193 lines
22 KiB
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193 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.841.32364" ID-GBIF-Dataset="ac2013d2-401a-4317-b2c0-e4efdaed4341" ID-PMC="PMC6491414" ID-Pensoft-Pub="1313-2970-841-1" ID-PubMed="31097912" ID-ZBK="4D023A1CB64C42788C8AF23D55266E2F" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-841-1" ModsDocOrigin="ZooKeys 841" ModsDocTitle="Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species" checkinTime="1556044701643" checkinUser="pensoft" docAuthor="van der Wal, Serita, Smit, Nico J. & Hadfield, Kerry A." docDate="2019" docId="6500A2B282AC6F837AA0A85591159B18" docLanguage="en" docName="ZooKeys 841: 1-37" docOrigin="ZooKeys 841" docSource="http://dx.doi.org/10.3897/zookeys.841.32364" docTitle="Elthusa xena Wal, Smit & Hadfield, 2019, sp. n." docType="treatment" docUuid="338A44A2-746F-4D9B-B890-5372D1E45B4C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FB3006256C0DFFA4FFE9FFDDD8202656" masterDocTitle="Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species" masterLastPageNumber="37" masterPageNumber="1" pageNumber="1" updateTime="1668167181931" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the fish parasitic genus Elthusa Schioedte & Meinert, 1884 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>van der Wal, Serita</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smit, Nico J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hadfield, Kerry A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>841</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>37</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.841.32364</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.841.32364</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-841-1</mods:identifier>
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<mods:identifier type="ZBK">4D023A1CB64C42788C8AF23D55266E2F</mods:identifier>
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<mods:identifier type="ZooBank">4D023A1CB64C42788C8AF23D55266E2F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156828328" LSID="urn:lsid:zoobank.org:act:338A44A2-746F-4D9B-B890-5372D1E45B4C" httpUri="http://treatment.plazi.org/id/6500A2B282AC6F837AA0A85591159B18" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/338A44A2-746F-4D9B-B890-5372D1E45B4C" class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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Figures 4, 5, 6, 7, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype. SOUTH AFRICA • 1 ♀ (ovigerous, 34.0 mm TL, 17.0 mm W); Alexander Bay, mouth of the Orange River;
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.633333">28°38'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="16.45">16°27'E</geoCoordinate>
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; July 1993; coll. J Laubscher; from the super klipfish,
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Clinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clinus superciliosus" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="superciliosus">Clinus superciliosus</taxonomicName>
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(Linnaeus, 1758); SAMC-A089958.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Paratype. SOUTH AFRICA • 1 ♂ (intermoult, 8.0 mm TL, 4.0 mm W); same data as holotype; SAMC-A089959.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description</paragraph>
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<paragraph pageId="0" pageNumber="1">(ovigerous ♀). Figs 4-5. Body slightly twisted to the left, elongated ovoid, twice as long as greatest width; dorsal surfaces smooth and polished in appearance, widest at pereonite 5, most narrow at pereonite 1, pereonite lateral margins mostly rounded, medially indented. Cephalon 0.8 times longer than wide, visible from dorsal view, sub-triangular with blunt anterior point. Frontal margin thickened, ventrally folded. Eyes oval with distinct margins; one eye 0.1 times width of cephalon, 0.3 times length of cephalon. Pereonite 1 smooth, anterior border slightly concave; anterolateral angle rounded, extending to the medial region of eyes. Posterior margins of pereonites smooth, slightly curved laterally. Coxae 2-3 narrow with posteroventral angles narrowly rounded; coxae 4-7 with rounded point, not extending past pereonite margin. Pereonites 2-5 subequal, pereonites 6 and 7 slightly narrower. Pleon 0.4 times as long as total body length, with pleonite 1 same width as other pleonites, lateral margins concealed by pereonite 7, slightly visible in dorsal view; pleonites posterior margin smooth, slightly curved laterally. Pleonite 2 partially overlapped by pereonite 7; posterolateral angles of pleonite 2 rounded. Pleonites 3-5 similar in form to pleonite 2; pleonite 5 longest, free, not overlapped by lateral margins of pleonite 4, with posterolateral angles narrowly rounded, posterior margin with 3 indentations. Pleotelson 0.6 times as long as anterior width, dorsal surface smooth; lateral margins convex; posterior margin evenly rounded.</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 4.
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. holotype ♀ (ovigerous, 34.0 mm TL, 17.0 mm W) (SAMC-A089958) from Alexander Bay, South Africa A dorsal body B lateral body C oostegites D dorsal view of cephalon and pereonite 1 E uropod F ventral cephalon G pleopod 1 H dorsal view of pleon I pereopod 1 J pereopod 7.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 5. Photos of
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. holotype ♀ (ovigerous, 34.0 mm TL, 17.0 mm W) (SAMC-A089958) from Alexander Bay, South Africa A dorsal view B ventral view C lateral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">Antennula shorter than antenna, consisting of eight articles; peduncle articles I and II distinct and articulated, extending to anterior of pereonite 1. Antenna consists of eleven articles, extending to past anterior margin of pereonite 1.</paragraph>
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<paragraph pageId="0" pageNumber="1">Pereopod 1 basis 1.8 times as long as greatest width; ischium 0.7 times as long as basis; merus proximal margin without bulbous protrusion; carpus with rounded proximal margin; propodus 1.8 times as long as wide; dactylus slender, 0.8 times as long as propodus, 2.3 times as long as basal width. Pereopods 2-3 similar to pereopod 1, all pereopods without robust or simple setae. Pereopod 7 basis with carina, 1.5 times as long as greatest width; ischium without protrusions, 0.9 times as long as basis; merus proximal margin with slight bulbous protrusion, 0.6 times as long as wide, 0.3 times as long as ischium; carpus with bulbous protrusion, 0.9 times as long as wide, 0.5 times as long as ischium; propodus as long as wide, 0.4 times as long as ischium; dactylus slender, 1.9 times as long as propodus, 3.1 times as long as basal width.</paragraph>
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<paragraph pageId="0" pageNumber="1">Pleopods simple, exopod larger than endopod. Pleopod 1 exopod 1.1 times as long as wide, lateral margin strongly convex, distally broadly rounded, mesial margin weakly convex; peduncle 2.8 times as wide as long.</paragraph>
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<paragraph pageId="0" pageNumber="1">Uropod more than half the length of pleotelson, peduncle 0.8 times longer than rami, peduncle lateral margin without setae; rami not extending beyond pleotelson, apices narrowly rounded. Endopod apically rounded, 2.5 times as long as greatest width, lateral margin weakly convex, mesial margin straight, terminating without setae. Exopod extending beyond end of endopod, twice as long as greatest width, apically rounded, lateral margin weakly convex, mesial margin straight, terminating without setae.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description</paragraph>
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<paragraph pageId="0" pageNumber="1">(paratype intermoult ♂). Figs 6, 7. Male similar to female but smaller. Specimen mid-moult. Body rectangular, not twisted, twice as long as greatest width. Pereonite lateral margins mostly subparallel. Cephalon 0.7 times longer than wide. Frontal margin rounded to form blunt rostrum. Eyes oval with distinct margins; one eye 0.2 times width of cephalon; 0.5 times length of cephalon. Pereonite 1 smooth, anterior border concave, extending past base of cephalon. Posterior margins of pereonites smooth and straight, except pereonite 4 and 5. Coxae 2-3 wide, with posteroventral angles rounded; coxae 4-7 rounded. Pereonites 6 and 7 narrower, becoming more progressively rounded posteriorly. Pleon 0.3 times as long as total body length, with pleonite 1 largely concealed by pereonite 7, slightly visible in dorsal view; pleonites 1-3 posterior margin posteriorly concave, smooth and slightly curved laterally. Pleonite 5 overlapped by lateral margins of pleonite 4, with posterolateral angles narrowly rounded, posterior margin straight. Pleotelson 0.8 times as long as anterior width, lateral margins straight or weakly convex, posterior margin broadly truncate.</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 6.
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. paratype ♂ (intermoult) (8 mm TL, 4 mm W) (SAMC-A089959) from Alexander Bay, South Africa A dorsal body B lateral body C pereopod 1 D pereopod 7 E dorsal view of cephalon F penes G uropod H ventral cephalon I dorsal view of pleon J ventral pleopod 2 K dorsal pleopod 2.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 7. Photos of
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. paratype ♂ (intermoult) (8.0 mm TL, 4.0 mm W) (SAMC-A089959) from Alexander Bay, South Africa A dorsal view B ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">Antennula shorter than antenna, consisting of eight articles. Antenna consists of ten articles, extending to middle of pereonite 1.</paragraph>
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<paragraph pageId="0" pageNumber="1">Pereopod 1 basis twice as long as greatest width; ischium 0.6 times as long as basis; propodus 1.6 times as long as wide; dactylus 1.1 times as long as propodus, 3 times as long as basal width. Pereopod 7 twice as long as greatest width; ischium 0.7 times as long as basis; merus proximal margin without bulbous protrusion, 0.7 times as long as wide, 0.4 times as long as ischium; carpus without bulbous protrusion, 0.7 times as long as wide, 0.4 times as long as ischium; propodus 1.3 times as long as wide, 0.6 as long as ischium; dactylus slender, 1.4 times as long as propodus, 2.7 times as long as basal width.</paragraph>
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<paragraph pageId="0" pageNumber="1">Pleopod 1 exopod 1.2 times as long as wide, lateral margin weakly convex, distally broadly rounded, mesial margin straight; endopod 2.1 times as long as wide, lateral margin weakly convex, mesial margin straight, peduncle 2.2 times as wide as long. Pleopod 2 appendix masculina with parallel margins, 1.1 times as long as endopod, distally narrowly rounded.</paragraph>
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<paragraph pageId="0" pageNumber="1">Uropod same length or slightly longer than the pleotelson, peduncle 0.4 times longer than rami, rami extending slightly beyond pleotelson, apices narrowly rounded. Endopod apically slightly pointed, 3 times as long as greatest width. Exopod 2.6 times as long as greatest width.</paragraph>
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<paragraph pageId="0" pageNumber="1">Penes medially adjacent; penial process 0.7 times as long as basal width.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The epithet is constructed in a possessive form of a personal name. This species is named after Xena, the warrior princess, in reference to the strong nature of the female cymothoid isopod.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Size.</paragraph>
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<paragraph pageId="0" pageNumber="1">Ovigerous female (34.0 mm TL, 17.0 mm W), male (8.0 mm TL, 4.0 mm W).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Currently only known from the mouth of the Orange River, Alexander Bay, South Africa (Atlantic Ocean).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="host">
