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<document id="856D92D5C3A76A4820D0F924FCB5044E" ID-DOI="10.5281/zenodo.183373" ID-GBIF-Dataset="0d7eca74-c851-4e3a-9503-9c3909a8ccc8" ID-ISSN="1175-5326" ID-Zenodo-Dep="183373" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459904382763" checkinUser="plazi" docAuthor="Artois, Tom J. &amp; Tessens, Bart S." docDate="2008" docId="03C687D8FFA5FF56FF69E70AFB010E2B" docLanguage="en" docName="zt01849p027.pdf" docOrigin="Zootaxa 1849" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Gyratrix hermaphroditus Ehrenberg 1831" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FFFFFFA0FFA3FF5EFFFEE43FFF870F79" masterDocTitle="Polycystididae (Rhabditophora: Rhabdocoela: Kalyptorhynchia) from the Indian Ocean, with the description of twelve new species" masterLastPageNumber="27" masterPageNumber="1" pageNumber="7" updateTime="1698228922093" updateUser="plazi">
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<mods:title id="2B912DCCFCCF5EC03AEF34E08BB9CEBA">Polycystididae (Rhabditophora: Rhabdocoela: Kalyptorhynchia) from the Indian Ocean, with the description of twelve new species</mods:title>
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<mods:namePart id="77ABF0974DE60812E294BE715003B57F">Artois, Tom J.</mods:namePart>
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<mods:namePart id="C58027E26841D1A28DF580DE9E75F5A1">Tessens, Bart S.</mods:namePart>
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<mods:date id="F8518F35C20F379B1440FE1DF880A31B">2008</mods:date>
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<mods:number id="7DCA531D01FA2C7946E16870C9A1D238">1849</mods:number>
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<treatment id="03C687D8FFA5FF56FF69E70AFB010E2B" ID-DOI="http://doi.org/10.5281/zenodo.6229703" ID-GBIF-Taxon="119373284" ID-Zenodo-Dep="6229703" LSID="urn:lsid:plazi:treatment:03C687D8FFA5FF56FF69E70AFB010E2B" httpUri="http://treatment.plazi.org/id/03C687D8FFA5FF56FF69E70AFB010E2B" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C3756545FFA5FF58FF69E70AFAB50CE3" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BD036CEFFA5FF58FF69E70AFCED0C36" blockId="6.[151,874,821,847]" box="[151,874,821,847]" pageId="6" pageNumber="7">
<heading id="D09881A2FFA5FF58FF69E70AFCED0C36" bold="true" box="[151,874,821,847]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B91BEADCFFA5FF58FF69E70AFCED0C36" bold="true" box="[151,874,821,847]" pageId="6" pageNumber="7">
<taxonomicName id="4C6F4D4DFFA5FF58FF69E70AFD190C36" ID-CoL="6KVFM" authority="Ehrenberg, 1831" authorityName="Ehrenberg" authorityYear="1831" box="[151,670,821,847]" class="Rhabditophora" family="Polycystididae" genus="Gyratrix" kingdom="Animalia" order="Rhabdocoela" pageId="6" pageNumber="7" phylum="Platyhelminthes" rank="species" species="hermaphroditus">
<emphasis id="B91BEADCFFA5FF58FF69E70AFE400C36" bold="true" box="[151,455,821,847]" italics="true" pageId="6" pageNumber="7">Gyratrix hermaphroditus</emphasis>
<bibRefCitation id="EFFE4B3FFFA5FF58FE30E70AFD190C36" author="Ehrenberg" box="[462,670,821,847]" pageId="6" pageNumber="25" refString="Ehrenberg C. G. (1831) Animalia evertebrata exclusis insectis recensuit C. G. Ehrenberg. Series prima cum Tabularum decade prima. In: Hemprich, F. G. &amp; Ehrenberg C. G. (Eds.), Symbolae Physicae 2. Phytozoa Turbellaria. Officina Academica, Berolina, pp. 1 - 15." type="journal article" year="1831">Ehrenberg, 1831</bibRefCitation>
</taxonomicName>
species complex
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036CEFFA5FF58FF69E740FAB50CE3" blockId="6.[151,1437,895,2002]" box="[151,1330,895,922]" pageId="6" pageNumber="7">
<emphasis id="B91BEADCFFA5FF58FF69E740FE0A0CE0" bold="true" box="[151,397,895,921]" pageId="6" pageNumber="7">Known distribution</emphasis>
. Cosmopolitan and euryhaline, found from pure marine to pure limnic habitats.
