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<document ID-DOI="http://doi.org/10.5281/zenodo.4335214" ID-GBIF-Dataset="6790c5bd-2f80-4161-bb6f-2330f95620ed" ID-GBIF-Taxon="164845041" ID-Zenodo-Dep="4335214" LSID="urn:lsid:zoobank.org:act:A386695B-6001-4E85-971B-964992422A3D" approvalRequired="13" approvalRequired_for_document="1" approvalRequired_for_textStreams="10" approvalRequired_for_treatments="2" checkinTime="1593596629447" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2020" docId="75207B3EB007FFFDA17CF972FEBC1E5A" docLanguage="en" docName="arachnology.18.4.368-386.pdf" docOrigin="Arachnology 18 (4)" docSource="http://dx.doi.org/10.13156/arac.2020.18.4.368" docStyle="DocumentStyle{}" docTitle="Persiscape Zamani &amp; Marusik 2020, gen. n." docType="treatment" docUuid="A386695B-6001-4E85-971B-964992422A3D" docUuidSource="ZooBank" docVersion="4" lastPageId="5" lastPageNumber="373" masterDocId="89190346B003FFF8A101FFBEFF801909" masterDocTitle="A review of Agelenini (Araneae: Agelenidae: Ageleninae) of Iran and Tajikistan, with descriptions of four new genera" masterLastPageNumber="386" masterPageNumber="368" pageId="4" pageNumber="372" updateTime="1632024453243" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A review of Agelenini (Araneae: Agelenidae: Ageleninae) of Iran and Tajikistan, with descriptions of four new genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arachnology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>18</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:extent unit="page">
<mods:start>368</mods:start>
<mods:end>386</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.13156/arac.2020.18.4.368</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.13156/arac.2020.18.4.368</mods:identifier>
<mods:identifier type="GBIF-Dataset">6790c5bd-2f80-4161-bb6f-2330f95620ed</mods:identifier>
<mods:identifier type="Zenodo-Dep">3925217</mods:identifier>
<mods:identifier type="ZooBank">CC0827D2-6390-43DC-A02D-D538246814B9</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4335214" ID-GBIF-Taxon="164845041" ID-Zenodo-Dep="4335214" LSID="urn:lsid:zoobank.org:act:A386695B-6001-4E85-971B-964992422A3D" httpUri="http://treatment.plazi.org/id/75207B3EB007FFFDA17CF972FEBC1E5A" lastPageId="5" lastPageNumber="373" pageId="4" pageNumber="372">
<subSubSection box="[125,323,1740,1764]" pageId="4" pageNumber="372" type="nomenclature">
<paragraph blockId="4.[125,323,1740,1764]" box="[125,323,1740,1764]" pageId="4" pageNumber="372">
<heading bold="true" box="[125,323,1740,1764]" fontSize="10" level="3" pageId="4" pageNumber="372" reason="2">
<emphasis bold="true" box="[125,323,1740,1764]" pageId="4" pageNumber="372">
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[125,242,1740,1763]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[125,242,1740,1763]" italics="true" pageId="4" pageNumber="372">Persiscape</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[248,323,1740,1764]" pageId="4" pageNumber="372" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="372" type="reference_group">
<paragraph blockId="4.[125,760,1803,2100]" pageId="4" pageNumber="372">
<materialsCitation ID-GBIF-Occurrence="2802170317" country="Azerbaijan" location="Azerbaijan" pageId="4" pageNumber="372" specimenCount="1" typeStatus="holotype">
<emphasis box="[155,296,1805,1828]" italics="true" pageId="4" pageNumber="372">
<typeStatus box="[155,205,1805,1828]" pageId="4" pageNumber="372">Type</typeStatus>
species
</emphasis>
:
<taxonomicName authority="Guseinov, Marusik &amp; Koponen, 2005" authorityName="Guseinov, Marusik &amp; Koponen" authorityYear="2005" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="species" species="levyi">
<emphasis box="[317,502,1803,1827]" italics="true" pageId="4" pageNumber="372">Agelescape levyi</emphasis>