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<paragraph pageId="0" pageNumber="1">Hosts.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Clinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clinus superciliosus" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="superciliosus">Clinus superciliosus</taxonomicName>
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(Linnaeus, 1758). This is the first record of a klipfish (of the genus
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Clinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clinus" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">Clinus</taxonomicName>
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Cuvier, 1816), and of the intertidal super klipfish,
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Clinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clinus supercilious" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="supercilious">Clinus supercilious</taxonomicName>
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, as a fish host of a species of
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Elthusa</taxonomicName>
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. This host belongs to the fish order
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<taxonomicName lsidName="Cymothooidea" order="Perciformes" pageId="0" pageNumber="1" rank="order">Perciformes</taxonomicName>
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, and is endemic to the Southeast Atlantic Ocean, from northern Namibia to the Kei River of South Africa (
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<bibRefCitation author="Smith, MM" journalOrPublisher="Springer, South Africa" pageId="0" pageNumber="1" title="Smith's sea fishes." year="1986">Smith and Heemstra 1986</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. female can be identified by the elongate, ovoid body shape; coxae 7 that do not extend past the posterior margin of pereonite 7; a bluntly pointed anterior margin of the cephalon; evenly rounded, slightly concave anterior margin of pereonite 1; uropod rami with apices narrowly rounded and more than half the length of pleotelson; pleonite 5 posterior margin with indentations; and the pleotelson is short, roughly quadrate, with margins that curl upward.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Two other
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Elthusa</taxonomicName>
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species have been recorded from related perciform fish hosts from the family
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<taxonomicName family="Clinidae" lsidName="" pageId="0" pageNumber="1" rank="family">Clinidae</taxonomicName>
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Swainson, 1839 (blennies).
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa californica" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="californica">Elthusa californica</taxonomicName>
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(Schioedte & Meinert, 1884) was noted from the striped kelpfish
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Gibbonsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbonsia metzi" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="metzi">Gibbonsia metzi</taxonomicName>
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Hubbs, 1927; and
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa menziesi" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="menziesi">Elthusa menziesi</taxonomicName>
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(Brusca, 1981) from both the spotted kelpfish
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Gibbonsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbonsia elegans" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="elegans">Gibbonsia elegans</taxonomicName>
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(Cooper, 1864) and the crevice kelpfish
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Gibbonsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbonsia montereyensis" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="montereyensis">Gibbonsia montereyensis</taxonomicName>
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Hubbs, 1927. However, this is the first record of
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Elthusa</taxonomicName>
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collected from a
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<taxonomicName class="Actinopterygii" family="Clinidae" genus="Clinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clinus" order="Perciformes" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">Clinus</taxonomicName>
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sp.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. can be distinguished from
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<taxonomicName lsidName="E. raynaudii" pageId="0" pageNumber="1" rank="species" species="raynaudii">E. raynaudii</taxonomicName>
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by having a bluntly pointed cephalon anterior margin, compared to the narrowly truncate margin of
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<taxonomicName lsidName="E. raynaudii" pageId="0" pageNumber="1" rank="species" species="raynaudii">E. raynaudii</taxonomicName>
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. Other differences include the shape of the pleotelson (which is quadrate, wide and short for
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<taxonomicName lsidName="E. xena" pageId="0" pageNumber="1" rank="species" species="xena">E. xena</taxonomicName>
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sp. n., and evenly rounded for
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<taxonomicName lsidName="E. raynaudii" pageId="0" pageNumber="1" rank="species" species="raynaudii">E. raynaudii</taxonomicName>
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); pleonite 1 is the same length as the other pleonites in
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<taxonomicName class="Malacostraca" family="Cymothoidae" genus="Elthusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elthusa xena" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="xena">Elthusa xena</taxonomicName>
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sp. n. but narrower in
|
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<taxonomicName lsidName="E. raynaudii" pageId="0" pageNumber="1" rank="species" species="raynaudii">E. raynaudii</taxonomicName>
|
|
; and the uropod apices of
|
|
<taxonomicName lsidName="E. xena" pageId="0" pageNumber="1" rank="species" species="xena">E. xena</taxonomicName>
|
|
sp. n. are narrowly rounded compared to the broadly rounded apices of
|
|
<taxonomicName lsidName="E. raynaudii" pageId="0" pageNumber="1" rank="species" species="raynaudii">E. raynaudii</taxonomicName>
|
|
uropods. See Table 1 for further morphological variation and comparisons.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |