treatments-xml/data/0E/4D/87/0E4D87D0FFE5FFB72FEC0DD7FE778AC1.xml
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<document id="F14C9A478940B3E34A9034A1670464FC" ID-CLB-Dataset="173353" ID-DOI="10.11646/phytotaxa.570.1.3" ID-GBIF-Dataset="5948903b-2326-4611-bd09-2d6314cfe677" ID-ISSN="1179-3163" ID-Zenodo-Dep="7251466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666767854836" checkinUser="plazi" docAuthor="Herman, Paul P. J." docDate="2022" docId="0E4D87D0FFE5FFB72FEC0DD7FE778AC1" docLanguage="en" docName="phytotaxa.570.1.3.pdf" docOrigin="Phytotaxa 570 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.570.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Hilliardiella oligocephala var. besteri P. P. J. Herman 2022, var. nov." docType="treatment" docVersion="5" lastPageNumber="41" masterDocId="F274FFA8FFECFFBB2F640D77FFD68A1D" masterDocTitle="Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety" masterLastPageNumber="42" masterPageNumber="29" pageNumber="38" updateTime="1699409107501" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="89D6A33ED44049DB0E3C55656C7225D4">Hilliardiella elaeagnoides and H. oligocephala (Asteraceae, Vernonieae), two distinct species and the description of a new variety</mods:title>
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<mods:name id="82117AEED71F2386AC61C4B43D59AB7F" type="personal">
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<mods:namePart id="5AE7D6EC71EB119979330EA26D14CB7D">Herman, Paul P. J.</mods:namePart>
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<mods:title id="A0D3F338233A55635627359176DCE947">Phytotaxa</mods:title>
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<mods:date id="B2DF071536C2795BA22D92A2831BD42D">2022</mods:date>
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<treatment id="0E4D87D0FFE5FFB72FEC0DD7FE778AC1" ID-DOI="http://doi.org/10.5281/zenodo.7258183" ID-GBIF-Taxon="203687367" ID-Zenodo-Dep="7258183" LSID="urn:lsid:plazi:treatment:0E4D87D0FFE5FFB72FEC0DD7FE778AC1" httpUri="http://treatment.plazi.org/id/0E4D87D0FFE5FFB72FEC0DD7FE778AC1" lastPageId="12" lastPageNumber="41" pageId="9" pageNumber="38">
<subSubSection id="CEFE654DFFE5FFB22FEC0DD7FC0B8AA6" box="[136,989,160,188]" pageId="9" pageNumber="38" type="nomenclature">
<paragraph id="865B36C6FFE5FFB22FEC0DD7FC0B8AA6" blockId="9.[136,989,160,188]" box="[136,989,160,188]" pageId="9" pageNumber="38">
<heading id="DD1381AAFFE5FFB22FEC0DD7FC0B8AA6" box="[136,989,160,188]" fontSize="11" level="2" pageId="9" pageNumber="38" reason="3">
<taxonomicName id="41E44D45FFE5FFB22FEC0DD7FCA68AA1" ID-CoL="B2WGK" authority="P.P.J.Herman" authorityName="P. P. J. Herman" authorityYear="2022" box="[136,880,160,188]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="variety" species="oligocephala" status="var. nov." variety="besteri">
<emphasis id="B490EAD4FFE5FFB22FEC0DD7FE678AA6" bold="true" box="[136,433,160,187]" italics="true" pageId="9" pageNumber="38">Hilliardiella oligocephala</emphasis>
(DC.) H.Rob. var.
<emphasis id="B490EAD4FFE5FFB22DE30DD7FD058AA6" bold="true" box="[647,723,160,187]" italics="true" pageId="9" pageNumber="38">besteri</emphasis>
P.P.J.Herman
</taxonomicName>
,
<emphasis id="B490EAD4FFE5FFB22C1F0DD6FC0B8AA6" bold="true" box="[891,989,161,187]" italics="true" pageId="9" pageNumber="38">
<taxonomicNameLabel id="AFA357AFFFE5FFB22C1F0DD6FC0B8AA6" box="[891,989,161,187]" pageId="9" pageNumber="38" rank="variety">var. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FEC0D9AFDEA8B3A" pageId="9" pageNumber="38" type="diagnosis">
<paragraph id="865B36C6FFE5FFB22FEC0D9AFDEA8B3A" blockId="9.[136,1452,236,367]" pageId="9" pageNumber="38">
Similar to
<taxonomicName id="41E44D45FFE5FFB22F8F0D9AFDF98B1F" authority="var. oligocephala" box="[235,559,237,258]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE5FFB22F8F0D9AFE528B1F" box="[235,388,237,258]" italics="true" pageId="9" pageNumber="38">H. oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE5FFB22ED60D9AFDF98B1F" box="[434,559,237,258]" italics="true" pageId="9" pageNumber="38">oligocephala</emphasis>
</taxonomicName>
but with narrowly elliptic to linear-elliptic leaves up to 10(12) mm broad, 3(rarely 5)-veined, 3(5) veins prominently raised abaxially.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FEC0C42FAF48B73" pageId="9" pageNumber="38" type="materials_examined">
<paragraph id="865B36C6FFE5FFB22FEC0C42FAF48B73" blockId="9.[136,1452,236,367]" pageId="9" pageNumber="38">
<emphasis id="B490EAD4FFE5FFB22FEC0C42FF148B57" bold="true" box="[136,194,309,330]" pageId="9" pageNumber="38">Type:</emphasis>
<materialsCitation id="368C3C9BFFE5FFB22FBF0C42FAC88B73" ID-GBIF-Occurrence="3947843305" collectingDate="2008-12-09" collectorName="CC Komatipoort &amp; Bester" country="South Africa" county="Barberton District" elevation="1591" location="Farm Ameide" municipality="Barberton" pageId="9" pageNumber="38" specimenCount="1" stateProvince="Mpumalanga" typeStatus="holotype">
<collectingCountry id="FEF37656FFE5FFB22FBF0C42FE5E8B57" box="[219,392,309,330]" name="South Africa" pageId="9" pageNumber="38">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="4420F824FFE5FFB22EEB0C42FDC48B56" box="[399,530,309,331]" country="South Africa" name="Mpumalanga" pageId="9" pageNumber="38">Mpumalanga</collectingRegion>
:
<collectingCounty id="6F3A4E4AFFE5FFB22D7C0C42FD118B56" box="[536,711,309,331]" pageId="9" pageNumber="38">Barberton District</collectingCounty>
, ±
<quantity id="411C9B23FFE5FFB22DBB0C43FCF68B56" box="[735,800,308,331]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="38" unit="km" value="8.5">8.5 km</quantity>
directly SSW of
<collectingMunicipality id="663FACBCFFE5FFB22CD90C42FBF68B56" box="[957,1056,309,331]" pageId="9" pageNumber="38">Barberton</collectingMunicipality>
,
<location id="833B601DFFE5FFB22B430C42FB7E8B56" LSID="urn:lsid:plazi:treatment:0E4D87D0FFE5FFB72FEC0DD7FE778AC1:833B601DFFE5FFB22B430C42FB7E8B56" box="[1063,1192,309,331]" country="South Africa" county="Barberton District" municipality="Barberton" name="Farm Ameide" pageId="9" pageNumber="38" stateProvince="Mpumalanga">Farm Ameide</location>
717JT,
<locationDeviation id="97E92FE4FFE5FFB22B940C42FF388B72" location="Shokhohlwa Peak" pageId="9" pageNumber="38">NW of Shokhohlwa Peak</locationDeviation>
, S hill slope, grassland,
<quantity id="411C9B23FFE5FFB22EB30C2FFDF48B73" box="[471,546,344,366]" metricMagnitude="3" metricUnit="m" metricValue="1.591" pageId="9" pageNumber="38" unit="m" value="1591.0">
<elevation id="0DC9D1F5FFE5FFB22EB30C2FFDF48B73" box="[471,546,344,366]" metricMagnitude="3" metricUnit="m" metricValue="1.591" pageId="9" pageNumber="38" unit="m" value="1591.0">1591 m</elevation>
</quantity>
, (QDS: 2531
<collectorName id="2B115310FFE5FFB22DC40C2FFC938B72" box="[672,837,344,367]" pageId="9" pageNumber="38">CC Komatipoort</collectorName>
),
<date id="F25A1006FFE5FFB22C310C2EFBD78B73" box="[853,1025,345,367]" pageId="9" pageNumber="38" value="2008-12-09">
<collectingDate id="E21EE9EEFFE5FFB22C310C2EFBD78B73" box="[853,1025,345,367]" pageId="9" pageNumber="38" value="2008-12-09">9 December 2008</collectingDate>
</date>
,
<emphasis id="B490EAD4FFE5FFB22B6F0C2EFBA98B73" box="[1035,1151,345,366]" italics="true" pageId="9" pageNumber="38">
<collectorName id="2B115310FFE5FFB22B6F0C2EFB9E8B73" box="[1035,1096,345,366]" pageId="9" pageNumber="38">Bester</collectorName>
8817
</emphasis>
(
<typeStatus id="595F8864FFE5FFB22BEF0C2EFB378B72" box="[1163,1249,345,367]" pageId="9" pageNumber="38" type="holotype">holotype</typeStatus>
PRE)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FEC0CEAFBE48BAA" box="[136,1074,413,440]" pageId="9" pageNumber="38" type="etymology">
<paragraph id="865B36C6FFE5FFB22FEC0CEAFBE48BAA" blockId="9.[136,1452,413,1735]" box="[136,1074,413,440]" pageId="9" pageNumber="38">
<emphasis id="B490EAD4FFE5FFB22FEC0CEAFEDF8BAA" bold="true" box="[136,265,413,439]" pageId="9" pageNumber="38">Etymology</emphasis>
:—Named for S.P. (Pieter) Bester, colleague and diligent plant collector.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FD90CB6FF328FF2" pageId="9" pageNumber="38" type="description">
<paragraph id="865B36C6FFE5FFB22FD90CB6FF328FF2" blockId="9.[136,1452,413,1735]" pageId="9" pageNumber="38">
Slender perennial herb with few, soft, annual, herbaceous stems from small underground rootstock.
<emphasis id="B490EAD4FFE5FFB22A590CB6FA7D8BC6" box="[1341,1451,449,475]" italics="true" pageId="9" pageNumber="38">Rootstock</emphasis>
also with fusiform tubers.