<bibRefCitation author="GUSEINOV, E. &amp; MARUSIK, Y. M. &amp; KOPONEN, S." journalOrPublisher="Faunistic review of the funnelweb spiders (Agelenidae) with the description of a new genus and species. Arthropoda Selecta" pageId="4" pageNumber="372" pagination="153 - 177" part="14" refId="ref12216" refString="GUSEINOV, E., MARUSIK, Y. M. &amp; KOPONEN, S. 2005: Spiders (Arachnida: Aranei) ofAzerbaijan 5. Faunistic review of the funnelweb spiders (Agelenidae) with the description of a new genus and species. Arthropoda Selecta 14: 153 - 177." title="Spiders (Arachnida: Aranei) ofAzerbaijan 5" type="journal article" year="2005">Guseinov, Marusik &amp; Koponen, 2005</bibRefCitation>
</taxonomicName>
from
<collectingCountry box="[353,474,1840,1862]" name="Azerbaijan" pageId="4" pageNumber="372">Azerbaijan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="372" type="etymology">
<paragraph blockId="4.[125,760,1803,2100]" pageId="4" pageNumber="372">
<emphasis box="[155,269,1871,1895]" italics="true" pageId="4" pageNumber="372">Etymology</emphasis>
: A combination of
<collectingCountry box="[476,541,1874,1896]" name="Iran" pageId="4" pageNumber="372">Persia</collectingCountry>
and scape, referring to the similarity with
<taxonomicName authorityName="Levy" authorityYear="1996" box="[354,475,1905,1929]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis box="[354,475,1905,1929]" italics="true" pageId="4" pageNumber="372">Agelescape</emphasis>
</taxonomicName>
. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="372" type="diagnosis">
<paragraph blockId="4.[125,760,1803,2100]" lastBlockId="4.[828,1464,1239,2100]" pageId="4" pageNumber="372">
<emphasis box="[155,263,1941,1964]" italics="true" pageId="4" pageNumber="372">Diagnosis</emphasis>
:
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[277,394,1941,1964]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[277,394,1941,1964]" italics="true" pageId="4" pageNumber="372">Persiscape</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[401,474,1943,1963]" pageId="4" pageNumber="372" rank="genus">gen. n.</taxonomicNameLabel>
differ from
<taxonomicName authorityName="Levy" authorityYear="1996" box="[605,726,1939,1963]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis box="[605,726,1939,1963]" italics="true" pageId="4" pageNumber="372">Agelescape</emphasis>
</taxonomicName>
by the number of the teeth on the promargin of the chelicera,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="372" unit="in" value="2.0">2 in</quantity>
the new genus and
<quantity box="[356,422,2010,2032]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="4" pageNumber="372" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">34 in</quantity>
<taxonomicName authorityName="Levy" authorityYear="1996" box="[422,543,2007,2031]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis box="[422,543,2007,2031]" italics="true" pageId="4" pageNumber="372">Agelescape</emphasis>
</taxonomicName>
. Males of this genus differ from those of
<taxonomicName authorityName="Levy" authorityYear="1996" box="[338,459,2041,2065]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis box="[338,459,2041,2065]" italics="true" pageId="4" pageNumber="372">Agelescape</emphasis>
</taxonomicName>
by having a complex embolus with a thumb-like base, a short embolus proper (stick- like portion) (not longer than tegular apophysis), an embolar lamellar process (Pe), conductor twisted around its axis with a thumb-like base and rounded tip (
<taxonomicName box="[1211,1327,1305,1328]" pageId="4" pageNumber="372" rank="variety" variety="embolus">
<emphasis box="[1211,1223,1305,1328]" italics="true" pageId="4" pageNumber="372">v</emphasis>
. embolus
</taxonomicName>
unmodified, filamentous, longer than tegular apophysis, conductor gradually tapering, untwisted, with a sharp tip) and by the shape of the tegular apophysis in lateral view: apophysis straight, longer than wide, with parallel lateral margins and an abrupt tip (
<taxonomicName box="[874,1008,1473,1496]" pageId="4" pageNumber="372" rank="variety" variety="apophysis">
<emphasis box="[874,890,1473,1496]" italics="true" pageId="4" pageNumber="372">v.</emphasis>
apophysis
</taxonomicName>
with diverging margins and rounded tip) (
