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<document id="E9F9AF14517021A5ACED5482F00A7007" ID-DOI="10.5281/zenodo.276822" ID-GBIF-Dataset="f5489381-8fe8-48a0-9fb6-5addeebaa3f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="276822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237113383" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03C687E4FFD5FF9EFF1BEE641296FBC5" docLanguage="en" docName="zt02763p033.pdf" docOrigin="Zootaxa 2763" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hlavaciellus adelphos Jałoszyński, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="FFFFFF9CFFDFFF95FF8CEB58101EFFCA" masterDocTitle="Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="11" updateTime="1698239820214" updateUser="plazi">
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<mods:title id="E566DEB11FAA8E8C0682088C2E4EDED3">Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="C470E19CEDD41DBB8DC1EE50FC7544F2">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="2CE55666FEE97351C99AC3E9C18D0CE0">2011</mods:date>
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<treatment id="03C687E4FFD5FF9EFF1BEE641296FBC5" ID-DOI="http://doi.org/10.5281/zenodo.6192605" ID-GBIF-Taxon="119388733" ID-Zenodo-Dep="6192605" LSID="urn:lsid:plazi:treatment:03C687E4FFD5FF9EFF1BEE641296FBC5" httpUri="http://treatment.plazi.org/id/03C687E4FFD5FF9EFF1BEE641296FBC5" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="C3756579FFD5FF9FFF1BEE641146FABC" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BD036F2FFD5FF9FFF1BEE6411F4FA9F" blockId="10.[151,490,1339,1398]" box="[151,490,1339,1366]" pageId="10" pageNumber="11">
<heading id="D098819EFFD5FF9FFF1BEE6411F4FA9F" bold="true" box="[151,490,1339,1366]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B91BEAE0FFD5FF9FFF1BEE6411F4FA9F" bold="true" box="[151,490,1339,1366]" pageId="10" pageNumber="11">
<taxonomicName id="4C6F4D71FFD5FF9FFF1BEE6411BEFA9C" ID-CoL="3M5PN" box="[151,416,1340,1366]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="adelphos" status="sp. nov.">
<emphasis id="B91BEAE0FFD5FF9FFF1BEE6411BEFA9C" bold="true" box="[151,416,1340,1366]" italics="true" pageId="10" pageNumber="11">Hlavaciellus adelphos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579BFFD5FF9FFE24EE6311F4FA9F" box="[424,490,1339,1365]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F2FFD5FF9FFF1BEE061146FABC" blockId="10.[151,490,1339,1398]" box="[151,344,1374,1398]" pageId="10" pageNumber="11">
(
<figureCitation id="13542A77FFD5FF9FFF13EE0610E3FABC" box="[159,253,1374,1398]" captionStart="FIGURES 10 12" captionStartId="8.[151,264,1095,1118]" captionTargetBox="[161,1425,204,1060]" captionTargetId="figure@8.[151,1436,193,1074]" captionTargetPageId="8" captionText="FIGURES 10 12. Dorsal view of holotypes of Hlavaciellus, males. 10, H. decoratus sp. n. 11, H. smetanai sp. n. 12, H. adelphos sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276825/files/figure.png" pageId="10" pageNumber="11">Figs. 12</figureCitation>
, 29, 30)
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD5FF9FFF1BEEF0128FF9CD" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BD036F2FFD5FF9FFF1BEEF0128FF9CD" blockId="10.[151,1436,1447,2012]" pageId="10" pageNumber="11">
<emphasis id="B91BEAE0FFD5FF9FFF1BEEF01274FA0A" bold="true" box="[151,618,1447,1472]" pageId="10" pageNumber="11">
<typeStatus id="54D48850FFD5FF9FFF1BEEF010CCFA0A" box="[151,210,1448,1472]" pageId="10" pageNumber="11">Type</typeStatus>
material.
