treatments-xml/data/03/C6/87/03C687E2FFF6EE00F0CB5144DBF81C88.xml
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<document id="E7A3C9AF3B227648B9CB1B9AE1A4E80B" ID-CLB-Dataset="8712" ID-DOI="10.11646/zootaxa.4908.4.5" ID-GBIF-Dataset="6925e575-3634-4492-8be6-b422b73fe888" ID-ISSN="1175-5326" ID-Zenodo-Dep="4447214" ID-ZooBank="FD97747A-BF20-4820-B35E-67B432FF9987" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610980108797" checkinUser="plazi" docAuthor="Sikorski, Andrey V., Pavlova, Lyudmila V., Sardá, Rafael, Langeneck, Joachim, Gil, João &amp; Ravara, Ascensão" docDate="2021" docId="03C687E2FFF6EE00F0CB5144DBF81C88" docLanguage="en" docName="zootaxa4908.4.5.pdf" docOrigin="Zootaxa 4908 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laonice (Appelloefia) barcinensis Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="521" masterDocId="FFFFFF9AFFF2EE07F05C5600DA261D01" masterDocTitle="Two new deep-sea species of Laonice (Annelida: Spionidae) from the Mediterranean Sea" masterLastPageNumber="526" masterPageNumber="515" pageNumber="519" updateTime="1698924590607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7B23CBD370EC8968D29A3A5881EB9FAD">Two new deep-sea species of Laonice (Annelida: Spionidae) from the Mediterranean Sea</mods:title>
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<mods:namePart id="55FDA892BD6E19BD96A2153C520A9D08">Sikorski, Andrey V.</mods:namePart>
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<mods:namePart id="7FD5FAEFFAB152539A8272C76FAB876F">Pavlova, Lyudmila V.</mods:namePart>
<mods:affiliation id="FE5C8D59E2076ABE3DBE8F479E1E3496">Russian Academy of Sciences, Kola Science Centre, Murmansk Marine Biological Institute, Vladimirskaya str. 17, Murmansk 183010, Russia. sea 1234 @ mail. ru; http: // orcid. org / 0000 - 0003 - 4422 - 0366</mods:affiliation>
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<mods:namePart id="90454DC7440704A5C87B0A0CA3C856F2">Sardá, Rafael</mods:namePart>
<mods:affiliation id="D725D9FF32E747C1B4A5477965EDBEEB">Centre dEstudis Avançats de Blanes (CEAB-CSIC), Carrer daccès a la Cala St. Francesc, 14, Blanes-Girona- 17300, Spain. sarda @ ceab. csic. es; http: // orcid. org / 0000 - 0003 - 2059 - 9017</mods:affiliation>
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<mods:namePart id="14F6487AA9D4ACACED7B7CC918EA5FA3">Langeneck, Joachim</mods:namePart>
<mods:affiliation id="BE61D678B48EB8034E71E0C61F9A8DE3">Dipartimento di Biologia, Universit di Pisa, via Derna 1, 56126 Pisa, Italy. jlangeneck @ biologia. unipi. it; http: // orcid. org / 0000 - 0003 - 3665 - 8683</mods:affiliation>
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<mods:namePart id="3F9FAF965BFC3913DD7A80B4C7DA400E">Gil, João</mods:namePart>
<mods:affiliation id="93C560EBE6C2DEDD843F2CA0229917C5">Centre of Marine Sciences (CCMAR), Universidade do Algarve, Campus de Gambelas, 8005 - 139 Faro, Portugal. joaocfgil @ gmail. com; http: // orcid. org / 0000 - 0002 - 1856 - 3333</mods:affiliation>
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<mods:namePart id="7BFAC621BF96D299490F205D22DC558D">Ravara, Ascensão</mods:namePart>
<mods:affiliation id="8006D66268A4C7445F4068EC694453C3">CESAM - Centro de Estudos do Ambiente e do Mar, Dep. Biologia, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - 193 Aveiro, Portugal. aravara @ ua. pt; http: // orcid. org / 0000 - 0002 - 1689 - 2985</mods:affiliation>
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<treatment id="03C687E2FFF6EE00F0CB5144DBF81C88" ID-DOI="http://doi.org/10.5281/zenodo.4455622" ID-GBIF-Taxon="177014014" ID-Zenodo-Dep="4455622" LSID="urn:lsid:plazi:treatment:03C687E2FFF6EE00F0CB5144DBF81C88" httpUri="http://treatment.plazi.org/id/03C687E2FFF6EE00F0CB5144DBF81C88" lastPageId="7" lastPageNumber="521" pageId="4" pageNumber="519">
<subSubSection id="C375657FFFF6EE03F0CB5144D9231A5F" box="[151,773,1860,1887]" pageId="4" pageNumber="519" type="nomenclature">
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<heading id="D0988198FFF6EE03F0CB5144D9231A5F" bold="true" box="[151,773,1860,1887]" fontSize="11" level="1" pageId="4" pageNumber="519" reason="1">
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<taxonomicName id="4C6F4D77FFF6EE03F0CB5144D8861A5F" ID-CoL="85RGR" authority="Sikorski" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[151,672,1860,1887]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="519" phylum="Annelida" rank="species" species="barcinensis" status="sp. nov." subGenus="Appelloefia">