<figureCitation box="[832,906,1508,1530]" captionStart="Fig" captionStartId="7.[125,156,1762,1781]" captionTargetBox="[140,1447,151,1739]" captionTargetId="figure@7.[140,1447,151,1739]" captionTargetPageId="7" captionText="Fig. 6: SEM micrographs of male palps. A Persiscape gideoni, bulb, ventral view; B P. ecbatana sp. n., bulb, ventral view; C P. levyi, bulb, ventral view; D Agelescape cf. livida, from Israel, bulb, ventral view; E P. ecbatana sp. n., bulb, retrolateral view; F Gorbiscape gorbachevi sp. n., bulb, ventral view; G Agelescape cf. livida, from Israel, whole palp with removed bulb; H same, bulb, anterior view; I Persiscape gideoni, bulb, anterior view. Scale bars = 0.1 mm, if not otherwise indicated. Abbreviations: Be = base of embolus, Cb = base of conductor, Co = conductor, Em = embolus, Pe = embolar process." figureDoi="http://doi.org/10.5281/zenodo.3925229" httpUri="https://zenodo.org/record/3925229/files/figure.png" pageId="4" pageNumber="372">Figs. 6</figureCitation>
AC, E, I, 7BC, 9AF, 10AF, 11AF). Females of the two genera differ by the presence of wide and straight copulatory ducts and the receptacles subdivided into 23 globular chambers in the new genus (
<emphasis box="[1230,1245,1607,1630]" italics="true" pageId="4" pageNumber="372">v.</emphasis>
tube-like, bent, true copulatory ducts and an undivided receptacle in
<taxonomicName authorityName="Levy" authorityYear="1996" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="372">Agelescape</emphasis>
</taxonomicName>
) (
<figureCitation box="[926,1019,1675,1697]" captionStart="Fig" captionStartId="13.[125,156,1465,1484]" captionTargetBox="[125,1450,151,1425]" captionTargetId="figure@13.[125,1463,151,1426]" captionTargetPageId="13" captionText="Fig. 12: Epigynes, different specimens showing variation.A Persiscape levyi, ventral view;B same, dorsal view; C same, ventral view;D same, dorsal view; E same, ventral view; F same, posterior view; G same, dorsal view; H P. gideoni, ventral view; I, D same, posterior views; K, L same, dorsal views; M P. caucasica, ventral view; N same, posterior view; O same, dorsal view. EG, HI, LO after Guseinov, Marusik &amp; Koponen (2005). Scale bars = 0.2 mm, unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3925241" httpUri="https://zenodo.org/record/3925241/files/figure.png" pageId="4" pageNumber="372">Figs. 12</figureCitation>
AO).
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[1089,1206,1674,1697]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[1089,1206,1674,1697]" italics="true" pageId="4" pageNumber="372">Persiscape</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1211,1282,1677,1697]" pageId="4" pageNumber="372" rank="genus">gen. n.</taxonomicNameLabel>
differs from
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Asiascape" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis italics="true" pageId="4" pageNumber="372">Asiascape</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[910,984,1710,1730]" pageId="4" pageNumber="372" rank="genus">gen. n.</taxonomicNameLabel>
by the presence of an embolar process (Pe) (
<taxonomicName box="[837,939,1742,1765]" pageId="4" pageNumber="372" rank="variety" variety="absent">
<emphasis box="[837,852,1742,1765]" italics="true" pageId="4" pageNumber="372">v.</emphasis>
absent
</taxonomicName>
) and globular receptacles subdivided into 23 chambers (
<taxonomicName box="[945,1045,1775,1798]" pageId="4" pageNumber="372" rank="variety" variety="tubular">
<emphasis box="[945,961,1775,1798]" italics="true" pageId="4" pageNumber="372">v.</emphasis>
tubular
</taxonomicName>
receptacles).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="373" pageId="4" pageNumber="372" type="description">
<paragraph blockId="4.[828,1464,1239,2100]" pageId="4" pageNumber="372">
<emphasis box="[858,984,1809,1832]" italics="true" pageId="4" pageNumber="372">Description</emphasis>
: Total length: male 5.308.55, female 6.78 11.20. Carapace length: male 2.504.25, female 2.304.30, width: male 1.603.00, female 1.903.00. Chelicera with 2 prolateral and 3 retrolateral teeth.
