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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="75080B2F5CE955999C9274B037E542BE" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Lachesana kavirensis Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1035</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>193</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
<mods:identifier type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181753555" LSID="urn:lsid:zoobank.org:act:3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" httpUri="http://treatment.plazi.org/id/75080B2F5CE955999C9274B037E542BE" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/3BC489D8-1BD7-46C1-A4D2-BCB9A3BCDE58" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis" status="sp. nov.">Lachesana kavirensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">Figs 1A-C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">, 3A-E</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">, 4I</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Qom Province</emphasis>
: Dasht-e Masileh,
<geoCoordinate degrees="34" direction="north" minutes="47" orientation="latitude" precision="925" value="34.783333">34°47'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="11" orientation="longitude" precision="925" value="51.183334">51°11'E</geoCoordinate>
, 11.2017 (M. Mirghazanfari).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">
The specific epithet is derived from
<normalizedToken originalValue="Kavir">'Kavir'</normalizedToken>
, a Persian word for desert, referring to the occurrence of the species in the central deserts of Iran.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
The male of
<taxonomicName lsidName="L. kavirensis" pageId="0" pageNumber="145" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">L. kavirensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from the similar
<taxonomicName lsidName="L. dyachkovi" pageId="0" pageNumber="145" rank="species" species="dyachkovi">
<emphasis italics="true" pageId="0" pageNumber="145">L. dyachkovi</emphasis>
</taxonomicName>
Fomichev &amp; Marusik, 2019 by having a gently bent cheliceral fang (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
bent over 90°, cf. Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
and fig. 15 in
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.08" author="Fomichev, AA" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="145" pagination="556 - 561" refId="B5" refString="Fomichev, AA, Marusik, YM, 2019. A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan. Arthropoda Selecta 28 (4): 556 - 561, DOI: https://doi.org/10.15298/arthsel.28.4.08" title="A new species of Lachesana Strand, 1932 (Aranei: Zodariidae) from southern Kazakhstan." url="https://doi.org/10.15298/arthsel.28.4.08" volume="28" year="2019">Fomichev and Marusik 2019</bibRefCitation>
). The shape of the RTA is also different from other species of the region (stalk with a gentle curve ventrally and tip straightly pointed; cf. Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
and Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4F-H, J</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
Habitus and chelicerae of male
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Lachesana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachesana kavirensis" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="kavirensis">
<emphasis italics="true" pageId="0" pageNumber="145">Lachesana kavirensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="L. perseus" pageId="0" pageNumber="145" rank="species" species="perseus">
<emphasis italics="true" pageId="0" pageNumber="145">L. perseus</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, B, D, E</emphasis>
habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
chelicerae, ventral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1A, B</figureCitation>
. Total length 11.55. Carapace 5.80 long, 2.60 wide at pars cephalica, 3.75 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.23, ALE: 0.20, PME: 0.17, PLE: 0.23, PME-PME: 0.14. Carapace, sternum, labium, chelicerae and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and chelicerae of male Lachesana kavirensis sp. nov. (A-C) and L. perseus sp. nov. (D-F) A, B, D, E habitus, dorsal C, F chelicerae, ventral." figureDoi="10.3897/zookeys.1035.65767.figure1" httpUri="https://binary.pensoft.net/fig/536878" pageId="0" pageNumber="145">1C</figureCitation>
) with dense, black setae and one promarginal tooth; fangs gently curved, almost straight. Legs the same color as carapace, without annulations and with numerous spines. Abdomen pale, dorsally with a longitudinal dark stripe and several small spots. Spinnerets slightly lighter than abdomen, uniform in color. Measurements of legs: I: 15.49 (4.47, 1.88, 3.33, 3.29, 2.52), II: 15.03 (4.20, 1.79, 3.02, 3.50, 2.52), III: 15.78 (4.08, 1.96, 2.48, 4.71, 2.55), IV: 18.30 (4.58, 2.07, 3.20, 5.24, 3.21).
</paragraph>
<paragraph pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palps of Lachesana kavirensis sp. nov. (A-E) and L. perseus sp. nov. (F-J) A, B, G, F retrolateral C, H prolateral D, I ventral E, J dorsal." figureDoi="10.3897/zookeys.1035.65767.figure3" httpUri="https://binary.pensoft.net/fig/536880" pageId="0" pageNumber="145">3A-E</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male palps of Lachesana dyachkovi (A-C, F), L. tarabaevi (D, E, G), L. blackwalli (H), L. kavirensis sp. nov. (I) and L. perseus sp. nov. (J) A, D ventral B, C, E dorsal, prolateral and retrolateral F-J retrolateral tibial apophysis A-G reproduced after Fomichev and Marusik (2019) H illustrated after Oezkuetuek et al. (2020). Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure4" httpUri="https://binary.pensoft.net/fig/536881" pageId="0" pageNumber="145">4I</figureCitation>
. RTA long, over 1.2 times longer than tibia, its widest part less than 2 times wider than stalk; cymbium with 2 spines in mesal proximal part; bulb elongate, 1.3 times longer than wide; base of embolus large, 1/3 of the tegulum's length; embolus filamentous, lying in chute formed by conductor.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Known only from the type locality in Qom Province, northern Iran (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Distribution records of Acanthinozodium spp. (partim 1 - 4, 11), Lachesana spp. (5 - 7), Parazodarion raddei (8), Pax spp. (9, 10) and Trygetus spp. (11, 12) in Iran and Turkmenistan: 1 A. atrisa sp. nov. 2 A. niusha sp. nov. 3 A. parysatis sp. nov. 4 A. sorani sp. nov. 5 L. insensibilis 6 L. kavirensis sp. nov. 7 L. perseus sp. nov. 8 P. raddei 9 P. ellipita sp. nov. 10 P. leila sp. nov. 11 A. ovtchinnikovi sp. nov., T. gromovi 12 T. susianus sp. nov." figureDoi="10.3897/zookeys.1035.65767.figure32" httpUri="https://binary.pensoft.net/fig/536909" pageId="0" pageNumber="145">32</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>