<emphasis id="B91BEAE6FFF6EE03F0CB5144DADF1A5F" bold="true" box="[151,249,1860,1886]" italics="true" pageId="4" pageNumber="519">Laonice</emphasis>
(
<emphasis id="B91BEAE6FFF6EE03F1565144DBB21A5E" bold="true" box="[266,404,1860,1887]" italics="true" pageId="4" pageNumber="519">Appelloefia</emphasis>
)
<emphasis id="B91BEAE6FFF6EE03F1FA5144D8171A5E" bold="true" box="[422,561,1860,1887]" italics="true" pageId="4" pageNumber="519">barcinensis</emphasis>
Sikorski
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<taxonomicNameLabel id="A228579DFFF6EE03F2FB5144D9231A5F" box="[679,773,1860,1886]" pageId="4" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BD036F4FFF6EE03F0CB5168DB431A83" blockId="4.[151,1053,1860,1959]" box="[151,357,1896,1922]" pageId="4" pageNumber="519">
(
<figureCitation id="13542A71FFF6EE03F0C35168DB2E1A83" box="[159,264,1896,1922]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="4" pageNumber="519">Figures 3</figureCitation>
AK, 4)
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LSID:
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urn:lsid:zoobank.org:act:
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<subSubSection id="C375657FFFF6EE02F0CB51D7DB131CAF" lastPageId="5" lastPageNumber="520" pageId="4" pageNumber="519" type="materials_examined">
<paragraph id="8BD036F4FFF6EE02F0CB51D7D9481DF8" blockId="4.[151,674,2006,2030]" lastBlockId="5.[151,1437,150,2013]" lastPageId="5" lastPageNumber="520" pageId="4" pageNumber="519">
<taxonomicName id="4C6F4D77FFF6EE03F0CB51D7DB431AEF" authorityName="Soderstrom" authorityYear="1920" box="[151,357,2006,2030]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="519" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B91BEAE6FFF6EE03F0CB51D7DB431AEF" box="[151,357,2006,2030]" italics="true" pageId="4" pageNumber="519">Laonice bahusiensis</emphasis>
</taxonomicName>
<bibRefCitation id="EFFE4B05FFF6EE03F12351D6D86B1AEF" author="Sarda, R. &amp; Gil, J. &amp; Taboada, S. &amp; Gili, J. M." box="[383,589,2006,2030]" pageId="4" pageNumber="519" pagination="1 - 21" refId="ref8503" refString="Sarda, R., Gil, J., Taboada, S. &amp; Gili, J. M. (2009) Polychaete species captured in sediment traps moored in northwestern Mediterranean submarine canyons. Zoological Journal of the Linnean Society, 155 (1), 1 - 21. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00442. x" type="journal article" year="2009">
Sardá
<emphasis id="B91BEAE6FFF6EE03F1E351D7DBD41AEF" box="[447,498,2006,2030]" italics="true" pageId="4" pageNumber="519">et al.</emphasis>
, 2009: 4
</bibRefCitation>
, table 2.
<emphasis id="B91BEAE6FFF7EE02F0CB5696DBA71DB1" bold="true" box="[151,385,150,176]" pageId="5" pageNumber="520">Material examined.</emphasis>
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<typeStatus id="54D48856FFF7EE02F1DB5697DBC91DB0" box="[391,495,151,177]" pageId="5" pageNumber="520" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBC99E8FFFF7EE02F1A25697D8BD1DB0" box="[510,667,150,177]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="520" type="Museum">MNCN 16.01</specimenCode>
/18548):
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,
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,
<location id="8EB0602FFFF7EE02F4985697DAE41DD4" LSID="urn:lsid:plazi:treatment:03C687E2FFF6EE00F0CB5144DBF81C88:8EB0602FFFF7EE02F4985697DAE41DD4" latitude="4.1993" longLatPrecision="7" longitude="-4.1994" name="NW Mediterranean Sea" pageId="5" pageNumber="520" stateProvince="SW Barcelona">NW Mediterranean Sea</location>
,
<collectorName id="269A5322FFF7EE02F08D56BBD94E1DD4" box="[209,872,187,213]" pageId="5" pageNumber="520">Projects EC-EUROMARGE-NB / CICYT-CONCENTRA</collectorName>
, St. 05C21,
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<geoCoordinate id="EE5B5033FFF7EE02F42056BBDED21DD4" box="[1148,1268,187,213]" degrees="01.9333" direction="east" orientation="longitude" pageId="5" pageNumber="520" precision="5" value="1.9333">01.9333ºE</geoCoordinate>
,
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moored
<quantity id="4C979B11FFF7EE02F0A856DFDB0B1DF8" box="[244,301,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="520" unit="m" value="30.0">
<elevation id="0042D1C7FFF7EE02F0A856DFDB0B1DF8" box="[244,301,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="520" unit="m" value="30.0">30 m</elevation>
</quantity>
above a sea ground
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deep, 04.199304.1994.