</paragraph>
<paragraph blockId="4.[828,1464,1239,2100]" lastBlockId="5.[125,760,877,1876]" lastPageId="5" lastPageNumber="373" pageId="4" pageNumber="372">
Carapace and abdomen with distinct pattern (
<figureCitation box="[1329,1403,1944,1966]" captionStart="Fig" captionStartId="1.[125,156,1514,1533]" captionTargetBox="[138,1457,160,1491]" captionTargetId="figure@1.[125,1463,151,1508]" captionTargetPageId="1" captionText="Fig. 1: A Persiscape caspica sp. n., ♀ habitus; B same, ♂ habitus; C Persilena sengleti sp. n., ♀ habitus; D Asiascape parthica sp. n., ♂ habitus; E Gorbiscape gorbachevi sp. n., ♂ habitus; F same, ♀ habitus." figureDoi="http://doi.org/10.5281/zenodo.3925219" httpUri="https://zenodo.org/record/3925219/files/figure.png" pageId="4" pageNumber="372">Figs. 1</figureCitation>
AB, 2AC, 3CD). Yellowish to reddish-brown, with two wide submarginal bands and a thin gray margin. Sternum grayish with yellow median stripe. Legs usually the same color as carapace, with obscure annulations in some species. Abdomen dark gray with a pattern of lighter spots dorsally, ventrally light gray. Spinnerets yellowish.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3925225" ID-Zenodo-Dep="3925225" httpUri="https://zenodo.org/record/3925225/files/figure.png" pageId="5" pageNumber="373" startId="5.[125,156,721,740]" targetBox="[148,733,152,698]" targetPageId="5">
<paragraph blockId="5.[125,760,717,813]" pageId="5" pageNumber="373">
Fig. 4: Bulb, ventral view.
<emphasis bold="true" box="[348,398,717,742]" pageId="5" pageNumber="373">A, B</emphasis>
<taxonomicName authorityName=": Levy" authorityYear="1996" box="[402,579,721,740]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="livida">
<emphasis box="[402,499,721,740]" italics="true" pageId="5" pageNumber="373">Agelescape</emphasis>
cf.