<emphasis id="B490EAD4FFE5FFB22ED70C92FD9F8BE2" box="[435,585,485,511]" italics="true" pageId="9" pageNumber="38">Annual stems</emphasis>
(100)120250(270) mm high, unbranched or rarely with one or two short side branches, faintly to distinctly ribbed, densely brown or silvery/whitish hairy, hairs short-stalked, medifixed Tshaped with arms parallel to surface.
<emphasis id="B490EAD4FFE5FFB22D4F0F59FDAF885A" box="[555,633,558,583]" italics="true" pageId="9" pageNumber="38">Leaves</emphasis>
alternate, sessile or very shortly petiolate, discolorous, narrowly elliptic, linear-elliptic, sometimes elliptic, rarely narrowly obovate or obovate, (10)1245(50) × (2.0)2.510.0(12.0) mm (including petiole), apex obtuse to acute, sometimes rounded or rarely attenuate, base cuneate or shortly attenuate, margin entire, prominently 3(rarely 5)-veined from base, these raised on abaxial surface, on adaxial surface faint or sunken, rarely some reticulate veins visible between main veins, adaxial surface sparsely to more densely hairy, abaxial surface densely hairy, hairs brownish or whitish, also short-stalked, medifixed T-shaped with arms parallel to surface, similar to those on stems.
<emphasis id="B490EAD4FFE5FFB22ED40E72FDC48902" box="[432,530,773,799]" italics="true" pageId="9" pageNumber="38">Capitula</emphasis>
homogamous, discoid, few, on very short peduncles, arranged in dense, terminal, umbellate synflorescences.
<emphasis id="B490EAD4FFE5FFB22EA50E5EFDFF895E" box="[449,553,809,835]" italics="true" pageId="9" pageNumber="38">Peduncle</emphasis>
(2.0)2.514.0 mm long, densely hairy.
<emphasis id="B490EAD4FFE5FFB22C9B0E5EFBBF895E" box="[1023,1129,809,835]" italics="true" pageId="9" pageNumber="38">Involucre</emphasis>
cupuliform to campanulate, (5)
<quantity id="411C9B23FFE5FFB22FD10E3AFEF58975" box="[181,291,845,872]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="9" pageNumber="38" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">610 mm</quantity>
wide across upper part.
<emphasis id="B490EAD4FFE5FFB22D580E3AFD29897A" box="[572,767,845,871]" italics="true" pageId="9" pageNumber="38">Involucral bracts</emphasis>
arranged in 4 or 5 rows, imbricate, outer shorter than inner, densely white or silvery hairy and glandular-punctate;
<emphasis id="B490EAD4FFE5FFB22C750E05FC528996" box="[785,900,882,907]" italics="true" pageId="9" pageNumber="38">outer row</emphasis>
narrowly triangular, linear-triangular, linear or narrowly ovate, (2.5)3.04.0(5.8) × 0.51.0 mm, apex usually cuspidate, reddish;
<emphasis id="B490EAD4FFE5FFB22B060EE2FB3089B2" box="[1122,1254,917,943]" italics="true" pageId="9" pageNumber="38">second row</emphasis>
linear-triangular, narrowly triangular, narrowly ovate or narrowly obovate, (3.0)4.05.0(6.5) × (0.8)1.01.5(2.0) mm, apex attenuate or cuspidate, upper part of margin and apex usually reddish, sometimes 3-veined, visible on inside;
<emphasis id="B490EAD4FFE5FFB22BBE0EAAFA9689EA" box="[1242,1344,989,1015]" italics="true" pageId="9" pageNumber="38">third row</emphasis>
narrowly ovate or narrowly obovate, 4.05.0(5.5) × 1.01.6(2.0) mm, apex acute or attenuate, cuspidate, upper part of margin and apex usually reddish, usually 3-veined, visible on inside;
<emphasis id="B490EAD4FFE5FFB22C2D0952FC178E22" box="[841,961,1061,1087]" italics="true" pageId="9" pageNumber="38">fourth row</emphasis>
narrowly ovate, narrowly elliptic or (very) narrowly obovate, (4)56 × 1.01.5(2.2) mm, apex acute or attenuate, cuspidate, apical part of margin and apex reddish, usually 3-veined, visible on inside;
<emphasis id="B490EAD4FFE5FFB22D13091AFD008E9A" box="[631,726,1133,1159]" italics="true" pageId="9" pageNumber="38">fifth row</emphasis>
narrowly elliptic or narrowly obovate, (4.2)5.5(6.0) × 1.0
<quantity id="411C9B23FFE5FFB22AEE091AFF608EB1" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="38" unit="mm" value="1.5">1.5 mm</quantity>
, apex attenuate, cuspidate, apical part of margin and apex reddish, usually 3-veined, visible on inside.
<emphasis id="B490EAD4FFE5FFB22A5509E6FA7A8EB6" box="[1329,1452,1169,1195]" italics="true" pageId="9" pageNumber="38">Receptacle</emphasis>
epaleate.