<typeStatus id="54D48850FFD5FF9FFEC1EEF011A0FA0A" box="[333,446,1448,1472]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3787662FFD5FF9FFE47EEFF127BFA0A" box="[459,613,1447,1472]" name="Malaysia" pageId="10" pageNumber="11">MALAYSIA</collectingCountry>
:
</emphasis>
ɗ, &quot;
<collectionCode id="ED7EAE37FFD5FF9FFD11EEFF131DFA75" box="[669,771,1447,1471]" pageId="10" pageNumber="11">SABAH</collectionCode>
: Crocker Ra. \
<quantity id="4C979B17FFD5FF9FFC22EEFF1418FA75" box="[942,1030,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="11" unit="m" value="1600.0">1600 m</quantity>
, km 51 rte \ Kota Kinabalu \ Tambunan,
<date id="FFD11032FFD5FF9FFF65EE941142FA2E" box="[233,348,1484,1508]" pageId="10" pageNumber="11" value="1987-05-18">18.V.1987</date>
\ Burckhardt - Löbl&quot; [white, printed], &quot;
<taxonomicName id="4C6F4D71FFD5FF9FFC9BEE9413C7FA2E" box="[791,985,1484,1508]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE0FFD5FF9FFC9BEE9413C7FA2E" box="[791,985,1484,1508]" italics="true" pageId="10" pageNumber="11">HLAVACIELLUS</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6F4D71FFD5FF9FFC7CEE94144BFA2E" box="[1008,1109,1484,1508]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="adelphos">
<emphasis id="B91BEAE0FFD5FF9FFC7CEE94144BFA2E" box="[1008,1109,1484,1508]" italics="true" pageId="10" pageNumber="11">adelphos</emphasis>
</taxonomicName>
m. \ det. P. JAŁOSZYŃSKI, '09 \
<typeStatus id="54D48850FFD5FF9FFF42EEB71169F9CD" box="[206,375,1519,1543]" pageId="10" pageNumber="11" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED7EAE37FFD5FF9FFDA5EEB7129BF9CD" box="[553,645,1519,1543]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="11">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD5FF9FFF4BED4B14FCF985" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BD036F2FFD5FF9FFF4BED4B14FCF985" blockId="10.[151,1436,1447,2012]" pageId="10" pageNumber="11">
<emphasis id="B91BEAE0FFD5FF9FFF4BED4B115FF9E6" bold="true" box="[199,321,1555,1580]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
Sides of vertex in males delimited from frons and supraantennal tubercles by pair of shallow but distinct transverse lateral impressions narrowly separated in middle; antennomere III subquadrate.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD5FF9EFF4BED0311BCFCB5" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="8BD036F2FFD5FF9FFF4BED031115F95D" blockId="10.[151,1436,1447,2012]" pageId="10" pageNumber="11">
<emphasis id="B91BEAE0FFD5FF9FFF4BED031147F9BE" bold="true" box="[199,345,1627,1652]" pageId="10" pageNumber="11">Description.</emphasis>
BL 1.83. Body of male (
<figureCitation id="13542A77FFD5FF9FFDE2ED0412A0F9BE" box="[622,702,1628,1652]" captionStart="FIGURES 10 12" captionStartId="8.[151,264,1095,1118]" captionTargetBox="[161,1425,204,1060]" captionTargetId="figure@8.[151,1436,193,1074]" captionTargetPageId="8" captionText="FIGURES 10 12. Dorsal view of holotypes of Hlavaciellus, males. 10, H. decoratus sp. n. 11, H. smetanai sp. n. 12, H. adelphos sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276825/files/figure.png" pageId="10" pageNumber="11">Fig. 12</figureCitation>
) strongly convex, pigmentation moderately light brown, vestiture yellowish.
</paragraph>
<paragraph id="8BD036F2FFD5FF9EFF4BEDFC1513FF1E" blockId="10.[151,1436,1447,2012]" lastBlockId="11.[151,1437,151,1040]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Head large, HL 0.30, HW 0.48; vertex convex, separated from supraantennal tubercles and lateral parts of frons by pair of very shallow transverse impressions narrowly interrupted in middle; frons flattened; clypeus convex; supraantennal tubercles prominent and sharply delimited from median part of frons; eyes large, strongly convex and coarsely faceted. Punctures on dorsal surface of head very fine but distinct, unevenly distributed, separated by spaces 1
<date id="FFD11032FFD5FF9FFEAFEC6C112CF886" box="[291,306,1844,1868]" pageId="10" pageNumber="11">2</date>
x as wide as puncture diameters; setae short, relatively dense, suberect. Antennae as long as 0.71 BL, moderately slender, gradually but only slightly thickened up to antennomere XI, uniformly covered with thin, long, dense and suberect setae, AnL 1.30, antennomere I 1.5x as long as broad; II distinctly narrower and shorter than I, subquadrate; IIIII nearly equal in length and width, each slightly larger than II, 1.1x as long as broad; V slightly larger than IV, 1.2x as long as broad; VI slightly larger than V, 1.3x as long as broad; VIIVIII subequal in length and width, each slightly larger than VI and 1.3x as long as broad; IX slightly larger than VIII, 1.2x as long as broad; X slightly larger than IX, 1.1x as long as broad; XI as broad as X, slightly shorter than IXX together.