<emphasis box="[530,579,721,740]" italics="true" pageId="5" pageNumber="373">livida</emphasis>
</taxonomicName>
, from Israel;
<emphasis bold="true" box="[691,710,717,741]" pageId="5" pageNumber="373">C</emphasis>
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Asiascape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="parthica" status="sp. n.">
<emphasis italics="true" pageId="5" pageNumber="373">Asiascape parthica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[318,367,746,765]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
;
<emphasis bold="true" box="[377,396,742,766]" pageId="5" pageNumber="373">D</emphasis>
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[403,605,745,764]" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="gorbachevi" status="sp. n.">
<emphasis box="[403,605,745,764]" italics="true" pageId="5" pageNumber="373">Gorbiscape gorbachevi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[611,660,746,765]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
;
<emphasis bold="true" box="[670,688,742,766]" pageId="5" pageNumber="373">E</emphasis>
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" class="Arachnida" family="Agelenidae" genus="Gorbiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="agelenoides">
<emphasis italics="true" pageId="5" pageNumber="373">G. agelenoides</emphasis>
</taxonomicName>
. A after
<bibRefCitation author="LEVY, G." box="[330,439,769,789]" journalOrPublisher="Zoologica Scripta" pageId="5" pageNumber="373" pagination="85 - 122" part="25" refId="ref12416" refString="LEVY, G. 1996: The agelenid funnel-weaver family and the spider genus Cedicus in Israel (Araneae, Agelenidae and Cybaeidae). Zoologica Scripta 25: 85 - 122." title="The agelenid funnel-weaver family and the spider genus Cedicus in Israel (Araneae, Agelenidae and Cybaeidae)" type="journal article" year="1996">Levy (1996)</bibRefCitation>
, E after
<bibRefCitation author="DE BLAUWE, R." box="[512,668,769,788]" journalOrPublisher="Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="373" pagination="1 - 28" part="52" refId="ref12131" refString="DE BLAUWE, R. 1980: Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie). Bulletin de lIInstitut Royal des Sciences Naturelles de Belgique 52 (11): 1 - 28." title="Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3 e partie)" type="journal article" year="1980">de Blauwe (1980)</bibRefCitation>
, both with modification. Scale bar = 0.2 mm
</paragraph>
</caption>
<paragraph blockId="5.[125,760,877,1876]" pageId="5" pageNumber="373">
Male palp (
<figureCitation box="[274,349,946,968]" captionStart="Fig" captionStartId="7.[125,156,1762,1781]" captionTargetBox="[140,1447,151,1739]" captionTargetId="figure@7.[140,1447,151,1739]" captionTargetPageId="7" captionText="Fig. 6: SEM micrographs of male palps. A Persiscape gideoni, bulb, ventral view; B P. ecbatana sp. n., bulb, ventral view; C P. levyi, bulb, ventral view; D Agelescape cf. livida, from Israel, bulb, ventral view; E P. ecbatana sp. n., bulb, retrolateral view; F Gorbiscape gorbachevi sp. n., bulb, ventral view; G Agelescape cf. livida, from Israel, whole palp with removed bulb; H same, bulb, anterior view; I Persiscape gideoni, bulb, anterior view. Scale bars = 0.1 mm, if not otherwise indicated. Abbreviations: Be = base of embolus, Cb = base of conductor, Co = conductor, Em = embolus, Pe = embolar process." figureDoi="http://doi.org/10.5281/zenodo.3925229" httpUri="https://zenodo.org/record/3925229/files/figure.png" pageId="5" pageNumber="373">Figs. 6</figureCitation>
AC, E, I, 7BC, 9AF, 10AF, 11A F) with one patellar and one tibial apophysis. Tegular apophysis strongly sclerotized, broad, with abrupt or rounded tip in lateral view; conductor complex, with thumblike base and roundly bent lamellate terminal part (Ct); embolus complex, with thumb-like base wider than base of conductor, with a lamellar process and short or stick-like embolus proper.