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</paragraph>
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<materialsCitation id="3B073CA9FFF7EE02F09B5703D94F1C67" ID-GBIF-Occurrence="3023040303" collectionCode="EC-EURO-, MARGE-NB" collectorName="Projects EC-EURO-MARGE-NB / CICYT-CONCENTRA" elevation="30" latitude="4.1993" longLatPrecision="7" longitude="-4.1994" pageId="5" pageNumber="520" specimenCode="MNCN 16.01" specimenCount="1" typeStatus="paratype">
Three
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from
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submarine canyon,
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Mediterranean Sea,
<collectorName id="269A5322FFF7EE02F4EA5703DF361C1C" box="[1206,1296,259,285]" pageId="5" pageNumber="520">Projects</collectorName>
<collectionCode id="ED7EAE31FFF7EE02F5465703DB0A1C40" pageId="5" pageNumber="520">EC-EURO-MARGE-NB</collectionCode>
/ CICYT-CONCENTRA: St. 01C22,
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,
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moored
<quantity id="4C979B11FFF7EE02F4A65727DF111C40" box="[1274,1335,295,321]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="520" unit="m" value="30.0">
<elevation id="0042D1C7FFF7EE02F4A65727DF111C40" box="[1274,1335,295,321]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="520" unit="m" value="30.0">30 m</elevation>
</quantity>
above a sea bed
<quantity id="4C979B11FFF7EE02F0A9574ADB6D1C64" box="[245,331,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.1949999999999998" pageId="5" pageNumber="520" unit="m" value="1195.0">
<elevation id="0042D1C7FFF7EE02F0A9574ADB6D1C64" box="[245,331,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.1949999999999998" pageId="5" pageNumber="520" unit="m" value="1195.0">1195 m</elevation>
</quantity>
deep, 04.199304.1994 (
<specimenCode id="DBC99E8FFFF7EE02F230574BD92C1C67" box="[620,778,331,358]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="520" type="Museum">MNCN 16.01</specimenCode>
/18676);
</materialsCitation>
<materialsCitation id="3B073CA9FFF7EE02F32E574BDB171CAF" ID-GBIF-Occurrence="3023040304" collectionCode="EC-EURO-, MARGE-NB" collectorName="Projects EC-EURO-MARGE-NB / CICYT-CONCENTRA" elevation="30" latitude="4.1993" longLatPrecision="7" longitude="-4.1994" pageId="5" pageNumber="520" specimenCode="MNCN 16.01" specimenCount="1" typeStatus="paratype">
St. 01C24 and St. 02C24,
<geoCoordinate id="EE5B5033FFF7EE02F4FC574BDF3E1C64" box="[1184,1304,331,357]" degrees="41.0333" direction="north" orientation="latitude" pageId="5" pageNumber="520" precision="5" value="41.0333">41.0333ºN</geoCoordinate>
<geoCoordinate id="EE5B5033FFF7EE02F57C574BDFBE1C64" box="[1312,1432,331,357]" degrees="01.9333" direction="east" orientation="longitude" pageId="5" pageNumber="520" precision="5" value="1.9333">01.9333ºE</geoCoordinate>
,
<collectingMethod id="522E4EE3FFF7EE02F0CB576FDB141C88" box="[151,306,367,393]" pageId="5" pageNumber="520">sediment trap</collectingMethod>
moored
<quantity id="4C979B11FFF7EE02F1C7576EDBC31C88" box="[411,485,366,393]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="520" unit="m" value="500.0">
<elevation id="0042D1C7FFF7EE02F1C7576EDBC31C88" box="[411,485,366,393]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="520" unit="m" value="500.0">500 m</elevation>
</quantity>
above a sea bed
<quantity id="4C979B11FFF7EE02F2ED576FD8DD1C88" box="[689,763,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="520" unit="m" value="990.0">
<elevation id="0042D1C7FFF7EE02F2ED576FD8DD1C88" box="[689,763,367,393]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="520" unit="m" value="990.0">990 m</elevation>
</quantity>
deep, 04.199304.1994 (
<specimenCode id="DBC99E8FFFF7EE02F443576FDE9A1C88" box="[1055,1212,367,393]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="520" type="Museum">MNCN 16.01</specimenCode>
/18677 and
<specimenCode id="DBC99E8FFFF7EE02F514576FDAF01CAF" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="520" type="Museum">MNCN 16.01</specimenCode>
/18617)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF7EE01F09B57B6DBEA1B0C" lastPageId="6" lastPageNumber="521" pageId="5" pageNumber="520" type="description">
<paragraph id="8BD036F4FFF7EE02F09B57B6DE891F18" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
<emphasis id="B91BEAE6FFF7EE02F09B57B6DB7F1CD1" bold="true" box="[199,345,438,464]" pageId="5" pageNumber="520">Description.</emphasis>
<typeStatus id="54D48856FFF7EE02F13E57B7DBEC1CD0" box="[354,458,439,465]" pageId="5" pageNumber="520" type="holotype">Holotype</typeStatus>
complete, 126 chaetigers,
<quantity id="4C979B11FFF7EE02F2A357B7D9761CD0" box="[767,848,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="520" unit="mm" value="32.0">32 mm</quantity>
long, and
<quantity id="4C979B11FFF7EE02F39057B7DE031CD0" box="[972,1061,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="520" unit="mm" value="1.2">1.2 mm</quantity>
wide at about chaetiger 10. One
<typeStatus id="54D48856FFF7EE02F0CB57DBDAD11CF4" box="[151,247,475,501]" pageId="5" pageNumber="520" type="paratype">paratype</typeStatus>
(MNCN 16.01/18676) complete—99 chaetigers, two other
<typeStatus id="54D48856FFF7EE02F3C157DBDE2E1CF4" box="[925,1032,475,501]" pageId="5" pageNumber="520" type="paratype">paratypes</typeStatus>
fragmented, however with pygidial fragments. Worms up to
<quantity id="4C979B11FFF7EE02F1F057FFDBDD1F18" box="[428,507,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="520" unit="mm" value="61.0">61 mm</quantity>
long for 142 chaetigers, and
<quantity id="4C979B11FFF7EE02F31E57FFD9BE1F18" box="[834,920,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="520" unit="mm" value="1.6">1.6 mm</quantity>
wide at chaetigers 712.
