241 lines
28 KiB
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241 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="2FE0B42C907B557494E84C289EFD1EE0" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole tampony Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="05966990-3AEA-4921-888E-941D151DFCC1" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090192" LSID="urn:lsid:zoobank.org:act:05966990-3AEA-4921-888E-941D151DFCC1" httpUri="http://treatment.plazi.org/id/2FE0B42C907B557494E84C289EFD1EE0" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="2FE0B42C-907B-5574-94E8-4C289EFD1EE0" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony" status="sp. nov.">Pheidole tampony</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">Figs 56A-F</figureCitation>
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<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">, 64N</figureCitation>
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<figureCitation captionStart="Figure 67" captionStartId="F67" captionText="Figure 67. Distribution. Pheidole renirano sp. nov. (A). Pheidole sava sp. nov. (B). Pheidole sikorae Forel (C). Pheidole sofia sp. nov. (D). Pheidole sparsa sp. nov. (E). Pheidole tampony sp. nov. (F). Pheidole trichotos sp. nov. (G). Pheidole tsaravoniana sp. nov. (H). Pheidole vadum sp. nov. (I). Pheidole veteratrix Forel (J). Pheidole volontany sp. nov. (K). Pheidole vony sp. nov. (L)." figureDoi="10.3897/zookeys.949.51269.figure67" httpUri="https://binary.pensoft.net/fig/432236" pageId="0" pageNumber="1">, 67F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Sava Region: Parc National de Marojejy, near summit, 25.4 km 20.1°NE Andapa;
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<geoCoordinate degrees="14.44918" direction="south" orientation="latitude" precision="1" value="-14.44918">-14.44918</geoCoordinate>
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,
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<geoCoordinate degrees="49.73243" direction="east" orientation="longitude" precision="1" value="49.73243">49.73243</geoCoordinate>
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; alt. 2100 m; 10 Feb 2018; B. L. Fisher et al. leg.; montane shrubland, under rootmat, on stone; BLF40950; CASENT0809550 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0923285 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tampony</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923285) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0809550).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval and not widening posteriorly, with anterior and posterior sides slightly convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with thick, interrupted, dense, and longitudinal rugae with smooth to indistinctly rugulate interspaces; occipital lobes predominantly smooth, only anterior part with indistinct longitudinal rugae; denser and thinner longitudinal rugae with rugulate interspaces; area posterolateral from eyes with reduced sculpture and predominantly smooth; scape, when laid back, exceeding the midlength of head by approximately two-fifths of its length; inner hypostomal teeth distinct, distinct, moderately large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower than and approximately as high as inner teeth, apex directed outward; inner and outer hypostomal teeth closely spaced and not connected by concavity; mesosoma rugofoveolate; pronotum with sparse foveolae, its dorsum with smooth notch; katepisternum with large, smooth notch; body brown.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate; median frons with short and indistinct longitudinal rugulae; vertex with fading sculpture; area posterolateral from eyes smooth; scape, when laid back, exceeding the posterior head margin by one-third of its length; promesonotum low and moderately long; promesonotal groove present; propodeal spines small and triangular; mesosoma foveolate; body bright brown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 1.36; HW: 1.36; SL: 0.87; EL: 0.19; WL: 1.34; PSL: 0.23; MTL: 0.89; PNW: 0.5; PTW: 0.18; PPW: 0.34; CI: 100.1; SI: 64.0; PSLI: 17.0; PPI: 52.1; PNI: 36.6; MTI: 65.4.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, not widening posteriorly, with anterior and posterior sides slightly convex (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, long, erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, interrupted, dense, and longitudinal rugae with smooth to indistinctly rugulate interspaces; lateral sides of frons with dense, thick, and longitudinal rugae, interspaces with dense rugulae. Occipital lobes predominantly smooth, only anterior part with indistinct longitudinal rugae. Area posterolateral from eyes with denser and thinner longitudinal rugae with rugulate interspaces, posteriormost parts with reduced sculpture and predominantly smooth. Gena with relatively sparse, thick, and longitudinal rugae with distinctly rugulate interspaces. Centre of clypeus indistinctly foveolate and having shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderately large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower than and approximately as high as inner teeth, apex directed outward; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 64" captionStartId="F64" captionText="Figure 64. Major worker, hypostomal teeth. Pheidole masoandro sp. nov. (A). Pheidole mavohavoana sp. nov. (B). Pheidole midongy sp. nov. (C). Pheidole mikros sp. nov. (D). Pheidole mivory sp. nov. (E). Pheidole nitidobruna sp. nov. (F). Pheidole parvula sp. nov. (G). Pheidole parvulogibba sp. nov. (H). Pheidole renirano sp. nov. (I). Pheidole sava sp. nov. (J). Pheidole sikorae Forel (K). Pheidole sofia sp. nov. (L). Pheidole sparsa sp. nov. (M). Pheidole tampony sp. nov. (N). Pheidole trichotos sp. nov. (O). Pheidole tsaravoniana sp. nov. (P). Pheidole vadum sp. nov. (Q). Pheidole veteratrix Forel (R). Pheidole volontany sp. nov. (S). Pheidole vony sp. nov. (T)." figureDoi="10.3897/zookeys.949.51269.figure64" httpUri="https://binary.pensoft.net/fig/432233" pageId="0" pageNumber="1">64N</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, with narrow base and acute apex; humeral area produced (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D</figureCitation>
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). Surface shiny and rugofoveolate; pronotum with sparse foveolae, its dorsum with smooth notch; katepisternum with large, smooth notch. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Brown, gaster slightly darker (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56D, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.68; HW: 0.57; SL: 0.81; EL: 0.13; WL: 0.93; PSL: 0.12; MTL: 0.65; PNW: 0.4; PTW: 0.09; PPW: 0.16; CI: 119.5; SI: 141.6; PSLI: 17.0; PPI: 54.7; PNI: 69.6; MTI: 114.2.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin slightly convex (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56A</figureCitation>
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). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and foveolate; median frons with short and indistinct longitudinal rugulae; vertex with fading sculpture; area posterolateral from eyes smooth; antennal sockets with few indistinct, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-third of its length; pilosity dense, subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56A, C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum low and moderately long, arched; promesonotal groove indistinct; metanotal groove distinct; propodeal spines small and triangular (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56C</figureCitation>
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). Sculpture shiny and foveolate. Pilosity very sparse, moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Bright brown (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Pheidole tampony sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923285) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0809550)." figureDoi="10.3897/zookeys.949.51269.figure56" httpUri="https://binary.pensoft.net/fig/432225" pageId="0" pageNumber="1">56C, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Malagasy for summit in reference to the type locality located close to the mountain peak.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
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<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species was collected at 2100 m in elevation, in montane shrubland. Nest was located under rootmat on stone.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tampony</emphasis>
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</taxonomicName>
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sp. nov. is a member of a group of species characterised by distinctly reduced head sculpture in major workers with occipital lobes entirely or predominantly smooth, area posterolateral from eyes partially or entirely smooth and shiny or with reduced sculpture and smooth notches. The group consists of four species:
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<taxonomicName lsidName="P. litigiosa" pageId="0" pageNumber="1" rank="species" species="litigiosa">
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<emphasis italics="true" pageId="0" pageNumber="1">P. litigiosa</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="P. masoandro" pageId="0" pageNumber="1" rank="species" species="masoandro">
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<emphasis italics="true" pageId="0" pageNumber="1">P. masoandro</emphasis>
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</taxonomicName>
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sp. nov.,
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<taxonomicName lsidName="P. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
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<emphasis italics="true" pageId="0" pageNumber="1">P. gracilis</emphasis>
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</taxonomicName>
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sp. nov., and
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<taxonomicName lsidName="P. tampony" pageId="0" pageNumber="1" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">P. tampony</emphasis>
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</taxonomicName>
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sp. nov.
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tampony</emphasis>
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</taxonomicName>
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|
sp. nov. is known only from Parc National de Marojejy in Antsiranana and its distribution does not overlap with other members of this group. Both minor and major workers of this species can be easily separated from other members of this group based on darker, bright brown to brown body colouration. However, major workers of
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<taxonomicName lsidName="P. tampony" pageId="0" pageNumber="1" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">P. tampony</emphasis>
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</taxonomicName>
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sp. nov. can be confused with majors of sympatric
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|
<taxonomicName lsidName="P. manantenina" pageId="0" pageNumber="1" rank="species" species="manantenina">
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<emphasis italics="true" pageId="0" pageNumber="1">P. manantenina</emphasis>
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|
</taxonomicName>
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sp. nov.
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tampony" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tampony">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole tampony</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be separated based on presence of distinct smooth notch on posteriormost part of area posterolateral from eyes and partially smooth occipital lobes (
|
|
<taxonomicName lsidName="P. manantenina" pageId="0" pageNumber="1" rank="species" species="manantenina">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. manantenina</emphasis>
|
|
</taxonomicName>
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|
sp. nov. has posteriormost part of area posterolateral from eyes and occipital lobes with sparse and reduced sculpture that are never smooth), lower promesonotum with more distinctly developed mesonotal process (
|
|
<taxonomicName lsidName="P. manantenina" pageId="0" pageNumber="1" rank="species" species="manantenina">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. manantenina</emphasis>
|
|
</taxonomicName>
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|
sp. nov. has promesonotum higher and lacking mesonotal process), and smooth notch on katepisternum (
|
|
<taxonomicName lsidName="P. manantenina" pageId="0" pageNumber="1" rank="species" species="manantenina">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. manantenina</emphasis>
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|
</taxonomicName>
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|
sp. nov. has katepisternum entirely sculptured). Both taxa distinctly differ in minor workers:
|
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<taxonomicName lsidName="P. tampony" pageId="0" pageNumber="1" rank="species" species="tampony">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. tampony</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. has minors with predominantly foveolate head and mesosoma and long and low promesonotum, while minors of
|
|
<taxonomicName lsidName="P. manantenina" pageId="0" pageNumber="1" rank="species" species="manantenina">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. manantenina</emphasis>
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</taxonomicName>
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|
sp. nov. have head and mesosoma predominantly smooth and shorter and higher promesonotum.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |