treatments-xml/data/0E/4D/87/0E4D87D0FFE8FFB32FEC083FFE4D8215.xml
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<document id="FCF0C3F7747267C97CCCDB9C7ED486BE" ID-CLB-Dataset="173353" ID-DOI="10.11646/phytotaxa.570.1.3" ID-GBIF-Dataset="5948903b-2326-4611-bd09-2d6314cfe677" ID-ISSN="1179-3163" ID-Zenodo-Dep="7251466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666767854836" checkinUser="plazi" docAuthor="Herman, Paul P. J." docDate="2022" docId="0E4D87D0FFE8FFB32FEC083FFE4D8215" docLanguage="en" docName="phytotaxa.570.1.3.pdf" docOrigin="Phytotaxa 570 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.570.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Hilliardiella oligocephala var. oligocephala var. oligocephala" docType="treatment" docVersion="5" lastPageNumber="37" masterDocId="F274FFA8FFECFFBB2F640D77FFD68A1D" masterDocTitle="Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety" masterLastPageNumber="42" masterPageNumber="29" pageNumber="33" updateTime="1699409107501" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="676185198B7E669ADAF8F6B58F463A41">Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety</mods:title>
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<treatment id="0E4D87D0FFE8FFB32FEC083FFE4D8215" ID-DOI="http://doi.org/10.5281/zenodo.7258181" ID-GBIF-Taxon="203687366" ID-Zenodo-Dep="7258181" LSID="urn:lsid:plazi:treatment:0E4D87D0FFE8FFB32FEC083FFE4D8215" httpUri="http://treatment.plazi.org/id/0E4D87D0FFE8FFB32FEC083FFE4D8215" lastPageId="8" lastPageNumber="37" pageId="4" pageNumber="33">
<subSubSection id="CEFE654DFFE8FFBF2FEC083FFDAA8F7E" box="[136,636,1352,1379]" pageId="4" pageNumber="33" type="nomenclature">
<paragraph id="865B36C6FFE8FFBF2FEC083FFDAA8F7E" blockId="4.[136,636,1352,1379]" box="[136,636,1352,1379]" pageId="4" pageNumber="33">
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<taxonomicName id="41E44D45FFE8FFBF2FEC083FFDAA8F7E" ID-CoL="3LXGS" authority="var. oligocephala" box="[136,636,1352,1379]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE8FFBF2FEC083FFE678F7E" bold="true" box="[136,433,1352,1379]" italics="true" pageId="4" pageNumber="33">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE8FFBF2E8C083FFDAA8F7E" bold="true" box="[488,636,1352,1379]" italics="true" pageId="4" pageNumber="33">oligocephala</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="CEFE654DFFE8FFBE2FEC08E6FD138E7D" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="description">
<paragraph id="865B36C6FFE8FFBE2FEC08E6FD138E7D" blockId="4.[136,1452,1425,2064]" lastBlockId="5.[136,1452,158,1481]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
Slender perennial herb with few soft, annual, herbaceous stems from small underground rootstock.
<emphasis id="B490EAD4FFE8FFBF2A6208E6FAA28FB6" box="[1286,1396,1425,1451]" italics="true" pageId="4" pageNumber="33">Rootstock</emphasis>
also with narrow fusiform tubers.
<emphasis id="B490EAD4FFE8FFBF2EBD08C2FDA78FD2" box="[473,625,1461,1487]" italics="true" pageId="4" pageNumber="33">Annual stems</emphasis>
80330, very rarely to 380, mm high, unbranched or rarely with one or two short side branches, faintly to distinctly ribbed, sparsely hairy to glabrescent basally, upwards densely silvery/ whitish hairy, hairs short-stalked, medifixed T-shaped with arms parallel to surface or very rarely sometimes with long-stalked, asymmetrically, Y-shaped hairs with arms shorter than stalk and stalk with reddish transverse cell walls.
<emphasis id="B490EAD4FFE8FFBF2FEC0B31FF008C42" box="[136,214,1606,1631]" italics="true" pageId="4" pageNumber="33">Leaves</emphasis>
alternate, sessile or very shortly petiolate, densely set to more scattered, discolorous, ovate, broadly ovate, cordate or narrowly ovate, elliptic or broadly elliptic, lowest pair often orbicular, 1028 × 3.520.0(25.0) mm (including petiole), apex acute, attenuate, mucronate apiculate, acuminate, obtuse or rounded, often shortly cuspidate, base cuneate, attenuate, obtuse, cordate or subcordate, margin entire, often undulate and distantly, sparsely serrulate, prominently pinnately veined, these raised on abaxial surface, on adaxial surface faint or sunken, adaxial surface sparsely hairy or glabrescent, hairs scattered over surface or confined to veins, often glandular punctate towards the apex, very rarely sometimes on adaxial side near apex and along margins of apex with long-stalked, asymmetrically, Y-shaped hairs with arms shorter than stalk and stalk with reddish transverse cell walls, abaxial surface densely hairy, hairs silvery/whitish, hairs short-stalked, medifixed, T-shaped with arms parallel to surface, similar to those on stems.
<emphasis id="B490EAD4FFE8FFBF2FEC0AFEFF3C8DBE" box="[136,234,1929,1955]" italics="true" pageId="4" pageNumber="33">Capitula</emphasis>
homogamous, discoid, very rarely heterogamous, discoid, few, on very short peduncles, arranged in dense, terminal, umbellate synflorescences or sometimes densely umbellate-racemosely arranged, rarely synflorescence more open and lax.
<emphasis id="B490EAD4FFE8FFBF2E430AA6FE598DF6" box="[295,399,2001,2027]" italics="true" pageId="4" pageNumber="33">Peduncle</emphasis>
(2)312(15), very rarely to 28, mm long, densely hairy, often hidden by uppermost leaves.
<emphasis id="B490EAD4FFE8FFBF2FEC0A82FF248212" box="[136,242,2037,2063]" italics="true" pageId="4" pageNumber="33">Involucre</emphasis>
broadly obconical to campanulate, 59(10) mm wide across upper part.