<emphasis id="B490EAD4FFE5FFB22F9609C1FEA28ED2" box="[242,372,1205,1231]" italics="true" pageId="9" pageNumber="38">Disc florets</emphasis>
bisexual, fertile, actinomorphic; corolla pinkish or purple, tubular;
<emphasis id="B490EAD4FFE5FFB22B0D09C2FB4F8ED2" box="[1129,1177,1205,1231]" italics="true" pageId="9" pageNumber="38">tube</emphasis>
3.54.5(5.0) mm long, upper part of tube with few sessile glands and T-shaped hairs, 5-lobed;
<emphasis id="B490EAD4FFE5FFB22CD209AEFC278EEE" box="[950,1009,1241,1267]" italics="true" pageId="9" pageNumber="38">lobes</emphasis>
<quantity id="411C9B23FFE5FFB22C9D09AEFB818EE9" box="[1017,1111,1241,1268]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="38" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, apex acute, with sessile glands and T-shaped hairs but around apex with unicellular hairs.
<emphasis id="B490EAD4FFE5FFB22CE4098AFC018F0A" box="[896,983,1277,1303]" italics="true" pageId="9" pageNumber="38">Anthers</emphasis>
(1.5)2.0
<quantity id="411C9B23FFE5FFB22B31098AFB7B8F05" box="[1109,1197,1277,1304]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="38" unit="mm" value="2.5">2.5 mm</quantity>
long, with (narrowly) ovate apical appendages, bases caudate.
<emphasis id="B490EAD4FFE5FFB22D320856FD5A8F26" box="[598,652,1313,1339]" italics="true" pageId="9" pageNumber="38">Style</emphasis>
4.06.0(7.5) mm long, exserted, bifurcate;
<emphasis id="B490EAD4FFE5FFB22BE90856FAE68F26" box="[1165,1328,1313,1339]" italics="true" pageId="9" pageNumber="38">style branches</emphasis>
reddish or purple, filiform, 1.01.5(1.8) mm long, recurved, with acute hairs on outside of branches and onto the shaft below furcation, total inner surface stigmatic.
<emphasis id="B490EAD4FFE5FFB22D22081EFD3E8F9E" box="[582,744,1385,1411]" italics="true" pageId="9" pageNumber="38">Ovary/cypsela</emphasis>
brown, ± obconical, 1.0
<quantity id="411C9B23FFE5FFB22B68081EFBB48F99" box="[1036,1122,1385,1412]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="38" unit="mm" value="1.8">1.8 mm</quantity>
long, densely twin hairy and glandular punctate, with distinct carpopodium.
<taxonomicName id="41E44D45FFE5FFB22DFA08F9FD248FBA" box="[670,754,1422,1447]" class="Insecta" family="Miridae" genus="Pappus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B490EAD4FFE5FFB22DFA08F9FD248FBA" box="[670,754,1422,1447]" italics="true" pageId="9" pageNumber="38">Pappus</emphasis>
</taxonomicName>
dimorphic, an outer row of laciniate scales, 0.51.0 mm long and inner row of barbellate bristles
<quantity id="411C9B23FFE5FFB22D7C08C6FDA38FD1" box="[536,629,1457,1484]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="9" pageNumber="38" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long.
<emphasis id="B490EAD4FFE5FFB22DD808C6FCBE8FD6" box="[700,872,1457,1483]" italics="true" pageId="9" pageNumber="38">Flowering time</emphasis>
: (August)September to December. (
<figureCitation id="1EDF2A43FFE5FFB22B9B08C6FAAE8FD6" box="[1279,1400,1457,1483]" captionStart="FIGURE 1" captionStartId="2.[136,229,1566,1588]" captionTargetBox="[151,1436,190,1542]" captionTargetId="figure-20@2.[151,1436,190,1542]" captionTargetPageId="2" captionText="FIGURE 1. Digital images illustrating the habit and synflorescences of Hilliardiella elaeagnoides and H. oligocephala var. besteri. A. Hilliardiella elaeagnoides growing in the Pretoria National Botanical Garden. Photographer M. Koekemoer. B. H. oligocephala var. besteri growing in Mpumalanga near the eSwatini border. Photographer D. Oosthuizen. C. The few-headed, umbellate synflorescence of H. oligocephala var. besteri growing on the farm Schultzenhorst 718 JT, Barberton District (Mpumalanga). Photographer D. Oosthuizen." figureDoi="http://doi.org/10.5281/zenodo.7251468" httpUri="https://zenodo.org/record/7251468/files/figure.png" pageId="9" pageNumber="38">Fig. 1B, C</figureCitation>
,
