber="163">Z. bactrianum</emphasis>
</taxonomicName>
and 
<taxonomicName id="39C04D1DFFAFE317FC87FE80441F2ECE" authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[892,1036,367,392]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="turanicum" status="sp. nov.">
<emphasis id="CCB4EA8CFFAFE317FC87FE80441F2ECE" box="[892,1036,367,392]" italics="true" pageId="2" pageNumber="163">Z. turanicum</emphasis>
</taxonomicName>
<emphasis id="CCB4EA8CFFAFE317FBE9FE8044432ECF" bold="true" box="[1042,1104,367,393]" pageId="2" pageNumber="163">
<taxonomicNameLabel id="D78757F7FFAFE317FBE9FE8044432ECF" box="[1042,1104,367,393]" pageId="2" pageNumber="163" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; in both species, RTA has two processes originating posteriorly, and an abrupt tip (
<figureCitation id="66FB2A1BFFAFE317FD26FE7C434F2EEB" box="[733,860,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1852,1876]" captionTargetBox="[191,1400,424,1829]" captionTargetId="figure-65@3.[173,1414,412,1829]" captionTargetPageId="3" captionText="FIGURE 2. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B, F), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E). A–E ventral view; F dorsal view of cymbium, showing cymbial diverticulum. Scale bars = 0.2 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branch(es) of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, To—triangular outgrowth, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022243" httpUri="https://zenodo.org/record/7022243/files/figure.png" pageId="2" pageNumber="163">Figs 2B, D</figureCitation>
).
</paragraph>
<paragraph id="FE7F369EFFAFE317FF3CFE5844602EB3" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="163">
3) 
<taxonomicName id="39C04D1DFFAFE317FF0CFE58417A2E96" ID-CoL="7GF4P" baseAuthorityName="Andreeva &amp; Tyshchenko" baseAuthorityYear="1968" box="[247,361,439,464]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="surprisum">
<emphasis id="CCB4EA8CFFAFE317FF0CFE58417A2E96" box="[247,361,439,464]" italics="true" pageId="2" pageNumber="163">surprisum</emphasis>
</taxonomicName>
species-group, comprising the rest of the species with known males; in all species of this group, RTA has one or two processes, and a roundly or almost roundly bent tip (
<figureCitation id="66FB2A1BFFAFE317FC33FE3444752EB3" box="[968,1126,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1852,1876]" captionTargetBox="[191,1400,424,1829]" captionTargetId="figure-65@3.[173,1414,412,1829]" captionTargetPageId="3" captionText="FIGURE 2. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B, F), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E). A–E ventral view; F dorsal view of cymbium, showing cymbial diverticulum. Scale bars = 0.2 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branch(es) of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, To—triangular outgrowth, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022243" httpUri="https://zenodo.org/record/7022243/files/figure.png" pageId="2" pageNumber="163">Figs 2A, C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFAFE317FF3CFE11415A2D7B" pageId="2" pageNumber="163" type="distribution">
<paragraph id="FE7F369EFFAFE317FF3CFE11415A2D7B" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="163">
<emphasis id="CCB4EA8CFFAFE317FF3CFE1141722D5E" bold="true" box="[199,353,510,536]" pageId="2" pageNumber="163">Distribution.</emphasis>
From Volga River’s delta to northeastern 
<collectingCountry id="86D7760EFFAFE317FCC5FE1043972D5F" box="[830,900,511,537]" name="China" pageId="2" pageNumber="163">China</collectingCountry>
, south to northeastern 
<collectingCountry id="86D7760EFFAFE317FB7EFE1044A12D5F" box="[1157,1202,511,537]" name="Iran" pageId="2" pageNumber="163">Iran</collectingCountry>
and south 
<collectingCountry id="86D7760EFFAFE317FAD4FE1045612D5F" box="[1327,1394,511,537]" name="China" pageId="2" pageNumber="163">China</collectingCountry>
(ca 25ºN) (
<figureCitation id="66FB2A1BFFAFE317FF12FDCC412F2D7B" box="[233,316,547,573]" captionStart="FIGURE 12" captionStartId="12.[151,250,1866,1890]" captionTargetBox="[164,1424,453,1842]" captionTargetId="figure-144@12.[164,1424,453,1842]" captionTargetPageId="12" captionText="FIGURE 12. Distribution records of Zodariellum spp. A: 1) Z. asiaticum, 2) Z. bactrianum, 3) Z. bekuzini, 4) Z. continentale, 5) Z. proszynskii, 6) Z. spasskyi and Z. zebra, 7) Z. surprisum, 8) Z. sytchevskajae, 9) Z. turanicum sp. n., 10) Z. turkmenicum sp. n., 11) Z. tadzhikum, 12) Z. testaceofasciatum, 13) Z. volgouralense, 14) Z. nenilini (in part, excluding records from Mongolia). B: 1) Z. nenilini (in part, records from Mongolia), 2) Z. schmidti, 3) Z. mongolicum, 4) Z. chaoyangense, 5) Z. furcum, 6) Z. hunanense, 7) Z. planum. A single point may refer to several closely located records." figureDoi="http://doi.org/10.5281/zenodo.7022273" httpUri="https://zenodo.org/record/7022273/files/figure.png" pageId="2" pageNumber="163">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>