</paragraph>
<paragraph blockId="5.[125,760,877,1876]" pageId="5" pageNumber="373">
Epigyne (
<figureCitation box="[256,350,1226,1248]" captionStart="Fig" captionStartId="13.[125,156,1465,1484]" captionTargetBox="[125,1450,151,1425]" captionTargetId="figure@13.[125,1463,151,1426]" captionTargetPageId="13" captionText="Fig. 12: Epigynes, different specimens showing variation.A Persiscape levyi, ventral view;B same, dorsal view; C same, ventral view;D same, dorsal view; E same, ventral view; F same, posterior view; G same, dorsal view; H P. gideoni, ventral view; I, D same, posterior views; K, L same, dorsal views; M P. caucasica, ventral view; N same, posterior view; O same, dorsal view. EG, HI, LO after Guseinov, Marusik &amp; Koponen (2005). Scale bars = 0.2 mm, unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3925241" httpUri="https://zenodo.org/record/3925241/files/figure.png" pageId="5" pageNumber="373">Figs. 12</figureCitation>
AO) with distinct fovea (atrium) and subdivided scape that is wider than long; copulatory opening hidden, originating laterad of scape, anterior portion of copulatory duct membranous, as long as wide; other part of copulatory duct straight, broad; receptacle subdivided into several globular chambers.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="373" type="synonymic_list">
<paragraph blockId="5.[125,760,877,1876]" pageId="5" pageNumber="373">
<emphasis box="[155,332,1433,1457]" italics="true" pageId="5" pageNumber="373">Included species</emphasis>
:
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[346,550,1434,1457]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="caspica" status="sp. n.">
<emphasis box="[346,550,1434,1457]" italics="true" pageId="5" pageNumber="373">Persiscape caspica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[557,614,1437,1457]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authority="(Guseinov, Marusik &amp; Koponen, 2005)" authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Guseinov, Marusik &amp; Koponen" baseAuthorityYear="2005" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis box="[627,760,1434,1457]" italics="true" pageId="5" pageNumber="373">P. caucasica</emphasis>
(
<bibRefCitation author="GUSEINOV, E. &amp; MARUSIK, Y. M. &amp; KOPONEN, S." box="[128,545,1470,1492]" journalOrPublisher="Faunistic review of the funnelweb spiders (Agelenidae) with the description of a new genus and species. Arthropoda Selecta" pageId="5" pageNumber="373" pagination="153 - 177" part="14" refId="ref12216" refString="GUSEINOV, E., MARUSIK, Y. M. &amp; KOPONEN, S. 2005: Spiders (Arachnida: Aranei) ofAzerbaijan 5. Faunistic review of the funnelweb spiders (Agelenidae) with the description of a new genus and species. Arthropoda Selecta 14: 153 - 177." title="Spiders (Arachnida: Aranei) ofAzerbaijan 5" type="journal article" year="2005">Guseinov, Marusik &amp; Koponen, 2005</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[562,687,1468,1492]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="ecbatana" status="sp. n.">
<emphasis box="[562,687,1468,1492]" italics="true" pageId="5" pageNumber="373">P. ecbatana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[695,753,1472,1492]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authority="(Levy, 1996)" authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Levy" baseAuthorityYear="1996" box="[125,378,1503,1527]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="gideoni">
<emphasis box="[125,232,1503,1527]" italics="true" pageId="5" pageNumber="373">P. gideoni</emphasis>
(
<bibRefCitation author="LEVY, G." box="[243,374,1505,1526]" journalOrPublisher="Zoologica Scripta" pageId="5" pageNumber="373" pagination="85 - 122" part="25" refId="ref12416" refString="LEVY, G. 1996: The agelenid funnel-weaver family and the spider genus Cedicus in Israel (Araneae, Agelenidae and Cybaeidae). Zoologica Scripta 25: 85 - 122." title="The agelenid funnel-weaver family and the spider genus Cedicus in Israel (Araneae, Agelenidae and Cybaeidae)" type="journal article" year="1996">Levy, 1996</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[392,551,1503,1527]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="nassirkhanii" status="sp. n.">
<emphasis box="[392,551,1503,1527]" italics="true" pageId="5" pageNumber="373">P. nassirkhanii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[558,616,1507,1527]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName authorityName="Guseinov, Marusik &amp; Koponen" authorityYear="2005" box="[630,707,1503,1527]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="levyi">
<emphasis box="[630,707,1503,1527]" italics="true" pageId="5" pageNumber="373">P. levyi</emphasis>
</taxonomicName>
, and
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[125,274,1539,1562]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. n.">
<emphasis box="[125,274,1539,1562]" italics="true" pageId="5" pageNumber="373">P. zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[281,339,1541,1561]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
Based on the shape of the embolus, we consider two species groups in this genus: the
<taxonomicName authorityName="Guseinov, Marusik &amp; Koponen" authorityYear="2005" box="[638,688,1572,1596]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="levyi">
<emphasis box="[638,688,1572,1596]" italics="true" pageId="5" pageNumber="373">levyi</emphasis>
</taxonomicName>
-group and the
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Levy" baseAuthorityYear="1996" box="[210,290,1607,1631]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="gideoni">
<emphasis box="[210,290,1607,1631]" italics="true" pageId="5" pageNumber="373">gideoni</emphasis>
</taxonomicName>
-group. Members of the former have a short spine- or claw-like embolus and include
<taxonomicName authorityName="Guseinov, Marusik &amp; Koponen" authorityYear="2005" box="[622,712,1642,1667]" class="Arachnida" family="Agelenidae" genus="Agelescape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="levyi">
<emphasis box="[622,638,1644,1667]" italics="true" pageId="5" pageNumber="373">P</emphasis>
.