<emphasis id="B490EAD4FFE8FFBF2B4A0A82FB268212" box="[1070,1264,2037,2063]" italics="true" pageId="4" pageNumber="33">Involucral bracts</emphasis>
arranged in (4)5 rows, imbricate, outer shorter than inner, densely white or silvery hairy and glandular-punctate;
<emphasis id="B490EAD4FFE9FFBE2B8C0DE8FA8F8AA5" box="[1256,1369,159,184]" italics="true" pageId="5" pageNumber="34">outer row</emphasis>
lineartriangular or very narrowly ovate, (2.2)2.54.0 ×
<quantity id="411C9B23FFE9FFBE2DB70DB5FC898AC0" box="[723,863,194,221]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="34" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 mm</quantity>
, apex attenuate, usually cuspidate, usually reddish, hairy, glabrous or glabrescent, upper part of margin sometimes reddish, surface densely silvery hairy and glandular punctate, main vein sometimes obvious;
<emphasis id="B490EAD4FFE9FFBE2D3A0C7DFD378B39" box="[606,737,266,292]" italics="true" pageId="5" pageNumber="34">second row</emphasis>
linear-triangular, very narrowly ovate or rarely very narrowly obovate, 3.04.5(4.8) ×
<quantity id="411C9B23FFE9FFBE2ED40C59FDEB8B54" box="[432,573,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="34" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
, apex acuminate or attenuate, cuspidate, usually reddish, hairy, glabrous or glabrescent, upper part of margin sometimes reddish, surface densely silvery hairy and glandular punctate, main vein sometimes obvious and sometimes keeled in lower part;
<emphasis id="B490EAD4FFE9FFBE2C670C01FCBC8B8D" box="[771,874,374,400]" italics="true" pageId="5" pageNumber="34">third row</emphasis>
narrowly triangular, very narrowly ovate, narrowly elliptic or rarely narrowly obovate, (3.5)4.05.4 × 0.51.4(1.5) mm, apex attenuate or acuminate, cuspidate, usually reddish, sparsely hairy, glabrous or glabrescent, upper part of margin usually reddish or pink-purple; surface whitish hairy and glandular punctate, main vein usually visible, often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined;
<emphasis id="B490EAD4FFE9FFBE2EB00F71FD9A883D" box="[468,588,518,544]" italics="true" pageId="5" pageNumber="34">fourth row</emphasis>
narrowly ovate, narrowly elliptic or rarely narrowly obovate, 4.25.5(6.0) × 0.51.4(1.5) mm, apex acuminate or attenuate, cuspidate, usually reddish, glabrous or glabrescent, upper part of margin reddish or pink-purple, surface whitish hairy and glandular punctate, main vein usually visible, often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined;
<emphasis id="B490EAD4FFE9FFBE2CBD0F05FBE18891" box="[985,1079,626,652]" italics="true" pageId="5" pageNumber="34">fifth row</emphasis>
narrowly ovate, narrowly elliptic or rarely narrowly obovate or narrowly oblong, (4.0)4.55.5(6.0) ×
<quantity id="411C9B23FFE9FFBE2CF90FE1FBF188AC" box="[925,1063,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="5" pageNumber="34" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.61.2 mm</quantity>
, apex attenuate, cuspidate, usually reddish, glabrous or glabrescent, upper part of margin reddish or pink-purple, surface sometimes reddish in upper part, whitish hairy and glandular punctate, main vein ± prominent and often keeled in lower half, inside surface with prominent main vein or sometimes 3-veined.
<emphasis id="B490EAD4FFE9FFBE2DEF0E75FCD08901" box="[651,774,770,796]" italics="true" pageId="5" pageNumber="34">Receptacle</emphasis>
epaleate.
<emphasis id="B490EAD4FFE9FFBE2C1C0E74FC2D8901" box="[888,1019,770,796]" italics="true" pageId="5" pageNumber="34">Disc florets</emphasis>
bisexual, fertile, sometimes few outer florets female, actinomorphic; corolla pinkish or purple, funnelform;
<emphasis id="B490EAD4FFE9FFBE2CF20E51FC10895D" box="[918,966,806,832]" italics="true" pageId="5" pageNumber="34">tube</emphasis>
3.55.0(5.5) mm long, upper part of tube with few sessile glands and T-shaped hairs, 5-lobed;
<emphasis id="B490EAD4FFE9FFBE2DAE0E3DFCD38979" box="[714,773,842,868]" italics="true" pageId="5" pageNumber="34">lobes</emphasis>
(1.5)2.03.5(4.0) mm long, apex acute, with sessile glands and T-shaped hairs but around apex with unicellular hairs.
<emphasis id="B490EAD4FFE9FFBE2C530E19FC588995" box="[823,910,878,904]" italics="true" pageId="5" pageNumber="34">Anthers</emphasis>
(1.5)1.82.2(2.5) mm long, with (narrowly) ovate apical appendages, bases caudate.
<emphasis id="B490EAD4FFE9FFBE2D2D0EE5FDA989B1" box="[585,639,914,940]" italics="true" pageId="5" pageNumber="34">Style</emphasis>
(4.0)4.56.5(8.5) mm long, exserted, bifurcate;
<emphasis id="B490EAD4FFE9FFBE2BD70EE5FA8589B1" box="[1203,1363,914,940]" italics="true" pageId="5" pageNumber="34">style branches</emphasis>
pinkish purple, filiform, 1.0
<quantity id="411C9B23FFE9FFBE2E160EC1FE1089CC" box="[370,454,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="34" unit="mm" value="1.8">1.8 mm</quantity>
long, recurved, with acute hairs on outside of branches and onto the shaft below furcation, total inner surface stigmatic.
<emphasis id="B490EAD4FFE9FFBE2EBE0EADFDAA89E9" box="[474,636,986,1012]" italics="true" pageId="5" pageNumber="34">Ovary/cypsela</emphasis>
brown, ± obconical, 1.0
<quantity id="411C9B23FFE9FFBE2CCC0EADFC2989E8" box="[936,1023,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="34" unit="mm" value="1.5">1.5 mm</quantity>
long, ribbed, densely twin hairy and glandular punctate, with distinct carpopodium.
<taxonomicName id="41E44D45FFE9FFBE2DFA0E88FD248E05" box="[670,754,1023,1048]" class="Insecta" family="Miridae" genus="Pappus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B490EAD4FFE9FFBE2DFA0E88FD248E05" box="[670,754,1023,1048]" italics="true" pageId="5" pageNumber="34">Pappus</emphasis>
</taxonomicName>
dimorphic, an outer row of laciniate scales, 0.51.0 mm long and inner row of barbellate bristles, 3.55.0(5.5) mm long.
<emphasis id="B490EAD4FFE9FFBE2C510955FC378E21" box="[821,993,1058,1084]" italics="true" pageId="5" pageNumber="34">Flowering time</emphasis>
: July to December with one record each for January, February and April. (
<figureCitation id="1EDF2A43FFE9FFBE2E9A0931FD908E7D" box="[510,582,1094,1120]" captionStart="FIGURE 3" captionStartId="6.[136,229,2028,2050]" captionTargetBox="[215,1371,196,2004]" captionTargetId="figure-20@6.[215,1371,190,2004]" captionTargetPageId="6" captionText="FIGURE 3. Scan of Hilliardiella oligocephala var. oligocephala specimen (Koekemoer 4695, PRE) to show the slender habit and fewflowered, umbellate synflorescences." figureDoi="http://doi.org/10.5281/zenodo.7251474" httpUri="https://zenodo.org/record/7251474/files/figure.png" pageId="5" pageNumber="34">Figs 3</figureCitation>
,
<figureCitation id="1EDF2A43FFE9FFBE2D370931FD6F8E7D" box="[595,697,1094,1121]" captionStart="FIGURE 4" captionStartId="7.[136,229,1439,1461]" captionTargetBox="[151,1436,190,1415]" captionTargetId="figure-16@7.[151,1436,190,1415]" captionTargetPageId="7" captionText="FIGURE 4. Leaves and involucral bracts of Hilliardiella oligocephala. A. Abaxial surface of leaves of H. oligocephala var. oligocephala (Bester 8832, PRE) showing the pinnate veins(Nikondigital camera,photographer P.Herman).B.Abaxial surface of leaf of H.oligocephala var. oligocephala (Medley Wood 5755, PRE) showing the dense hairs and pinnate veins (Olympus digital microscope). C.Abaxial surface of the leaf of H. oligocephala var. besteri (Smuts &amp; Gillett 2406, PRE) showing the 3 main veins (Nikon digital camera, photographer P. Herman). D. Abaxial surface of the leaf of H. oligocephala var. besteri (Smuts &amp; Gillett 2406, PRE) showing the dense hairs and the 3 main veins (Olympus digital microscope). E. Inner surface of the inner involucral bract of H. oligocephala var. oligocephala (Codd 1796, PRE) showing the 3 main veins (Olympus digital microscope). F. Inner surface of the inner involucral bract of H. oligocephala var. besteri (Compton 26095, PRE) showing the 3 main veins (Olympus digital microscope)." figureDoi="http://doi.org/10.5281/zenodo.7251476" httpUri="https://zenodo.org/record/7251476/files/figure.png" pageId="5" pageNumber="34">4A, B, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE9FFBE2FD9091DFF228F41" pageId="5" pageNumber="34" type="diagnosis">
<paragraph id="865B36C6FFE9FFBE2FD9091DFF228F41" blockId="5.[136,1452,158,1481]" pageId="5" pageNumber="34">
<emphasis id="B490EAD4FFE9FFBE2FD9091DFE128E99" bold="true" box="[189,452,1130,1156]" pageId="5" pageNumber="34">Diagnostic characters</emphasis>
:—
<taxonomicName id="41E44D45FFE9FFBE2E8C091DFC098E99" authority="var. oligocephala" box="[488,991,1130,1157]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE9FFBE2E8C091DFCD98E99" box="[488,783,1130,1156]" italics="true" pageId="5" pageNumber="34">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE9FFBE2C29091DFC098E99" box="[845,991,1130,1156]" italics="true" pageId="5" pageNumber="34">oligocephala</emphasis>
</taxonomicName>
is a slender plant up to about
<quantity id="411C9B23FFE9FFBE2A2F091DFA7A8E98" box="[1355,1452,1130,1157]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="34" unit="mm" value="350.0">350 mm</quantity>
high, with only a few soft, herbaceous, slender stems from the underground rootstock, with narrowly or broadly ovate, cordate, elliptic or broadly elliptic, prominent pinnately veined, leaves and a few capitula arranged in dense terminal umbellate synflorescences.
<taxonomicName id="41E44D45FFE9FFBE2EDB09A0FDA18EED" box="[447,631,1238,1264]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="elaeagnoides">
<emphasis id="B490EAD4FFE9FFBE2EDB09A0FDA18EED" box="[447,631,1238,1264]" italics="true" pageId="5" pageNumber="34">H. elaeagnoides</emphasis>
</taxonomicName>
is a rigid, robust plant up to
<quantity id="411C9B23FFE9FFBE2CA709A1FC398EEC" box="[963,1007,1238,1265]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="34" unit="m" value="1.0">1 m</quantity>
high, with clumps of many rigid, erect stems from the underground rootstock, with many capitula in large, open corymbose synflorescences. Some specimens were seen with capitula more densely arranged in almost umbellate synflorescences, but their stems are still quite hard and rigid.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE9FFBE2FD90811FCAC8FD4" pageId="5" pageNumber="34" type="distribution">
<paragraph id="865B36C6FFE9FFBE2FD90811FCAC8FD4" blockId="5.[136,1452,158,1481]" pageId="5" pageNumber="34">
<emphasis id="B490EAD4FFE9FFBE2FD90811FE998F9D" bold="true" box="[189,335,1382,1408]" pageId="5" pageNumber="34">Distribution</emphasis>
:—
<taxonomicName id="41E44D45FFE9FFBE2E170811FC8B8F9D" authority="var. oligocephala" box="[371,861,1382,1409]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE9FFBE2E170811FD408F9D" box="[371,662,1382,1408]" italics="true" pageId="5" pageNumber="34">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE9FFBE2DAF0811FC8B8F9D" box="[715,861,1382,1408]" italics="true" pageId="5" pageNumber="34">oligocephala</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="FEF37656FFE9FFBE2C980811FB5C8F9D" box="[1020,1162,1382,1408]" name="South Africa" pageId="5" pageNumber="34">South Africa</collectingCountry>
and occurs in a small area around Barberton (
<collectingRegion id="4420F824FFE9FFBE2E3A08FDFE2E8FB9" box="[350,504,1418,1444]" country="South Africa" name="Mpumalanga" pageId="5" pageNumber="34">Mpumalanga</collectingRegion>
), through
<collectingRegion id="4420F824FFE9FFBE2D0F08FDFCCD8FB9" box="[619,795,1418,1444]" country="South Africa" name="KwaZulu-Natal" pageId="5" pageNumber="34">KwaZulu-Natal</collectingRegion>
and into the
<collectingRegion id="4420F824FFE9FFBE2CD608FCFB9E8FB9" box="[946,1096,1418,1445]" country="South Africa" name="Eastern Cape" pageId="5" pageNumber="34">Eastern Cape</collectingRegion>
(
<figureCitation id="1EDF2A43FFE9FFBE2B3C08FDFB488FB8" box="[1112,1182,1418,1445]" captionStart="FIGURE 5" captionStartId="8.[136,229,1530,1552]" captionTargetBox="[166,1422,448,1492]" captionTargetId="figure-124@8.[151,1436,425,1505]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map of Hilliardiella oligocephala var. oligocephala in South Africa." figureDoi="http://doi.org/10.5281/zenodo.7251478" httpUri="https://zenodo.org/record/7251478/files/figure.png" pageId="5" pageNumber="34">Fig. 5</figureCitation>
). Not as common as
<taxonomicName id="41E44D45FFE9FFBE2AF408FCFECA8FD5" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="34" phylum="Tracheophyta" rank="species" species="elaeagnoides">
<emphasis id="B490EAD4FFE9FFBE2AF408FCFECA8FD5" italics="true" pageId="5" pageNumber="34">H. elaeagnoides</emphasis>
</taxonomicName>
and usually overlooked because of its smaller stature.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE9FFB32FEC0B5BFE4D8215" lastPageId="8" lastPageNumber="37" pageId="5" pageNumber="34" type="materials_examined">
<paragraph id="865B36C6FFE9FFBE2FEC0B5BFDC38C5B" blockId="5.[136,533,1580,1606]" box="[136,533,1580,1606]" pageId="5" pageNumber="34">
<heading id="DD1381AAFFE9FFBE2FEC0B5BFDC38C5B" bold="true" box="[136,533,1580,1606]" fontSize="11" level="1" pageId="5" pageNumber="34" reason="1">
<emphasis id="B490EAD4FFE9FFBE2FEC0B5BFDC38C5B" bold="true" box="[136,533,1580,1606]" pageId="5" pageNumber="34">Additional specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="865B36C6FFE9FFBE2FED0B03FBA98D5F" blockId="5.[136,1452,1652,2075]" pageId="5" pageNumber="34">
<collectingCountry id="FEF37656FFE9FFBE2FED0B03FE8C8C93" box="[137,346,1652,1678]" name="South Africa" pageId="5" pageNumber="34">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="4420F824FFE9FFBE2E030B03FE298C93" box="[359,511,1652,1678]" country="South Africa" name="Mpumalanga" pageId="5" pageNumber="34">Mpumalanga</collectingRegion>
: Barberton District, ca.
<quantity id="411C9B23FFE9FFBE2C660B03FC988C93" box="[770,846,1652,1678]" metricMagnitude="3" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="34" unit="km" value="9.8">9.8 km</quantity>
directly SW of Barberton, Farm Schultzenhorst 718JT, N facing mountain slope, grassland,
<quantity id="411C9B23FFE9FFBE2D4F0BEFFD528CAE" box="[555,644,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.6179999999999999" pageId="5" pageNumber="34" unit="m" value="1618.0">1618 m</quantity>
, (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE9FFBE2C960BEFFB168CAF" box="[1010,1216,1688,1714]" pageId="5" pageNumber="34" value="2008-12-09">9 December 2008</date>
,
<emphasis id="B490EAD4FFE9FFBE2BA90BEEFA838CAF" box="[1229,1365,1688,1714]" italics="true" pageId="5" pageNumber="34">Bester 8832</emphasis>
(PRE). Mukwana Heights north of house,
<quantity id="411C9B23FFE9FFBE2D710BCBFDB28CCB" box="[533,612,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="34" unit="km" value="2.4">2.4 km</quantity>
S of Barberton, burnt N facing mountain slope,
<quantity id="411C9B23FFE9FFBE2BF40BCBFB348CCB" box="[1168,1250,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.1277599999999999" pageId="5" pageNumber="34" unit="ft" value="3700.0">3700 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2B950BCBFA9B8CCA" box="[1265,1357,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.128" pageId="5" pageNumber="34" unit="m" value="1128.0">1128 m</quantity>
], (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE9FFBE2EF10B97FD9D8CE7" box="[405,587,1760,1786]" pageId="5" pageNumber="34" value="1974-08-18">18 August 1974</date>
,
<emphasis id="B490EAD4FFE9FFBE2D330B96FCD48CE7" box="[599,770,1760,1786]" italics="true" pageId="5" pageNumber="34">De Souza 434b</emphasis>
(PRE). Barberton, Saddleback Mountain,
<quantity id="411C9B23FFE9FFBE2B870B97FAE28CE7" box="[1251,1332,1760,1786]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="5" pageNumber="34" unit="ft" value="4000.0">4000 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2A270B97FA498CE6" box="[1347,1439,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="5" pageNumber="34" unit="m" value="1219.0">1219 m</quantity>
], (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE9FFBE2E870A73FD4B8D03" box="[483,669,1796,1822]" pageId="5" pageNumber="34" value="1889-09">September 1889</date>
,
<emphasis id="B490EAD4FFE9FFBE2DCC0A73FCF18D03" box="[680,807,1796,1822]" italics="true" pageId="5" pageNumber="34">Galpin 425</emphasis>
(PRE). Locality not found: “Brommers” Mountains SE of Barberton, steep, rocky grassland, recently burnt,
<date id="F25A1006FFE9FFBE2DB10A5FFCBE8D5E" box="[725,872,1832,1859]" pageId="5" pageNumber="34" value="1985-07-10">10 July 1985</date>
,
<emphasis id="B490EAD4FFE9FFBE2C170A5FFBFC8D5F" box="[883,1066,1832,1858]" italics="true" pageId="5" pageNumber="34">Onderstall 1247</emphasis>
(PRE).
</paragraph>
<paragraph id="865B36C6FFE9FFB32FD90A3BFD498B71" blockId="5.[136,1452,1652,2075]" lastBlockId="8.[136,1451,158,401]" lastPageId="8" lastPageNumber="37" pageId="5" pageNumber="34">
<collectingRegion id="4420F824FFE9FFBE2FD90A3BFEA78D7B" box="[189,369,1868,1894]" country="South Africa" name="KwaZulu-Natal" pageId="5" pageNumber="34">KwaZulu-Natal</collectingRegion>
: Louwsburg Distr., Itala Nature Reserve, Potwe, mountain,
<quantity id="411C9B23FFE9FFBE2B470A3BFBA08D7B" box="[1059,1142,1868,1894]" metricMagnitude="3" metricUnit="m" metricValue="1.15824" pageId="5" pageNumber="34" unit="ft" value="3800.0">3800 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2BE20A3BFB328D7A" box="[1158,1252,1868,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.158" pageId="5" pageNumber="34" unit="m" value="1158.0">1158 m</quantity>
], (QDS: 2731AC Louwsburg),
<date id="F25A1006FFE9FFBE2E440A07FE3B8D96" box="[288,493,1904,1931]" pageId="5" pageNumber="34" value="1975-12-08">8 December 1975</date>
,
<emphasis id="B490EAD4FFE9FFBE2E9D0A06FD308D97" box="[505,742,1904,1930]" italics="true" pageId="5" pageNumber="34">Brown &amp; Shapiro 34</emphasis>
(PRE). Ntonjaneni Distr., 3 ½ m [
<quantity id="411C9B23FFE9FFBE2B0B0A07FB128D97" box="[1135,1220,1904,1931]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="34" unit="km" value="5.6">5.6 km</quantity>
] S of Ntonjaneni on road to Melmoth, dense grassveld, recently burnt,
<quantity id="411C9B23FFE9FFBE2DDF0AE3FCDD8DB3" box="[699,779,1940,1966]" metricMagnitude="3" metricUnit="m" metricValue="1.03632" pageId="5" pageNumber="34" unit="ft" value="3400.0">3400 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2C7D0AE3FCA38DB2" box="[793,885,1940,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.036" pageId="5" pageNumber="34" unit="m" value="1036.0">1036 m</quantity>
], (QDS: 2831CB Nkandla),
<date id="F25A1006FFE9FFBE2BD60AE3FAB28DB3" box="[1202,1380,1940,1966]" pageId="5" pageNumber="34" value="1946-10-08">8 October 1946</date>
,
<emphasis id="B490EAD4FFE9FFBE2A0B0AE3FF168DCF" italics="true" pageId="5" pageNumber="34">Codd 1796</emphasis>
(PRE). Ngoye Forest [Umgoye/Ongoye Forest], grassland between exposed rock faces,
<quantity id="411C9B23FFE9FFBE2BC70ACFFB258DCF" box="[1187,1267,1976,2002]" metricMagnitude="2" metricUnit="m" metricValue="4.8768" pageId="5" pageNumber="34" unit="ft" value="1600.0">1600 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2A650ACFFA998DCE" box="[1281,1359,1976,2003]" metricMagnitude="2" metricUnit="m" metricValue="4.88" pageId="5" pageNumber="34" unit="m" value="488.0">488 m</quantity>
], (QDS: 2831DC Nkandla),
<date id="F25A1006FFE9FFBE2E050AABFDE38DEB" box="[353,565,2012,2038]" pageId="5" pageNumber="34" value="1973-12-11">11 December 1973</date>
,
<emphasis id="B490EAD4FFE9FFBE2D5B0AABFD6D8DEB" box="[575,699,2012,2038]" italics="true" pageId="5" pageNumber="34">Stirton 446</emphasis>
(PRE). Zululand, Hlabisa district, west of Charters Creek, amongst grasses in sandy soil near vlei,
<quantity id="411C9B23FFE9FFBE2E9C0577FDEB8207" box="[504,573,2048,2074]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="34" unit="ft" value="100.0">100 ft</quantity>
[
<quantity id="411C9B23FFE9FFBE2D2A0577FD478206" box="[590,657,2048,2075]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="34" unit="m" value="30.0">30 m</quantity>
], (QDS: 2832AB Mtubatuba),
<date id="F25A1006FFE9FFBE2C9C0577FB1F8206" box="[1016,1225,2048,2075]" pageId="5" pageNumber="34" value="1955-12-05">5 December 1955</date>
,
<emphasis id="B490EAD4FFE9FFBE2BB30577FA828207" box="[1239,1364,2048,2074]" italics="true" pageId="5" pageNumber="34">Ward 2847</emphasis>
(PRE). St Lucia, S of Lake, Dukuduku State forest, moist riverine forest/thicket,
<quantity id="411C9B23FFEBFFBC2CC70B97FC0A8CE7" box="[931,988,1760,1786]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="36" unit="m" value="20.0">20 m</quantity>
, (QDS: 2832AD Mtubatuba),
<date id="F25A1006FFEBFFBC2A420B97FF158D03" pageId="7" pageNumber="36" value="1989-12-09">9 December 1989</date>
,
<emphasis id="B490EAD4FFEBFFBC2FB40A74FE8C8D03" box="[208,346,1795,1822]" italics="true" pageId="7" pageNumber="36">Hobson 370</emphasis>
(PRE). Witbank, Balgowan, [Lions River Distr.], dolorite outcrop, grassland,
<quantity id="411C9B23FFEBFFBC2B840A73FAE48D00" box="[1248,1330,1795,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.46304" pageId="7" pageNumber="36" unit="ft" value="4800.0">4800 ft</quantity>
[
<quantity id="411C9B23FFEBFFBC2A260A74FA728D03" box="[1346,1444,1795,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="7" pageNumber="36" unit="m" value="1463.0">1463 m</quantity>
], (QDS: 2930AC Pietermaritzburg),
<date id="F25A1006FFEBFFBC2D770A5FFD058D5C" box="[531,723,1831,1858]" pageId="7" pageNumber="36" value="1964-10-12">12 October 1964</date>
,
<emphasis id="B490EAD4FFEBFFBC2DBB0A50FC858D5F" box="[735,851,1831,1858]" italics="true" pageId="7" pageNumber="36">Moll 1219</emphasis>
(PRE). Greytown,
<quantity id="411C9B23FFEBFFBC2B4A0A5FFBA98D5C" box="[1070,1151,1831,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.0424159999999998" pageId="7" pageNumber="36" unit="ft" value="3420.0">3420 ft</quantity>
[
<quantity id="411C9B23FFEBFFBC2BE90A5FFB3C8D5F" box="[1165,1258,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.042" pageId="7" pageNumber="36" unit="m" value="1042.0">1042 m</quantity>
], (QDS: 2930BA Pietermaritzburg), Oct-Nov 1931,
<emphasis id="B490EAD4FFEBFFBC2D620A3CFDB98D78" box="[518,623,1867,1893]" italics="true" pageId="7" pageNumber="36">Wylie s.n.</emphasis>
TvlMus 33875 (PRE). Pietermaritzburg, Silverglen Nature Reserve, NE of Umlazi Nursery, recently burnt grassland, (QDS: 2930DD Pietermaritzburg),
<date id="F25A1006FFEBFFBC2C950A07FB648D97" box="[1009,1202,1903,1930]" pageId="7" pageNumber="36" value="1985-10-21">21 October 1985</date>
,
<emphasis id="B490EAD4FFEBFFBC2BD90A07FA818D97" box="[1213,1367,1904,1930]" italics="true" pageId="7" pageNumber="36">Ngwenya 181</emphasis>
(PRE). [Pinetown], Kloof,
<quantity id="411C9B23FFEBFFBC2E000AE4FE638DB0" box="[356,437,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="7" pageNumber="36" unit="ft" value="1500.0">1500 ft</quantity>
[
<quantity id="411C9B23FFEBFFBC2EA00AE4FDC58DB3" box="[452,531,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="7" pageNumber="36" unit="m" value="457.0">457 m</quantity>
], (QDS: 2930DD Pietermaritzburg),
<date id="F25A1006FFEBFFBC2CD10AE3FB428DB3" box="[949,1172,1939,1966]" pageId="7" pageNumber="36" value="1893-09-14">14 September 1893</date>
,
<emphasis id="B490EAD4FFEBFFBC2BC40AE4FA808DB3" box="[1184,1366,1939,1966]" italics="true" pageId="7" pageNumber="36">Schlechter 3203</emphasis>
(PRE). Hill near Pinetown,
<quantity id="411C9B23FFEBFFBC2E0B0AC0FE178DCC" box="[367,449,1975,2002]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="7" pageNumber="36" unit="ft" value="1500.0">1500 ft</quantity>
[
<quantity id="411C9B23FFEBFFBC2EB50AC0FDF78DCF" box="[465,545,1975,2002]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="7" pageNumber="36" unit="m" value="457.0">457 m</quantity>
], (QDS: 2930DD Pietermaritzburg),
<date id="F25A1006FFEBFFBC2CA20ACFFBA88DCF" box="[966,1150,1976,2002]" pageId="7" pageNumber="36" value="1893-08-29">29 August 1893</date>
,
<emphasis id="B490EAD4FFEBFFBC2BEF0AC0FB2E8DCC" box="[1163,1272,1975,2001]" italics="true" pageId="7" pageNumber="36">Wylie s.n.</emphasis>
Nat. Herb. 495 (PRE,
<specimenCount id="90E2FD4FFFEBFFBC2FB10AABFEB18DE8" box="[213,359,2011,2038]" pageId="7" pageNumber="36" type="generic">2 specimens</specimenCount>
). Weza State Forest, foothills of Ngeli Mountain near Loveday [Lovedale?] lookout, grassland, (QDS: 3029DA Kokstad),
<date id="F25A1006FFEBFFBC2EA60577FD718204" box="[450,679,2047,2074]" pageId="7" pageNumber="36" value="1986-11-20">20 November 1986</date>
,
<emphasis id="B490EAD4FFEBFFBC2DD30577FC898204" box="[695,863,2047,2073]" italics="true" pageId="7" pageNumber="36">Van Wyk 7561</emphasis>
(PRE). District Alexandra, Station Dumisa, Farm Friedenau,
<quantity id="411C9B23FFE4FFB32E620DE9FE868AA4" box="[262,336,158,185]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="37" unit="m" value="600.0">600 m</quantity>
, (QDS: 3030AD Port Shepstone),
<date id="F25A1006FFE4FFB32DB10DE9FC478AA5" box="[725,913,158,185]" pageId="8" pageNumber="37" value="1908-01-31">31 January 1908</date>
,
<emphasis id="B490EAD4FFE4FFB32CF90DE9FBF28AA5" box="[925,1060,158,184]" italics="true" pageId="8" pageNumber="37">Rudatis 235</emphasis>
(PRE). District Alexandra, Station Dumisa, Campbellton,
<quantity id="411C9B23FFE4FFB32EEB0DB5FE0F8AC0" box="[399,473,194,221]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="37" unit="m" value="800.0">800 m</quantity>
, (QDS: 3030AD Port Shepstone),
<date id="F25A1006FFE4FFB32C000DB5FBE58AC1" box="[868,1075,194,220]" pageId="8" pageNumber="37" value="1912-11-08">8 November 1912</date>
,
<emphasis id="B490EAD4FFE4FFB32B240DB5FB008AC1" box="[1088,1238,194,220]" italics="true" pageId="8" pageNumber="37">Rudatis 1745</emphasis>
(PRE). Dududu,
<quantity id="411C9B23FFE4FFB32AF90DB5FF7A8B1D" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="37" unit="km" value="2.0">2 km</quantity>
W of the Cedars, gentle slope, recently burnt grassland, loamy soil, (QDS: 3030BA Port Shepstone),
<date id="F25A1006FFE4FFB32A440D91FF158B39" pageId="8" pageNumber="37" value="1992-11-05">5 November 1992</date>
,
<emphasis id="B490EAD4FFE4FFB32FB40C7DFE558B39" box="[208,387,266,292]" italics="true" pageId="8" pageNumber="37">Sikhakhane 268</emphasis>
(PRE). Port Edward, Clearwater, (QDS: 3130AA Port Edward),
<date id="F25A1006FFE4FFB32B0E0C7DFAF78B39" box="[1130,1313,266,292]" pageId="8" pageNumber="37" value="1982-08-22">22 August 1982</date>
,
<emphasis id="B490EAD4FFE4FFB32A4A0C7DFA7D8B39" box="[1326,1451,266,292]" italics="true" pageId="8" pageNumber="37">Abbott 241</emphasis>
(PRE). Just outside Port Edward on road to Bizana, hillside, grassland, recently burned, (QDS: 3130AA Port Edward),
<date id="F25A1006FFE4FFB32FEC0C25FEB38B71" box="[136,357,338,364]" pageId="8" pageNumber="37" value="1996-09-20">20 September 1996</date>
,
<emphasis id="B490EAD4FFE4FFB32E140C25FD9C8B71" box="[368,586,338,364]" italics="true" pageId="8" pageNumber="37">Germishuizen 8659</emphasis>
(PRE).
</paragraph>
<caption id="D29B664EFFEAFFBD2FEC0A9BFE3C823B" ID-DOI="http://doi.org/10.5281/zenodo.7251474" ID-Zenodo-Dep="7251474" httpUri="https://zenodo.org/record/7251474/files/figure.png" pageId="6" pageNumber="35" startId="6.[136,229,2028,2050]" targetBox="[215,1371,196,2004]" targetPageId="6">
<paragraph id="865B36C6FFEAFFBD2FEC0A9BFE3C823B" blockId="6.[136,1451,2028,2086]" pageId="6" pageNumber="35">
<emphasis id="B490EAD4FFEAFFBD2FEC0A9BFF2E821C" bold="true" box="[136,248,2028,2050]" pageId="6" pageNumber="35">FIGURE 3</emphasis>
. Scan of
<taxonomicName id="41E44D45FFEAFFBD2E320A9BFD29821C" authority="var. oligocephala" box="[342,767,2028,2050]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="35" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFEAFFBD2E320A9BFD87821C" box="[342,593,2028,2049]" italics="true" pageId="6" pageNumber="35">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEAFFBD2DE60A9BFD29821C" box="[642,767,2028,2049]" italics="true" pageId="6" pageNumber="35">oligocephala</emphasis>
</taxonomicName>
specimen (
<emphasis id="B490EAD4FFEAFFBD2C140A9BFBC2821C" box="[880,1044,2028,2049]" italics="true" pageId="6" pageNumber="35">Koekemoer 4695</emphasis>
, PRE) to show the slender habit and fewflowered, umbellate
<emphasis id="B490EAD4FFEAFFBD2E2A0567FE328238" box="[334,484,2064,2085]" italics="true" pageId="6" pageNumber="35">synflorescences</emphasis>
.
</paragraph>
</caption>
<caption id="D29B664EFFEBFFBC2FEC08E8FC458CAC" ID-DOI="http://doi.org/10.5281/zenodo.7251476" ID-Zenodo-Dep="7251476" httpUri="https://zenodo.org/record/7251476/files/figure.png" pageId="7" pageNumber="36" startId="7.[136,229,1439,1461]" targetBox="[151,1436,190,1415]" targetPageId="7">
<paragraph id="865B36C6FFEBFFBC2FEC08E8FC458CAC" blockId="7.[136,1451,1439,1713]" pageId="7" pageNumber="36">
<emphasis id="B490EAD4FFEBFFBC2FEC08E8FF218FA9" bold="true" box="[136,247,1439,1461]" pageId="7" pageNumber="36">FIGURE 4</emphasis>
. Leaves and involucral bracts of
<taxonomicName id="41E44D45FFEBFFBC2D5008E8FCF88FA9" authorityName="H. Robinson" authorityYear="1999" baseAuthorityName="DC." box="[564,814,1439,1460]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="species" species="oligocephala">
<emphasis id="B490EAD4FFEBFFBC2D5008E8FCF88FA9" box="[564,814,1439,1460]" italics="true" pageId="7" pageNumber="36">Hilliardiella oligocephala</emphasis>
</taxonomicName>
. A. Abaxial surface of leaves of
<taxonomicName id="41E44D45FFEBFFBC2B0108D7FA7D8FA9" authority="var. oligocephala" box="[1125,1451,1439,1461]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFEBFFBC2B0108D7FB298FA9" box="[1125,1279,1439,1461]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2A4A08E8FA7D8FA9" box="[1326,1451,1439,1460]" italics="true" pageId="7" pageNumber="36">oligocephala</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2FF408B3FED78FC5" box="[144,257,1475,1497]" italics="true" pageId="7" pageNumber="36">Bester 8832</emphasis>
, PRE) showing the pinnate veins (Nikon digital camera, photographer P. Herman). B.Abaxial surface of leaf of
<taxonomicName id="41E44D45FFEBFFBC2A7708B3FEF88FE1" authority="var. oligocephala" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFEBFFBC2A7708B3FA7D8FC5" box="[1299,1451,1475,1497]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2FD50890FEF88FE1" box="[177,302,1511,1532]" italics="true" pageId="7" pageNumber="36">oligocephala</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2E580890FE258FE1" box="[316,499,1511,1532]" italics="true" pageId="7" pageNumber="36">Medley Wood 5755</emphasis>
, PRE) showing the dense hairs and pinnate veins (Olympus digital microscope). C. Abaxial surface of the leaf of
<taxonomicName id="41E44D45FFEBFFBC2E6B0B7BFDC88C3D" authorityName="P. P. J. Herman" authorityYear="2022" box="[271,542,1547,1569]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFEBFFBC2E6B0B7BFE7D8C3D" box="[271,427,1547,1569]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2EB90B7CFDC88C3D" box="[477,542,1547,1568]" italics="true" pageId="7" pageNumber="36">besteri</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2D4A0B7CFD2B8C3D" box="[558,765,1547,1568]" italics="true" pageId="7" pageNumber="36">Smuts &amp; Gillett 2406</emphasis>
, PRE) showing the 3 main veins (Nikon digital camera, photographer P. Herman). D. Abaxial surface of the leaf of
<taxonomicName id="41E44D45FFEBFFBC2D5F0B47FC918C59" authorityName="P. P. J. Herman" authorityYear="2022" box="[571,839,1583,1605]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFEBFFBC2D5F0B47FD008C59" box="[571,726,1583,1605]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2C620B58FC918C59" box="[774,839,1583,1604]" italics="true" pageId="7" pageNumber="36">besteri</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2C310B58FBF78C59" box="[853,1057,1583,1604]" italics="true" pageId="7" pageNumber="36">Smuts &amp; Gillett 2406</emphasis>
, PRE) showing the dense hairs and the 3 main veins (Olympus digital microscope). E. Inner surface of the inner involucral bract of
<taxonomicName id="41E44D45FFEBFFBC2C8C0B23FAF88C75" authority="var. oligocephala" box="[1000,1326,1619,1641]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFEBFFBC2C8C0B23FB548C75" box="[1000,1154,1619,1641]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2BD50B24FAF88C75" box="[1201,1326,1619,1640]" italics="true" pageId="7" pageNumber="36">oligocephala</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2A580B24FA738C75" box="[1340,1445,1619,1640]" italics="true" pageId="7" pageNumber="36">Codd 1796</emphasis>
, PRE) showing the 3 main veins (Olympus digital microscope). F. Inner surface of the inner involucral bract of
<taxonomicName id="41E44D45FFEBFFBC2BC60B0FFA7D8C91" authorityName="P. P. J. Herman" authorityYear="2022" box="[1186,1451,1655,1677]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="36" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFEBFFBC2BC60B0FFAEA8C91" box="[1186,1340,1655,1677]" italics="true" pageId="7" pageNumber="36">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFEBFFBC2A0E0B00FA7D8C91" box="[1386,1451,1655,1676]" italics="true" pageId="7" pageNumber="36">besteri</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFEBFFBC2FF40BECFEFC8CAD" box="[144,298,1691,1712]" italics="true" pageId="7" pageNumber="36">Compton 26095</emphasis>
, PRE) showing the 3 main veins (Olympus digital microscope).
</paragraph>
</caption>
<paragraph id="865B36C6FFE4FFB32FD90C00FB2E8B8D" blockId="8.[136,1451,158,401]" box="[189,1272,374,401]" pageId="8" pageNumber="37">
Vague locality: Zululand,
<quantity id="411C9B23FFE4FFB32E840C01FD9A8B8D" box="[480,588,374,400]" metricMagnitude="2" metricUnit="m" metricValue="3.049524" metricValueMax="6.096" metricValueMin="0.003048" pageId="8" pageNumber="37" unit="ft" value="1000.5" valueMax="2000.0" valueMin="1.0">12000 ft</quantity>
[
<quantity id="411C9B23FFE4FFB32D3E0C01FD368B8C" box="[602,736,374,401]" metricMagnitude="2" metricUnit="m" metricValue="4.575" metricValueMax="6.1" metricValueMin="3.05" pageId="8" pageNumber="37" unit="m" value="457.5" valueMax="610.0" valueMin="305.0">305610 m</quantity>
],
<date id="F25A1006FFE4FFB32D950C01FCB88B8C" box="[753,878,374,401]" pageId="8" pageNumber="37" value="1895-04">April 1895</date>
,
<emphasis id="B490EAD4FFE4FFB32C1D0C01FB758B8C" box="[889,1187,374,401]" italics="true" pageId="8" pageNumber="37">Wylie / Medley Wood 5755</emphasis>
(PRE).
</paragraph>
<caption id="D29B664EFFE4FFB32FEC088DFBD88C0D" ID-DOI="http://doi.org/10.5281/zenodo.7251478" ID-Zenodo-Dep="7251478" box="[136,1038,1530,1552]" httpUri="https://zenodo.org/record/7251478/files/figure.png" pageId="8" pageNumber="37" startId="8.[136,229,1530,1552]" targetBox="[166,1422,448,1492]" targetPageId="8">
<paragraph id="865B36C6FFE4FFB32FEC088DFBD88C0D" blockId="8.[136,1038,1530,1552]" box="[136,1038,1530,1552]" pageId="8" pageNumber="37">
<emphasis id="B490EAD4FFE4FFB32FEC088DFF218C12" bold="true" box="[136,247,1530,1552]" pageId="8" pageNumber="37">FIGURE 5</emphasis>
. Distribution map of
<taxonomicName id="41E44D45FFE4FFB32EA3088DFCBB8C12" authority="var. oligocephala" box="[455,877,1530,1551]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="37" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE4FFB32EA3088DFD178C12" box="[455,705,1530,1551]" italics="true" pageId="8" pageNumber="37">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE4FFB32D94088DFCBB8C12" box="[752,877,1530,1551]" italics="true" pageId="8" pageNumber="37">oligocephala</emphasis>
</taxonomicName>
in South Africa.
</paragraph>
</caption>
<paragraph id="865B36C6FFE4FFB32FD90B48FE4D8215" blockId="8.[136,1452,1598,2056]" pageId="8" pageNumber="37">
<collectingRegion id="4420F824FFE4FFB32FD90B48FE838C45" box="[189,341,1598,1625]" country="South Africa" name="Eastern Cape" pageId="8" pageNumber="37">Eastern Cape</collectingRegion>
: Griqualand East, near Clydesdale, Mount Malowe, grassy slopes,
<quantity id="411C9B23FFE4FFB32B520B49FB538C45" box="[1078,1157,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.0668" pageId="8" pageNumber="37" unit="ft" value="3500.0">3500 ft</quantity>
[
<quantity id="411C9B23FFE4FFB32BF50B49FB3A8C44" box="[1169,1260,1598,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.069" pageId="8" pageNumber="37" unit="m" value="1069.0">1069 m</quantity>
], (QDS: 3029BD Kokstad),
<date id="F25A1006FFE4FFB32F990B15FE608C60" box="[253,438,1634,1661]" pageId="8" pageNumber="37" value="1885-11">November 1885</date>
,
<emphasis id="B490EAD4FFE4FFB32EA50B14FD948C61" box="[449,578,1634,1660]" italics="true" pageId="8" pageNumber="37">Tyson 2148</emphasis>
(PRE,
<specimenCount id="90E2FD4FFFE4FFB32DF00B15FCF28C61" box="[660,804,1634,1660]" pageId="8" pageNumber="37" type="generic">2 specimens</specimenCount>
). Bizana Commonage, grassveld,
<quantity id="411C9B23FFE4FFB32BC60B15FB248C61" box="[1186,1266,1634,1661]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="37" unit="ft" value="2500.0">2500 ft</quantity>
[
<quantity id="411C9B23FFE4FFB32A640B15FA998C60" box="[1280,1359,1634,1661]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="37" unit="m" value="762.0">762 m</quantity>
], (QDS: 3029DD Kokstad),
<date id="F25A1006FFE4FFB32E0D0BF1FD9F8CBC" box="[361,585,1670,1697]" pageId="8" pageNumber="37" value="1945-11-22">22 November 1945</date>
,
<emphasis id="B490EAD4FFE4FFB32D320BF1FD238CBD" box="[598,757,1670,1696]" italics="true" pageId="8" pageNumber="37">Acocks 12257</emphasis>
(PRE). Bizana District, flats
<quantity id="411C9B23FFE4FFB32B2E0BF1FB658CBD" box="[1098,1203,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="8" pageNumber="37" unit="mi" value="0.5">0.5 miles</quantity>
[
<quantity id="411C9B23FFE4FFB32BA70BF1FACF8CBD" box="[1219,1305,1670,1696]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="37" unit="km" value="0.8">0.8 km</quantity>
] W of town, sour grassveld,
<quantity id="411C9B23FFE4FFB32E530BDDFE568CD8" box="[311,384,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="37" unit="m" value="762.0">762 m</quantity>
, (QDS: 3029DD Kokstad),
<date id="F25A1006FFE4FFB32DDD0BDDFC428CD8" box="[697,916,1706,1733]" pageId="8" pageNumber="37" value="1945-11-22">22 November 1945</date>
,
<emphasis id="B490EAD4FFE4FFB32CFB0BDCFBDA8CD9" box="[927,1036,1706,1733]" italics="true" pageId="8" pageNumber="37">Story 587</emphasis>
(PRE). Ntsubane Forest Station near Lusikisiki, grassy plains, (QDS: 3129BC Port St Johns),
<date id="F25A1006FFE4FFB32C6D0BB9FC108CF5" box="[777,966,1742,1768]" pageId="8" pageNumber="37" value="1929-02-02">2 February 1929</date>
,
<emphasis id="B490EAD4FFE4FFB32CB60BB9FBBB8CF5" box="[978,1133,1742,1768]" italics="true" pageId="8" pageNumber="37">Galpin 10993</emphasis>
(PRE). Transkei, Mkambati Nature Reserve, burnt grassland,
<quantity id="411C9B23FFE4FFB32D620B85FD878D10" box="[518,593,1778,1805]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="37" unit="m" value="230.0">230 m</quantity>
, (QDS: 3129BD Port St Johns),
<date id="F25A1006FFE4FFB32CAF0B85FB4C8D10" box="[971,1178,1778,1805]" pageId="8" pageNumber="37" value="1985-02-19">19 February 1985</date>
,
<emphasis id="B490EAD4FFE4FFB32BC30B85FA838D11" box="[1191,1365,1778,1804]" italics="true" pageId="8" pageNumber="37">Shackleton 228</emphasis>
(PRE). Transkei, 1.3 myl [
<quantity id="411C9B23FFE4FFB32E070A61FE6F8D2D" box="[355,441,1814,1840]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="37" unit="km" value="2.1">2.1 km</quantity>
] vanaf die kantore by Mkambati op pad na die hospitaal, in onlangs gebrande grasveld [
<quantity id="411C9B23FFE4FFB32FEB0A4DFF368D49" box="[143,224,1850,1876]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="37" unit="km" value="2.1">2.1 km</quantity>
from the offices at Mkambati on the way to the hospital, in recently burnt grassveld], (QDS: 3129BD Port St Johns),
<date id="F25A1006FFE4FFB32FBB0A29FEB08D65" box="[223,358,1886,1912]" pageId="8" pageNumber="37" value="1976-07-09">9 July 1976</date>
,
<emphasis id="B490EAD4FFE4FFB32E150A28FDD98D65" box="[369,527,1886,1912]" italics="true" pageId="8" pageNumber="37">Van Wyk 1555</emphasis>
(PRE). Flora of Coffee Bay, Tembuland, on grassy slopes, (QDS: 3129CC Port St Johns),
<date id="F25A1006FFE4FFB32FBB0AF5FE408D81" box="[223,406,1922,1948]" pageId="8" pageNumber="37" value="1919-11">November 1919</date>
,
<emphasis id="B490EAD4FFE4FFB32EC50AF4FDD98D81" box="[417,527,1923,1948]" italics="true" pageId="8" pageNumber="37">Tyson s.n.</emphasis>
PRE44465 (PRE). Bizana District, Umtentu mouth, open grassveld,
<quantity id="411C9B23FFE4FFB32A750AF5FA858D81" box="[1297,1363,1922,1948]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="8" pageNumber="37" unit="ft" value="100.0">100 ft</quantity>
[
<quantity id="411C9B23FFE4FFB32A040AF5FA498D80" box="[1376,1439,1922,1949]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="37" unit="m" value="30.0">30 m</quantity>
], (QDS: 3130AA Port Edward),
<date id="F25A1006FFE4FFB32EBB0AD1FD478DDC" box="[479,657,1958,1985]" pageId="8" pageNumber="37" value="1953-08-17">17 August 1953</date>
,
<emphasis id="B490EAD4FFE4FFB32DFF0AD0FCC38DDD" box="[667,789,1958,1985]" italics="true" pageId="8" pageNumber="37">Story 4171</emphasis>
(PRE). District Kentani, margin of woods,
<quantity id="411C9B23FFE4FFB32B920AD1FA908DDD" box="[1270,1350,1958,1985]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="8" pageNumber="37" unit="ft" value="1500.0">1500 ft</quantity>
[
<quantity id="411C9B23FFE4FFB32A360AD1FA498DDC" box="[1362,1439,1958,1985]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="8" pageNumber="37" unit="m" value="457.0">457 m</quantity>
], (QDS: 3228CB Butterworth), 1901,
<emphasis id="B490EAD4FFE4FFB32D7B0ABDFD5A8DF9" box="[543,652,1994,2020]" italics="true" pageId="8" pageNumber="37">Pegler 64</emphasis>
(PRE). Kentani, (QDS: 3228CB Butterworth),
<date id="F25A1006FFE4FFB32BFA0ABDFA808DF9" box="[1182,1366,1994,2020]" pageId="8" pageNumber="37" value="1906-11">November 1906</date>
,
<emphasis id="B490EAD4FFE4FFB32A050ABDFF798215" italics="true" pageId="8" pageNumber="37">Pegler s.n.</emphasis>
TvlMus4126 (PRE).
</paragraph>
</subSubSection>
</treatment>
</document>