</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BEBB81237FE05" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="12">
Pronotum in dorsal view subrectangular with weakly convex anterior margin, broadest at base and in anterior third, PL 0.53, PW 0.75; sides strongly rounded in anterior fourth, weakly concave just behind middle; hind angles right and blunt; posterior margin shallowly bi-emarginate; ante-basal pits shallow but well defined, each located slightly closer to posterior than to lateral margin of pronotum; lateral edges not swollen. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 1.5
<date id="FFD11032FFD4FF9EFB95EA371439FE4D" box="[1049,1063,367,391]" pageId="11" pageNumber="12">2</date>
x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, those near posterior angles appear coarse. Setae long, moderately dense, suberect to erect.
</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BEA841267FD06" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="12">
Elytra oval, relatively short, broadest between middle and anterior third, EL 1.00, EW 0.88, EI 1.14; basal pit on each elytron small but distinct, located very close to scutellum; subhumeral lines short but distinct, as long as 0.35x EL, only slightly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron much more distinct than those on pronotum but very shallow, separated by spaces 11.5x as wide as puncture diameters, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum but slightly thicker and distinctly longer.
<collectingCountry id="F3787662FFD4FF9EFEB5E9EC116FFD06" box="[313,369,692,716]" name="India" pageId="11" pageNumber="12">Hind</collectingCountry>
wings fully developed.
</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BE9801291FD25" blockId="11.[151,1437,151,1040]" box="[199,655,727,752]" pageId="11" pageNumber="12">Legs slender and long, all tibiae straight.</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BE9A4151FFCDE" blockId="11.[151,1437,151,1040]" box="[199,1281,764,788]" pageId="11" pageNumber="12">Metaventrite with large and deep postmesocoxal impressions, occupying about half of its length.</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BE8471445FC96" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="12">Aedeagus (Figs. 29, 30) very slender, AeL 0.48; median lobe with short, trapezoidal apical part; internal armature nearly symmetrical; parameres in lateral view broad, with strongly curved apices.</paragraph>
<paragraph id="8BD036F2FFD4FF9EFF4BE83011BCFCB5" blockId="11.[151,1437,151,1040]" box="[199,418,871,896]" pageId="11" pageNumber="12">
<emphasis id="B91BEAE0FFD4FF9EFF4BE830113BFC4A" bold="true" box="[199,293,872,896]" pageId="11" pageNumber="12">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD4FF9EFF4BE8D312DEFC6E" box="[199,704,907,932]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BD036F2FFD4FF9EFF4BE8D312DEFC6E" blockId="11.[151,1437,151,1040]" box="[199,704,907,932]" pageId="11" pageNumber="12">
<emphasis id="B91BEAE0FFD4FF9EFF4BE8D3117EFC6E" bold="true" box="[199,352,907,932]" pageId="11" pageNumber="12">Distribution.</emphasis>
East
<collectingCountry id="F3787662FFD4FF9EFE12E8D41217FC6E" box="[414,521,908,932]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
: Borneo, Sabah.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD4FF9EFF4BE8E814E5FC26" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BD036F2FFD4FF9EFF4BE8E814E5FC26" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="12">
<emphasis id="B91BEAE0FFD4FF9EFF4BE8E81150FC02" bold="true" box="[199,334,944,968]" pageId="11" pageNumber="12">Etymology.</emphasis>
The Greek noun αδελφός (
<taxonomicName id="4C6F4D71FFD4FF9EFD0FE8F712F2FC0D" box="[643,748,943,967]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="adelphos">adelphos</taxonomicName>
), meaning &quot;a brother&quot;, was chosen to underline a close similarity of this species to other members of the genus showing the same
<typeStatus id="54D48850FFD4FF9EFC25E88D13C7FC27" box="[937,985,981,1005]" pageId="11" pageNumber="12">type</typeStatus>
of the head modification.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFD4FF9EFF4BE8A01296FBC5" box="[199,648,1015,1040]" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BD036F2FFD4FF9EFF4BE8A01296FBC5" blockId="11.[151,1437,151,1040]" box="[199,648,1015,1040]" pageId="11" pageNumber="12">
<emphasis id="B91BEAE0FFD4FF9EFF4BE8A01125FBDA" bold="true" box="[199,315,1016,1040]" pageId="11" pageNumber="12">Remarks.</emphasis>
See remarks for
<taxonomicName id="4C6F4D71FFD4FF9EFE76E8A1129FFBDA" box="[506,641,1017,1040]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91BEAE0FFD4FF9EFE76E8A1129FFBDA" box="[506,641,1017,1040]" italics="true" pageId="11" pageNumber="12">H. smetanai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>