<emphasis box="[662,712,1642,1666]" italics="true" pageId="5" pageNumber="373">levyi</emphasis>
</taxonomicName>
,
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="ecbatana" status="sp. n.">
<emphasis box="[737,753,1644,1667]" italics="true" pageId="5" pageNumber="373">P</emphasis>
.
<emphasis box="[125,223,1677,1701]" italics="true" pageId="5" pageNumber="373">ecbatana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[230,289,1681,1701]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
, and
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[349,460,1679,1702]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="caspica" status="sp. n.">
<emphasis box="[349,365,1679,1702]" italics="true" pageId="5" pageNumber="373">P</emphasis>
.
<emphasis box="[379,460,1679,1702]" italics="true" pageId="5" pageNumber="373">caspica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[469,527,1681,1701]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
, and the latter have a stick-like embolus and include
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2020" baseAuthorityName="Levy" baseAuthorityYear="1996" box="[472,581,1712,1737]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="gideoni">
<emphasis box="[472,581,1712,1737]" italics="true" pageId="5" pageNumber="373">P. gideoni</emphasis>
</taxonomicName>
,
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[598,760,1712,1737]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="nassirkhanii" status="sp. n.">
<emphasis box="[598,760,1712,1737]" italics="true" pageId="5" pageNumber="373">P. nassirkhanii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[125,182,1751,1771]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
, and
<taxonomicName authority="Zamani &amp; Marusik, 2020" authorityName="Zamani &amp; Marusik" authorityYear="2020" box="[240,390,1748,1772]" class="Arachnida" family="Agelenidae" genus="Persiscape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="373" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. n.">
<emphasis box="[240,390,1748,1772]" italics="true" pageId="5" pageNumber="373">P. zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[397,454,1751,1771]" pageId="5" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="373" type="distribution">
<paragraph blockId="5.[125,760,877,1876]" pageId="5" pageNumber="373">
<emphasis box="[155,285,1782,1806]" italics="true" pageId="5" pageNumber="373">Distribution</emphasis>
: The genus is known from Lesbos, Central
<collectingCountry box="[125,219,1819,1841]" name="Turkey" pageId="5" pageNumber="373">Anatolia</collectingCountry>
, and
<collectingCountry box="[280,339,1820,1841]" name="Israel" pageId="5" pageNumber="373">Israel</collectingCountry>
to
<collectingCountry box="[379,422,1821,1841]" name="Iran" pageId="5" pageNumber="373">Iran</collectingCountry>
(western, northern, and southeastern) (
<figureCitation box="[220,306,1854,1876]" captionStart="Fig" captionStartId="16.[125,156,1563,1582]" captionTargetBox="[126,1462,155,1538]" captionTargetId="figure@16.[126,1462,155,1538]" captionTargetPageId="16" captionText="Fig. 15: Distribution maps. A Agelescape afFnis, A. livida, and Persiscape caucasica; B P. caspica sp. n., P. ecbatana sp. n., P. nassirkhanii sp. n., P. levyi, P. zagrosensis sp. n., Asiascape parthica sp. n., Persilena sengleti sp. n., and Gorbiscape gorbachevi sp. n.; C Persiscape gideoni." figureDoi="http://doi.org/10.5281/zenodo.3925248" httpUri="https://zenodo.org/record/3925248/files/figure.png" pageId="5" pageNumber="373">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>