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B5423D8501E80" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Prostomium longer than wide, from almost triangular, slightly T-shaped (
<figureCitation id="13542A71FFF7EE02F3AA5423DE681F3C" box="[1014,1102,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3A</figureCitation>
) to almost quadrangular, with anterior margin truncated or slightly rounded with slight concavity, and completely separated from the peristomium at the anterior edges. Eyespots nearly invisible, smaller individuals may show one pair of faint (probably deeply embedded) eye-strips close to the base of the occipital antenna. Occipital antenna long and very thin (as long as or longer than notopodial postchaetal lamellae of chaetiger 1), ending in a thread-like tip, usually vertically erect, sometimes with middle bulbous enlargement, and inserted at the end of the prostomium. Intact palps lost from all specimens (one very short, torn off palp present in the sample MNCN 16.01/18617). Nuchal organs as long double U-shaped ciliary bands extending posteriorly to chaetigers 911 (
<figureCitation id="13542A71FFF7EE02F3D8551FD9C61E38" box="[900,992,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3A</figureCitation>
). Anterior region of body (i.e. anterior segments bearing the nuchal organs plus the following one to approximately 10 anterior chaetigers) approximately 1.5 times wider than subsequent segments.
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B558BDEDB1958" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Branchiae from chaetiger 2, present nearly along the entire body, although absent in the 1819 posterior-most chaetigers. Branchiae free, one third shorter than notopodial postchaetal lamellae until chaetiger 4, then gradually increasing in length and reaching about the same length as the notopodial postchaetal lamellae at chaetiger 79 (
<figureCitation id="13542A71FFF7EE02F53355D3DAF01910" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3A,C</figureCitation>
), and becoming 1.21.3 times longer than notopodial postchaetal lamellae on subsequent segments (
<figureCitation id="13542A71FFF7EE02F51C55F7DFBE1910" box="[1344,1432,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3B</figureCitation>
, DF), except in the very posterior ones, where they become about one third shorter (
<figureCitation id="13542A71FFF7EE02F40F521BDE881934" box="[1107,1198,1051,1077]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3G</figureCitation>
). Branchiae from the mid-body ending in a threadlike process (
<figureCitation id="13542A71FFF7EE02F235523FD8981958" box="[617,702,1087,1113]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3F</figureCitation>
), posterior-most branchiae thin, subulate (
<figureCitation id="13542A71FFF7EE02F4CA523FDED61958" box="[1174,1264,1087,1113]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B5263D8551830" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Notopodial prechaetal lamellae visible from chaetiger 3 to 1222, attaining maximum size on chaetigers 910 (approx. 0.4 the length of notopodial postchaetal lamellae). Longest notopodial postchaetal lamellae at chaetigers 725. Notopodial postchaetal lamellae of the first 47 chaetigers leaf-like with elongated pointed tips (
<figureCitation id="13542A71FFF7EE02F56952ABDFA819C4" box="[1333,1422,1195,1221]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3C</figureCitation>
), more obtuse afterwards, with broadly rounded upper margins on chaetigers posterior to nuchal organs (
<figureCitation id="13542A71FFF7EE02F54A52CFDF7019E8" box="[1302,1366,1231,1257]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3</figureCitation>
DE), lacking an upwardly extension after chaetiger 45 (
<figureCitation id="13542A71FFF7EE02F2E352F3D935180C" box="[703,787,1267,1293]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3F</figureCitation>
), becoming lanceolate at chaetiger 85 (
<figureCitation id="13542A71FFF7EE02F49D52F3DF31180C" box="[1217,1303,1267,1293]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3G</figureCitation>
) and finally disappearing 4 segments before pygidium.