<figureCitation id="1EDF2A43FFE5FFB22AE008C6FF6C8FED" captionStart="FIGURE 4" captionStartId="7.[136,229,1439,1461]" captionTargetBox="[151,1436,190,1415]" captionTargetId="figure-16@7.[151,1436,190,1415]" captionTargetPageId="7" captionText="FIGURE 4. Leaves and involucral bracts of Hilliardiella oligocephala. A. Abaxial surface of leaves of H. oligocephala var. oligocephala (Bester 8832, PRE) showing the pinnate veins(Nikondigital camera,photographer P.Herman).B.Abaxial surface of leaf of H.oligocephala var. oligocephala (Medley Wood 5755, PRE) showing the dense hairs and pinnate veins (Olympus digital microscope). C.Abaxial surface of the leaf of H. oligocephala var. besteri (Smuts &amp; Gillett 2406, PRE) showing the 3 main veins (Nikon digital camera, photographer P. Herman). D. Abaxial surface of the leaf of H. oligocephala var. besteri (Smuts &amp; Gillett 2406, PRE) showing the dense hairs and the 3 main veins (Olympus digital microscope). E. Inner surface of the inner involucral bract of H. oligocephala var. oligocephala (Codd 1796, PRE) showing the 3 main veins (Olympus digital microscope). F. Inner surface of the inner involucral bract of H. oligocephala var. besteri (Compton 26095, PRE) showing the 3 main veins (Olympus digital microscope)." figureDoi="http://doi.org/10.5281/zenodo.7251476" httpUri="https://zenodo.org/record/7251476/files/figure.png" pageId="9" pageNumber="38">4C, D, F</figureCitation>
,
<figureCitation id="1EDF2A43FFE5FFB22FA208A2FF008FF2" box="[198,214,1493,1519]" captionStart="FIGURE 6" captionStartId="10.[136,229,2028,2050]" captionTargetBox="[211,1375,190,2004]" captionTargetId="figure-20@10.[211,1375,190,2004]" captionTargetPageId="10" captionText="FIGURE 6. Scan of the holotype of Hilliardiella oligocephala var. besteri (Bester 8817, PRE)." figureDoi="http://doi.org/10.5281/zenodo.7251480" httpUri="https://zenodo.org/record/7251480/files/figure.png" pageId="9" pageNumber="38">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FD9088EFAE38C62" pageId="9" pageNumber="38" type="diagnosis">
<paragraph id="865B36C6FFE5FFB22FD9088EFAE38C62" blockId="9.[136,1452,413,1735]" pageId="9" pageNumber="38">
<emphasis id="B490EAD4FFE5FFB22FD9088EFE1E8C0E" bold="true" box="[189,456,1529,1555]" pageId="9" pageNumber="38">Diagnostic characters</emphasis>
:—
<taxonomicName id="41E44D45FFE5FFB22E8F088EFC7F8C0E" authorityName="P. P. J. Herman" authorityYear="2022" box="[491,937,1529,1556]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFE5FFB22E8F088EFCC18C0E" box="[491,791,1529,1555]" italics="true" pageId="9" pageNumber="38">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE5FFB22C39088EFC7F8C0E" box="[861,937,1529,1555]" italics="true" pageId="9" pageNumber="38">besteri</emphasis>
</taxonomicName>
has few, soft, slender, annual, herbaceous stems only up to 250(270) mm high from the underground rootstock, narrowly elliptic to linear-elliptic, abaxially prominently raised 3(rarely 5)-veined leaves up to 10(12) mm broad.
<taxonomicName id="41E44D45FFE5FFB22CF20B36FA568C46" authority="var. oligocephala" box="[918,1408,1601,1628]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="oligocephala">
<emphasis id="B490EAD4FFE5FFB22CF20B36FB6F8C46" box="[918,1209,1601,1627]" italics="true" pageId="9" pageNumber="38">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE5FFB22B8A0B36FA568C46" box="[1262,1408,1601,1627]" italics="true" pageId="9" pageNumber="38">oligocephala</emphasis>
</taxonomicName>
has narrowly or broadly ovate, cordate, elliptic or broadly elliptic, prominently raised, pinnately veined leaves.
</paragraph>
</subSubSection>
<subSubSection id="CEFE654DFFE5FFB22FD90BFEFD088CDA" pageId="9" pageNumber="38" type="distribution">
<paragraph id="865B36C6FFE5FFB22FD90BFEFD088CDA" blockId="9.[136,1452,413,1735]" pageId="9" pageNumber="38">
<emphasis id="B490EAD4FFE5FFB22FD90BFEFE998CBE" bold="true" box="[189,335,1673,1699]" pageId="9" pageNumber="38">Distribution</emphasis>
:—
<taxonomicName id="41E44D45FFE5FFB22E170BFEFCCF8CBE" authorityName="P. P. J. Herman" authorityYear="2022" box="[371,793,1673,1700]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="38" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFE5FFB22E170BFEFD418CBE" box="[371,663,1673,1699]" italics="true" pageId="9" pageNumber="38">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE5FFB22DA90BFEFCCF8CBE" box="[717,793,1673,1699]" italics="true" pageId="9" pageNumber="38">besteri</emphasis>
</taxonomicName>
is near endemic to Mpumalanga and occurs in a small area in Mpumalanga and in
<collectingCountry id="FEF37656FFE5FFB22EE80BDAFE388CDA" box="[396,494,1709,1735]" name="Swaziland" pageId="9" pageNumber="38">eSwatini</collectingCountry>
(
<collectingCountry id="FEF37656FFE5FFB22E990BDAFDA38CDA" box="[509,629,1709,1735]" name="Swaziland" pageId="9" pageNumber="38">Swaziland</collectingCountry>
) (
<figureCitation id="1EDF2A43FFE5FFB22DEF0BDAFD068CDA" box="[651,720,1709,1735]" captionStart="FIGURE 7" captionStartId="11.[136,229,1291,1313]" captionTargetBox="[172,1420,215,1252]" captionTargetId="figure-16@11.[151,1436,190,1267]" captionTargetPageId="11" captionText="FIGURE 7. Distribution map of Hilliardiella oligocephala var. besteri in southern Africa." figureDoi="http://doi.org/10.5281/zenodo.7251482" httpUri="https://zenodo.org/record/7251482/files/figure.png" pageId="9" pageNumber="38">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D29B664EFFE6FFB12FEC0A9BFBCB821F" ID-DOI="http://doi.org/10.5281/zenodo.7251480" ID-Zenodo-Dep="7251480" box="[136,1053,2028,2050]" httpUri="https://zenodo.org/record/7251480/files/figure.png" pageId="10" pageNumber="39" startId="10.[136,229,2028,2050]" targetBox="[211,1375,190,2004]" targetPageId="10">
<paragraph id="865B36C6FFE6FFB12FEC0A9BFBCB821F" blockId="10.[136,1053,2028,2050]" box="[136,1053,2028,2050]" pageId="10" pageNumber="39">
<emphasis id="B490EAD4FFE6FFB12FEC0A9BFF21821C" bold="true" box="[136,247,2028,2050]" pageId="10" pageNumber="39">FIGURE 6</emphasis>
. Scan of the holotype of
<taxonomicName id="41E44D45FFE6FFB12E8D0A9BFC82821C" authorityName="P. P. J. Herman" authorityYear="2022" box="[489,852,2028,2050]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="39" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFE6FFB12E8D0A9BFD35821C" box="[489,739,2028,2049]" italics="true" pageId="10" pageNumber="39">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE6FFB12C770A9BFC82821C" box="[787,852,2028,2049]" italics="true" pageId="10" pageNumber="39">besteri</emphasis>
</taxonomicName>
(
<emphasis id="B490EAD4FFE6FFB12C060A9AFC00821F" box="[866,982,2029,2050]" italics="true" pageId="10" pageNumber="39">Bester 8817</emphasis>
, PRE).
</paragraph>
</caption>
<caption id="D29B664EFFE7FFB02FEC087CFC3A8F3C" ID-DOI="http://doi.org/10.5281/zenodo.7251482" ID-Zenodo-Dep="7251482" box="[136,1004,1291,1313]" httpUri="https://zenodo.org/record/7251482/files/figure.png" pageId="11" pageNumber="40" startId="11.[136,229,1291,1313]" targetBox="[172,1420,215,1252]" targetPageId="11">
<paragraph id="865B36C6FFE7FFB02FEC087CFC3A8F3C" blockId="11.[136,1004,1291,1313]" box="[136,1004,1291,1313]" pageId="11" pageNumber="40">
<emphasis id="B490EAD4FFE7FFB02FEC087CFF218F3C" bold="true" box="[136,247,1291,1313]" pageId="11" pageNumber="40">FIGURE 7</emphasis>
. Distribution map of
<taxonomicName id="41E44D45FFE7FFB02EA3087CFCE78F3D" authorityName="P. P. J. Herman" authorityYear="2022" box="[455,817,1291,1313]" class="Magnoliopsida" family="Asteraceae" genus="Hilliardiella" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="40" phylum="Tracheophyta" rank="variety" species="oligocephala" variety="besteri">
<emphasis id="B490EAD4FFE7FFB02EA3087CFD178F3D" box="[455,705,1291,1312]" italics="true" pageId="11" pageNumber="40">Hilliardiella oligocephala</emphasis>
var.
<emphasis id="B490EAD4FFE7FFB02D94087CFCE78F3D" box="[752,817,1291,1312]" italics="true" pageId="11" pageNumber="40">besteri</emphasis>
</taxonomicName>
in southern Africa.
</paragraph>
</caption>
<subSubSection id="CEFE654DFFE7FFB72FEC08FBFE778AC1" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40" type="materials_examined">
<paragraph id="865B36C6FFE7FFB02FEC08FBFDC38FBB" blockId="11.[136,533,1420,1446]" box="[136,533,1420,1446]" pageId="11" pageNumber="40">
<heading id="DD1381AAFFE7FFB02FEC08FBFDC38FBB" bold="true" box="[136,533,1420,1446]" fontSize="11" level="1" pageId="11" pageNumber="40" reason="1">
<emphasis id="B490EAD4FFE7FFB02FEC08FBFDC38FBB" bold="true" box="[136,533,1420,1446]" pageId="11" pageNumber="40">Additional specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="865B36C6FFE7FFB02FED08A0FD858C9C" blockId="11.[136,1452,1494,2025]" pageId="11" pageNumber="40">
<collectingCountry id="FEF37656FFE7FFB02FED08A0FEE28FEC" box="[137,308,1495,1521]" name="Swaziland" pageId="11" pageNumber="40">SWAZILAND</collectingCountry>
[eSWATINI]. Havelock Mine, grassland,
<quantity id="411C9B23FFE7FFB02C7408A0FC898FED" box="[784,863,1494,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.3106399999999998" pageId="11" pageNumber="40" unit="ft" value="4300.0">4300 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02C0908A0FC118FEC" box="[877,967,1495,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.311" pageId="11" pageNumber="40" unit="m" value="1311.0">1311 m</quantity>
], (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02A5008A1FF158C08" pageId="11" pageNumber="40" value="1955-09">September 1955</date>
,
<emphasis id="B490EAD4FFE7FFB02FAA088DFE858C08" box="[206,339,1530,1557]" italics="true" pageId="11" pageNumber="40">Miller 3013</emphasis>
(PRE). Havelock Mine, grassland,
<quantity id="411C9B23FFE7FFB02D86088CFCE58C09" box="[738,819,1530,1557]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="11" pageNumber="40" unit="ft" value="4000.0">4000 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02C25088CFC4A8C08" box="[833,924,1531,1557]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="11" pageNumber="40" unit="m" value="1219.0">1219 m</quantity>
], (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02A6F088DFA718C08" box="[1291,1447,1530,1557]" pageId="11" pageNumber="40" value="1955-10">October 1955</date>
,
<emphasis id="B490EAD4FFE7FFB02FEC0B69FEDB8C24" box="[136,269,1566,1593]" italics="true" pageId="11" pageNumber="40">Miller 3057</emphasis>
(PRE). Mbabane District, Waverley Mine,
<quantity id="411C9B23FFE7FFB02D950B69FC978C25" box="[753,833,1566,1593]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="40" unit="ft" value="5000.0">5000 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02C2A0B69FC7F8C24" box="[846,937,1566,1593]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="11" pageNumber="40" unit="m" value="1524.0">1524 m</quantity>
], (QDS: 2630BD Carolina),
<date id="F25A1006FFE7FFB02B8D0B68FA718C24" box="[1257,1447,1566,1593]" pageId="11" pageNumber="40" value="1956-10-17">17 October 1956</date>
,
<emphasis id="B490EAD4FFE7FFB02FEC0B34FEE98C41" box="[136,319,1602,1629]" italics="true" pageId="11" pageNumber="40">Compton 26095</emphasis>
(PRE). Mbabane District, Ngwenya Mountain,
<quantity id="411C9B23FFE7FFB02C0E0B35FC6B8C41" box="[874,957,1602,1629]" metricMagnitude="3" metricUnit="m" metricValue="1.6764" pageId="11" pageNumber="40" unit="ft" value="5500.0">5500 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02CAA0B35FBFB8C40" box="[974,1069,1602,1629]" metricMagnitude="3" metricUnit="m" metricValue="1.6759999999999997" pageId="11" pageNumber="40" unit="m" value="1676.0">1676 m</quantity>
], (QDS: 2632AA Bela Vista),
<date id="F25A1006FFE7FFB02AEB0B34FE968C9C" pageId="11" pageNumber="40" value="1956-11-18">18 November 1956</date>
,
<emphasis id="B490EAD4FFE7FFB02E2F0B10FE298C9D" box="[331,511,1638,1665]" italics="true" pageId="11" pageNumber="40">Compton 26332</emphasis>
(PRE).
</paragraph>
<paragraph id="865B36C6FFE7FFB02FD90BFCFDC28DF4" blockId="11.[136,1452,1494,2025]" pageId="11" pageNumber="40">
<collectingCountry id="FEF37656FFE7FFB02FD90BFCFE438CB8" box="[189,405,1675,1701]" name="South Africa" pageId="11" pageNumber="40">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="4420F824FFE7FFB02EC30BFDFDE98CB9" box="[423,575,1674,1700]" country="South Africa" name="Mpumalanga" pageId="11" pageNumber="40">Mpumalanga</collectingRegion>
: Pilgrims Rest District, Sabie, Mauchsberg,
<quantity id="411C9B23FFE7FFB02B2A0BFDFB3E8CB9" box="[1102,1256,1674,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.61544" metricValueMax="1.76784" metricValueMin="1.46304" pageId="11" pageNumber="40" unit="ft" value="5300.0" valueMax="5800.0" valueMin="4800.0">48005800 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02B9D0BFDFA498CB8" box="[1273,1439,1674,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.6155" metricValueMax="1.768" metricValueMin="1.463" pageId="11" pageNumber="40" unit="m" value="1615.5" valueMax="1768.0" valueMin="1463.0">14631768 m</quantity>
], (QDS: 2530BA Lydenburg),
<date id="F25A1006FFE7FFB02EAA0BD8FD708CD4" box="[462,678,1710,1737]" pageId="11" pageNumber="40" value="1932-12-22">22 December 1932</date>
,
<emphasis id="B490EAD4FFE7FFB02DD50BD8FC498CD4" box="[689,927,1710,1737]" italics="true" pageId="11" pageNumber="40">Smuts &amp; Gillett 2271</emphasis>
(PRE),
<emphasis id="B490EAD4FFE7FFB02C9F0BD8FB3F8CD4" box="[1019,1257,1710,1737]" italics="true" pageId="11" pageNumber="40">Smuts &amp; Gillett 2329</emphasis>
(PRE). Pilgrims Rest District, Mount Anderson,
<quantity id="411C9B23FFE7FFB02E910BA4FD908CF1" box="[501,582,1746,1773]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="11" pageNumber="40" unit="ft" value="7000.0">7000 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02D310BA4FD658CF0" box="[597,691,1747,1773]" metricMagnitude="3" metricUnit="m" metricValue="2.134" pageId="11" pageNumber="40" unit="m" value="2134.0">2134 m</quantity>
], (QDS: 2530BA Lydenburg),
<date id="F25A1006FFE7FFB02B760BA4FB3B8CF0" box="[1042,1261,1746,1773]" pageId="11" pageNumber="40" value="1932-12-24">24 December 1932</date>
,
<emphasis id="B490EAD4FFE7FFB02B9E0BA4FF168D0D" italics="true" pageId="11" pageNumber="40">Smuts &amp; Gillett 2406</emphasis>
(PRE). Barberton District, ±
<quantity id="411C9B23FFE7FFB02D7D0B80FDB48D0D" box="[537,610,1782,1809]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="40" unit="km" value="11.0">11 km</quantity>
E of Barberton, Farm Twello, Sappi Plantation, moderate NW mountain slope, grassland,
<quantity id="411C9B23FFE7FFB02E2C0A6CFE498D28" box="[328,415,1819,1845]" metricMagnitude="3" metricUnit="m" metricValue="1.449" pageId="11" pageNumber="40" unit="m" value="1449.0">1449 m</quantity>
, (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02C610A6CFC368D28" box="[773,992,1818,1845]" pageId="11" pageNumber="40" value="2014-11-13">13 November 2014</date>
,
<emphasis id="B490EAD4FFE7FFB02C8F0A6CFBA98D28" box="[1003,1151,1819,1845]" italics="true" pageId="11" pageNumber="40">Bester 12085</emphasis>
(PRE). Barberton District, at turn-off from R40 between Barberton and Josefsdal towards Diepgezet mine, ±
<quantity id="411C9B23FFE7FFB02B5F0A48FB528D45" box="[1083,1156,1854,1881]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="40" unit="km" value="18.0">18 km</quantity>
SSE of Barberton, gentle NW mountain slope, grassland,
<quantity id="411C9B23FFE7FFB02E990A15FD818D60" box="[509,599,1890,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="11" pageNumber="40" unit="m" value="1455.0">1455 m</quantity>
, (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02CAF0A15FB788D60" box="[971,1198,1890,1917]" pageId="11" pageNumber="40" value="2017-11-26">26 November 2017</date>
,
<emphasis id="B490EAD4FFE7FFB02BD80A14FA828D60" box="[1212,1364,1891,1917]" italics="true" pageId="11" pageNumber="40">Bester 13858</emphasis>
(PRE). Barberton District, Oosterbeek farm,
<quantity id="411C9B23FFE7FFB02D4C0AF1FDB68DBD" box="[552,608,1926,1953]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="40" unit="km" value="5.0">5 km</quantity>
S of Barberton, grassland,
<quantity id="411C9B23FFE7FFB02CEB0AF1FC088DBD" box="[911,990,1926,1953]" metricMagnitude="3" metricUnit="m" metricValue="1.58496" pageId="11" pageNumber="40" unit="ft" value="5200.0">5200 ft</quantity>
[
<quantity id="411C9B23FFE7FFB02C8F0AF1FB908DBC" box="[1003,1094,1926,1953]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="11" pageNumber="40" unit="m" value="1585.0">1585 m</quantity>
], (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02FEC0ADDFEB28DD8" box="[136,356,1962,1989]" pageId="11" pageNumber="40" value="1977-11-15">15 November 1977</date>
,
<emphasis id="B490EAD4FFE7FFB02E0B0ADCFDDD8DD9" box="[367,523,1962,1989]" italics="true" pageId="11" pageNumber="40">De Sousa 631</emphasis>
(PRE). Barberton, Angle station, hill above station, (QDS: 2531CC Komatipoort),
<date id="F25A1006FFE7FFB02FEC0AB8FEF88DF4" box="[136,302,1999,2025]" pageId="11" pageNumber="40" value="1978-08-02">2 August 1978</date>
,
<emphasis id="B490EAD4FFE7FFB02E5D0AB8FE168DF4" box="[313,448,1999,2025]" italics="true" pageId="11" pageNumber="40">Venter 4491</emphasis>
(PRE).
</paragraph>
<paragraph id="865B36C6FFE0FFB72FD90DE9FE778AC1" blockId="12.[136,1452,158,221]" pageId="12" pageNumber="41">
Locality not found:
<collectingRegion id="4420F824FFE0FFB72EF00DE9FDFD8AA5" box="[404,555,158,184]" country="South Africa" name="Mpumalanga" pageId="12" pageNumber="41">Mpumalanga</collectingRegion>
, Barberton, mountain tops near Boschs,
<quantity id="411C9B23FFE0FFB72C830DE9FBAC8AA5" box="[999,1146,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.2954" metricValueMax="1.3716" metricValueMin="1.2192" pageId="12" pageNumber="41" unit="ft" value="4250.0" valueMax="4500.0" valueMin="4000.0">40004500 ft</quantity>
[
<quantity id="411C9B23FFE0FFB72BE00DE9FAF58AA4" box="[1156,1315,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.2955" metricValueMax="1.3719999999999999" metricValueMin="1.219" pageId="12" pageNumber="41" unit="m" value="1295.5" valueMax="1372.0" valueMin="1219.0">12191372 m</quantity>
], September 1889,
<emphasis id="B490EAD4FFE0FFB72FAA0DB5FE9B8AC0" box="[206,333,194,221]" italics="true" pageId="12" pageNumber="41">Galpin 557</emphasis>
(PRE).
</paragraph>
</subSubSection>
</treatment>
</document>