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B533BD93C1B08" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Neuropodial prechaetal lamellae visible from chaetiger 2 to 1213, reaching maximum size in chaetigers 79. Neuropodial postchaetal lamellae reaching their maximum size on chaetigers 817 (
<figureCitation id="13542A71FFF7EE02F402535FDE9E1878" box="[1118,1208,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3C</figureCitation>
), then gradually diminishing further back, becoming very short on the posterior 4045 chaetigers (
<figureCitation id="13542A71FFF7EE02F4405383DE50189C" box="[1052,1142,1411,1437]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3G</figureCitation>
). Neuropodial postchaetal lamellae bearing acute upwardly turned tips on the first 7 chaetigers (
<figureCitation id="13542A71FFF7EE02F3C153A7D9D218C0" box="[925,1012,1447,1473]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3C</figureCitation>
), with a peak gradually shifting along the margin of the lamella afterwards and becoming less evident (
<figureCitation id="13542A71FFF7EE02F33C53CBD98C18E4" box="[864,938,1483,1509]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Figs. 3</figureCitation>
DE); neuropodial postchaetal lamellae with acute tips in the posterior half of body again (
<figureCitation id="13542A71FFF7EE02F2C953EFD8F31B08" box="[661,725,1519,1545]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3</figureCitation>
FG).
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B5013DF4F1B50" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Transverse dorsal crests connecting bases of notopodial postchaetal lamellae (
<figureCitation id="13542A71FFF7EE02F4775013DEA71B2C" box="[1067,1153,1555,1581]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3B</figureCitation>
) from chaetiger 1215 to chaetiger 5074. Transverse dorsal crests generally low, attaining maximum height between chaetigers 1547.
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B505BD8231BBC" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">Inter-parapodial pouches from chaetigers 58 to nearly the end of the body, ending 57 chaetigers before pygidium. The inter-parapodial pouches may start from unpaired segments on opposite sides of the same specimen (difference of up to 2 segments).</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B50C7D8711A94" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Anterior parapodia with notopodial capillary chaetae arranged in more than 2 vertical rows in chaetigers 115 (with up to 6 rows in chaetigers 810), in 2 vertical rows from chaetiger 16; and neuropodial capillaries arranged in more than 2 vertical rows in the 12 anterior most chaetigers (
<figureCitation id="13542A71FFF7EE02F31D510FD9BE1A28" box="[833,920,1807,1833]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3C</figureCitation>
), in 2 rows only on more posterior chaetigers. Neuropodial hooded hooks appearing from chaetiger 1922, numbering 911 per bundle, and bearing 3 apical teeth above the main fang (
<figureCitation id="13542A71FFF7EE02F1CA5157DBFF1A70" box="[406,473,1879,1905]" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3</figureCitation>
HI, K). Sabre chaetae appearing from chaetiger 1112, numbering at first 34 per bundle, and then 12 from chaetiger 16.
</paragraph>
<paragraph id="8BD036F4FFF7EE02F09B519FDAE61ADC" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="520">
Pygidium with a pair of short ventral lobes and two pairs of thin and long, threadlike lateral anal cirri (
<figureCitation id="13542A71FFF7EE02F533519FDA951ADC" captionStart="FIGURE 3" captionStartId="6.[151,250,1165,1189]" captionTargetBox="[156,1436,188,1133]" captionTargetId="figure-28@6.[151,1436,181,1141]" captionTargetPageId="6" captionText="FIGURE 3. Laonice (Appelloefia) barcinensis sp. nov. A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4." figureDoi="http://doi.org/10.5281/zenodo.4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="5" pageNumber="520">Fig. 3J</figureCitation>
).
</paragraph>
<caption id="DF10667CFFF4EE01F0CB528DD8031858" ID-DOI="http://doi.org/10.5281/zenodo.4447224" ID-Zenodo-Dep="4447224" httpUri="https://zenodo.org/record/4447224/files/figure.png" pageId="6" pageNumber="521" startId="6.[151,250,1165,1189]" targetBox="[156,1436,188,1133]" targetPageId="6">
<paragraph id="8BD036F4FFF4EE01F0CB528DD8031858" blockId="6.[151,1437,1165,1369]" pageId="6" pageNumber="521">
<emphasis id="B91BEAE6FFF4EE01F0CB528DDB3519A4" bold="true" box="[151,275,1165,1189]" pageId="6" pageNumber="521">FIGURE 3.</emphasis>
<taxonomicName id="4C6F4D77FFF4EE01F146528ED85519A4" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[282,627,1165,1189]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="barcinensis" status="sp. nov." subGenus="Appelloefia">
<emphasis id="B91BEAE6FFF4EE01F146528EDB4A19A4" box="[282,364,1166,1189]" italics="true" pageId="6" pageNumber="521">Laonice</emphasis>
(
<emphasis id="B91BEAE6FFF4EE01F127528DDBD619A4" box="[379,496,1165,1189]" italics="true" pageId="6" pageNumber="521">Appelloefia</emphasis>
)
<emphasis id="B91BEAE6FFF4EE01F1A3528DD85519A4" box="[511,627,1165,1189]" italics="true" pageId="6" pageNumber="521">barcinensis</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF4EE01F226528DD8ED19A4" bold="true" box="[634,715,1165,1189]" pageId="6" pageNumber="521">
<taxonomicNameLabel id="A228579DFFF4EE01F226528DD8ED19A4" box="[634,715,1165,1189]" pageId="6" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Anterior end, dorsal view. B. Midbody chaetigers 3740, dorso-lateral view, showing transverse dorsal crests. CG. Right parapodium, anterior view: C, chaetiger 7; D, chaetiger 18; E, chaetiger 30; F, chaetiger 57; G, chaetiger 111. H. Neuropodial hooded hook of chaetiger 52, side view. I. Neuropodial hooded hook of chaetiger 52, frontal view. J. Pygidium, dorsal view. K. Neuropodial hooded hook from posterior segment, SEM photo. Material: AI—paratype MNCN 16.01/18617; J—paratype MNCN 16.01/18677; K—paratype MNCN 16.01/18676. Scales (mm): AB = 1.0; CG = 0.5; HI = 0.005; J = 0.4.
</paragraph>
</caption>
<paragraph id="8BD036F4FFF4EE01F09B5388D87018A0" blockId="6.[151,1437,1415,2018]" box="[199,598,1415,1442]" pageId="6" pageNumber="521">No specific pigmentation observed.</paragraph>
<paragraph id="8BD036F4FFF4EE01F09B53ABDBEA1B0C" blockId="6.[151,1437,1415,2018]" pageId="6" pageNumber="521">
<emphasis id="B91BEAE6FFF4EE01F09B53ABD82318C4" bold="true" box="[199,517,1451,1477]" pageId="6" pageNumber="521">Staining in Methyl Green.</emphasis>
Anterior margin of prostomium and peristomium, occipital antenna, margins of postchaetal lamellae and small oval lateral spots between notopodia and neuropodia visibly staining more intensely than adjacent parts of body.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF4EE01F09B5017DEC61B54" pageId="6" pageNumber="521" type="etymology">
<paragraph id="8BD036F4FFF4EE01F09B5017DEC61B54" blockId="6.[151,1437,1415,2018]" pageId="6" pageNumber="521">
<emphasis id="B91BEAE6FFF4EE01F09B5017DB681B30" bold="true" box="[199,334,1559,1585]" pageId="6" pageNumber="521">Etymology.</emphasis>
The name
<taxonomicName id="4C6F4D77FFF4EE01F1925017D8681B30" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[462,590,1559,1585]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="barcinensis">
<emphasis id="B91BEAE6FFF4EE01F1925017D8681B30" box="[462,590,1559,1585]" italics="true" pageId="6" pageNumber="521">barcinensis</emphasis>
</taxonomicName>
is derived from Barcino, ancient Latin name of Barcelona, and is given to this species in honor of the city of Barcelona, located close to the
<typeStatus id="54D48856FFF4EE01F324503CD98E1B57" box="[888,936,1596,1622]" pageId="6" pageNumber="521">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF4EE01F09B505FD9D41B9C" pageId="6" pageNumber="521" type="distribution">
<paragraph id="8BD036F4FFF4EE01F09B505FD9D41B9C" blockId="6.[151,1437,1415,2018]" pageId="6" pageNumber="521">
<emphasis id="B91BEAE6FFF4EE01F09B505FDB471B78" bold="true" box="[199,353,1631,1657]" pageId="6" pageNumber="521">Distribution.</emphasis>
Mediterranean Sea (
<figureCitation id="13542A71FFF4EE01F214505FD8AB1B78" box="[584,653,1631,1657]" captionStart="FIGURE 4" captionStartId="7.[151,250,1296,1320]" captionTargetBox="[155,1430,422,1270]" captionTargetId="figure-128@7.[151,1436,417,1273]" captionTargetPageId="7" captionText="FIGURE 4. Distributional map of the Mediterranean deep-sea Laonice species: circles, Laonice (Sarsiana) alberti sp. nov.; triangles, Laonice (Appelloefia) barcinensis sp. nov.; star, Laonice (Laonice) sp.; T, type localities." figureDoi="http://doi.org/10.5281/zenodo.4447226" httpUri="https://zenodo.org/record/4447226/files/figure.png" pageId="6" pageNumber="521">Fig. 4</figureCitation>
): Foix submarine canyon (off NE Spain), in sediment traps moored
<quantity id="4C979B11FFF4EE01F5DC505FDA8B1B9F" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="521" unit="m" value="30.0">30 m</quantity>
above a seabed
<quantity id="4C979B11FFF4EE01F1385083DBD61B9F" box="[356,496,1667,1694]" metricMagnitude="3" metricUnit="m" metricValue="1.0925" metricValueMax="1.1949999999999998" metricValueMin="0.99" pageId="6" pageNumber="521" unit="m" value="1092.5" valueMax="1195.0" valueMin="990.0">9901195 m</quantity>
deep, and
<quantity id="4C979B11FFF4EE01F2355083D8961B9F" box="[617,688,1667,1694]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="521" unit="m" value="500.0">500 m</quantity>
above a seabed
<quantity id="4C979B11FFF4EE01F3345083D9891B9F" box="[872,943,1667,1694]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="6" pageNumber="521" unit="m" value="990.0">990 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF4EE00F09B50A7DBF81C88" lastPageId="7" lastPageNumber="522" pageId="6" pageNumber="521" type="discussion">
<paragraph id="8BD036F4FFF4EE01F09B50A7DBAE1ABF" blockId="6.[151,1437,1415,2018]" pageId="6" pageNumber="521">
<emphasis id="B91BEAE6FFF4EE01F09B50A7DB1D1BC0" bold="true" box="[199,315,1703,1729]" pageId="6" pageNumber="521">Remarks.</emphasis>
<taxonomicName id="4C6F4D77FFF4EE01F11A50A8D8EC1BC0" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[326,714,1703,1729]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="barcinensis" status="sp. nov." subGenus="Appelloefia">
<emphasis id="B91BEAE6FFF4EE01F11A50A8D8EC1BC0" box="[326,714,1703,1729]" italics="true" pageId="6" pageNumber="521">Laonice (Appelloefia) barcinensis</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF4EE01F28950A7D9141BC0" bold="true" box="[725,818,1703,1729]" pageId="6" pageNumber="521">
<taxonomicNameLabel id="A228579DFFF4EE01F28950A7D9141BC0" box="[725,818,1703,1729]" pageId="6" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically very similar to
<taxonomicName id="4C6F4D77FFF4EE01F48950A8DFBA1BC0" box="[1237,1436,1703,1730]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="appelloefi" subGenus="Appelloefia">
<emphasis id="B91BEAE6FFF4EE01F48950A8DFBA1BC0" box="[1237,1436,1703,1730]" italics="true" pageId="6" pageNumber="521">L. (A.) appelloefi</emphasis>
</taxonomicName>
Ŝderstr̂m, 1920 and
<taxonomicName id="4C6F4D77FFF4EE01F1C150CCD9D91BE7" authority="Aguirrezabalaga &amp; Ceberio, 2005" authorityName="Aguirrezabalaga &amp; Ceberio" authorityYear="2005" box="[413,1023,1739,1766]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="maciolekae" subGenus="Appelloefia">
<emphasis id="B91BEAE6FFF4EE01F1C150CCD8541BE4" box="[413,626,1739,1766]" italics="true" pageId="6" pageNumber="521">L. (A.) maciolekae</emphasis>
<bibRefCitation id="EFFE4B05FFF4EE01F22750CBD9D91BE7" author="Aguirrezabalaga, F. &amp; Ceberio, A." box="[635,1023,1739,1766]" pageId="6" pageNumber="521" pagination="267 - 280" refId="ref6577" refString="Aguirrezabalaga, F. &amp; Ceberio, A. (2005) Spionidae (Annelida: Polychaeta) from the Capbreton Canyon (Bay of Biscay, NE Atlantic) with descriptions of a new genus and three new species. Marine Biology Research, 1 (4), 267 - 280. https: // doi. org / 10.1080 / 17451000500262066" type="journal article" year="2005">Aguirrezabalaga &amp; Ceberio, 2005</bibRefCitation>
</taxonomicName>
(synonymized with
<taxonomicName id="4C6F4D77FFF4EE01F4B050CCDBA21A08" authority="in Meissner et al., 2014" authorityName="in Meissner" authorityYear="2014" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="appelloefi">
<emphasis id="B91BEAE6FFF4EE01F4B050CCDF5B1BE4" box="[1260,1405,1739,1765]" italics="true" pageId="6" pageNumber="521">L. appelloefi</emphasis>
in
<bibRefCitation id="EFFE4B05FFF4EE01F0CB50EFDBA21A08" author="Meissner, K. &amp; Bick, A. &amp; Guggolz, T. &amp; Gotting, M." box="[151,388,1775,1802]" pageId="6" pageNumber="521" pagination="201 - 245" refId="ref8341" refString="Meissner, K., Bick, A., Guggolz, T. &amp; Gotting, M. (2014) Spionidae (Polychaeta: Canalipalpata: Spionida) from seamounts in the NE Atlantic. Zootaxa, 3786 (3), 201 - 245. https: // doi. org / 10.11646 / zootaxa. 3786.3.1" type="journal article" year="2014">
Meissner
<emphasis id="B91BEAE6FFF4EE01F0A250F0DB1E1A08" box="[254,312,1775,1801]" italics="true" pageId="6" pageNumber="521">et al.</emphasis>
, 2014
</bibRefCitation>
</taxonomicName>
). However, branchiae and inter-parapodial pouches were only observed in
<taxonomicName id="4C6F4D77FFF4EE01F48850F0DF451A08" box="[1236,1379,1775,1801]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="appelloefi">
<emphasis id="B91BEAE6FFF4EE01F48850F0DF451A08" box="[1236,1379,1775,1801]" italics="true" pageId="6" pageNumber="521">L. appelloefi</emphasis>
</taxonomicName>
on a limited number of segments, whereas in the new species these structures are present through almost the entire length of the body, with the exception of the last posterior-most segments. These two characters suggest that the new species is similar to
<taxonomicName id="4C6F4D77FFF4EE01F101515CD94A1A74" authority="Sikorski, 2003" authorityName="Sikorski" authorityYear="2003" box="[349,876,1883,1910]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="norgensis" subGenus="Norgensia">
<emphasis id="B91BEAE6FFF4EE01F101515CD89D1A74" box="[349,699,1884,1910]" italics="true" pageId="6" pageNumber="521">Laonice (Norgensia) norgensis</emphasis>
<bibRefCitation id="EFFE4B05FFF4EE01F298515BD94A1A74" author="Sikorski, A. V." box="[708,876,1883,1909]" pageId="6" pageNumber="521" pagination="316 - 345" refId="ref8665" refString="Sikorski, A. V. (2003) Laonice (Polychaeta, Spionidae) in the Arctic and the North Atlantic. Sarsia, 88 (5), 316 - 345. https: // doi. org / 10.1080 / 00364820310002551" type="journal article" year="2003">Sikorski, 2003</bibRefCitation>
</taxonomicName>
, but this species has much longer nuchal organs, reaching chaetiger 1529, and the inter-parapodial pouches appear from chaetiger 817 (against chaetiger
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<taxonomicName id="4C6F4D77FFF4EE01F5DA5180DB311ABC" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="521" phylum="Annelida" rank="species" species="barcinensis" status="sp. nov.">
<emphasis id="B91BEAE6FFF4EE01F5DA5180DB311ABC" italics="true" pageId="6" pageNumber="521">L. barcinensis</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF4EE01F14251A3DB5E1ABC" bold="true" box="[286,376,1955,1981]" pageId="6" pageNumber="521">
<taxonomicNameLabel id="A228579DFFF4EE01F14251A3DB5E1ABC" box="[286,376,1955,1981]" pageId="6" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BD036F4FFF4EE00F09B51C8DBF81C88" blockId="6.[151,1437,1415,2018]" lastBlockId="7.[151,1437,150,393]" lastPageId="7" lastPageNumber="522" pageId="6" pageNumber="521">
It is remarkable to notice how large adult worms of several dozens of milligrams of wet weight (
<bibRefCitation id="EFFE4B05FFF4EE00F54551C7DAF51DB0" author="Sarda, R. &amp; Gil, J. &amp; Taboada, S. &amp; Gili, J. M." lastPageId="7" lastPageNumber="522" pageId="6" pageNumber="521" pagination="1 - 21" refId="ref8503" refString="Sarda, R., Gil, J., Taboada, S. &amp; Gili, J. M. (2009) Polychaete species captured in sediment traps moored in northwestern Mediterranean submarine canyons. Zoological Journal of the Linnean Society, 155 (1), 1 - 21. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00442. x" type="journal article" year="2009">
Sardá
<emphasis id="B91BEAE6FFF4EE01F53F51C8DFBB1AE0" box="[1379,1437,1991,2017]" italics="true" pageId="6" pageNumber="521">et al.</emphasis>
2009
</bibRefCitation>
), belonging to a typically benthic genus, seem able to reach sediment traps moored as high as
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above the seabed.
<bibRefCitation id="EFFE4B05FFF5EE00F14256BBDBD51DD4" author="Sarda, R. &amp; Gil, J. &amp; Taboada, S. &amp; Gili, J. M." box="[286,499,187,213]" pageId="7" pageNumber="522" pagination="1 - 21" refId="ref8503" refString="Sarda, R., Gil, J., Taboada, S. &amp; Gili, J. M. (2009) Polychaete species captured in sediment traps moored in northwestern Mediterranean submarine canyons. Zoological Journal of the Linnean Society, 155 (1), 1 - 21. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00442. x" type="journal article" year="2009">
Sardá
<emphasis id="B91BEAE6FFF5EE00F13A56BBDB861DD4" box="[358,416,187,213]" italics="true" pageId="7" pageNumber="522">et al.</emphasis>
(2009)
</bibRefCitation>
hypothesized that the existence of a highly turbulent environment associated with the transport of large amounts of water and sediment inside the Mediterranean submarine canyons (
<bibRefCitation id="EFFE4B05FFF5EE00F55056DFDAF51C1F" author="Canals, M. &amp; Puig, P. &amp; Durrieu de Madron, X. &amp; Heussner, S. &amp; Palanques, A. &amp; Fabres, J." pageId="7" pageNumber="522" pagination="354 - 357" refId="ref6966" refString="Canals, M., Puig, P., Durrieu de Madron, X., Heussner, S., Palanques, A. &amp; Fabres, J. (2006) Flushing submarine canyons. Nature, 444, 354 - 357. https: // doi. org / 10.1038 / nature 05271" type="journal article" year="2006">
Canals
<emphasis id="B91BEAE6FFF5EE00F53E56DFDFBB1DF8" box="[1378,1437,223,249]" italics="true" pageId="7" pageNumber="522">et al.</emphasis>
2006
</bibRefCitation>
), along with their strong bottom currents and resuspension events, would be responsible for the mobilization of sediment and benthic and epibenthic species into the water column. Such resuspension could even transport the specimens, up to hundreds of meters above the seabed where they normally live, reaching this way the sediment traps where they were found.
</paragraph>
</subSubSection>
</treatment>
</document>