</paragraph>
</subSubSection>
<subSubSection id="C3756545FFA5FF58FF3BE798FDCC0A8B" pageId="6" pageNumber="7" type="description">
<paragraph id="8BD036CEFFA5FF58FF3BE798FDCC0A8B" blockId="6.[151,1437,895,2002]" pageId="6" pageNumber="7">
<emphasis id="B91BEADCFFA5FF58FF3BE798FDFE0CB8" bold="true" box="[197,633,935,961]" pageId="6" pageNumber="7">New localities in the Indian Ocean</emphasis>
.
<collectingCountry id="F378765EFFA5FF58FD77E797FD760CBB" box="[649,753,936,962]" name="Tanzania" pageId="6" pageNumber="7">Zanzibar</collectingCountry>
Island (
<collectingCountry id="F378765EFFA5FF58FCACE797FC440CBB" box="[850,963,936,962]" name="Tanzania" pageId="6" pageNumber="7">Tanzania</collectingCountry>
): beach between the Institute of Marine Sciences and the Floating Restaurant,
<taxonomicName id="4C6F4D4DFFA5FF58FDA2E7EFFD120C90" box="[604,661,976,1001]" genus="Ulva" kingdom="Fungi" pageId="6" pageNumber="7" rank="genus">
<emphasis id="B91BEADCFFA5FF58FDA2E7EFFD120C90" box="[604,661,976,1001]" italics="true" pageId="6" pageNumber="7">Ulva</emphasis>
</taxonomicName>
-like algae with sand, from rocks beside a stair from the beach to the dike, lower eulittoral at low tide (
<date id="FFD1100EFFA5FF58FDBBE7C7FD560B6B" box="[581,721,1016,1042]" pageId="6" pageNumber="7" value="1995-08-03">03/08/1995</date>
); off Maruhubi Palace ruins, sand flat, coarse sand with some detritus, relatively dry with many crab holes (
<date id="FFD1100EFFA5FF58FD52E01FFCBF0B43" box="[684,824,1056,1082]" pageId="6" pageNumber="7" value="1995-08-05">05/08/1995</date>
); same locality, open beach with relatively fine sand (
<date id="FFD1100EFFA5FF58FF61E077FEAC0B1B" box="[159,299,1096,1122]" pageId="6" pageNumber="7" value="1995-08-05">05/08/1995</date>
).
<collectingCountry id="F378765EFFA5FF58FEC1E077FE460B1B" box="[319,449,1096,1122]" name="Seychelles" pageId="6" pageNumber="7">Seychelles</collectingCountry>
:
<collectingCountry id="F378765EFFA5FF58FE32E077FD950B1B" box="[460,530,1096,1122]" name="India" pageId="6" pageNumber="7">Mahé</collectingCountry>
, Grande Anse (Beolière), mangrove area, fine sediment from the upper eulittoral (
<date id="FFD1100EFFA5FF58FF22E04FFEE00BF3" box="[220,359,1136,1162]" pageId="6" pageNumber="7" value="1992-12-19">19/12/1992</date>
); same locality, between two mangrove areas, relatively fine sand (
<date id="FFD1100EFFA5FF58FB8BE04FFA870BF3" box="[1141,1280,1136,1162]" pageId="6" pageNumber="7" value="1992-12-15">15/12/1992</date>
), coarse sand (Amphioxus sand) from
<quantity id="4C979B2BFFA5FF58FE47E0A7FE720BCB" box="[441,501,1176,1202]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="m" value="19.0">19 m</quantity>
deep from an unknown locality in the
<collectingCountry id="F378765EFFA5FF58FC40E0A7FBB80BCB" box="[958,1087,1176,1202]" name="Seychelles" pageId="6" pageNumber="7">Seychelles</collectingCountry>
;
<collectingCountry id="F378765EFFA5FF58FBB5E0A7FB160BCB" box="[1099,1169,1176,1202]" name="India" pageId="6" pageNumber="7">Mahé</collectingCountry>
, Baie Beau Vallon, little pool between granite rocks, fine sand (
<date id="FFD1100EFFA5FF58FD74E0FFFC920BA3" box="[650,789,1216,1242]" pageId="6" pageNumber="7" value="1992-12-27">27/12/1992</date>
). McKenzie Point,
<collectingCountry id="F378765EFFA5FF58FC05E0FFFBE80BA3" box="[1019,1135,1216,1242]" name="Kenya" pageId="6" pageNumber="7">Mombasa</collectingCountry>
(
<collectingCountry id="F378765EFFA5FF58FB81E0FFFB540BA3" box="[1151,1235,1216,1242]" name="Kenya" pageId="6" pageNumber="7">Kenya</collectingCountry>
): fine, clear sand near to a mangrove area; very fine sand from a rock pool (
<date id="FFD1100EFFA5FF58FCB9E0D7FC550A7B" box="[839,978,1256,1282]" pageId="6" pageNumber="7" value="1992-10-12">12/10/1992</date>
); between rocks near the Four Seasons restaurant, dry coarse sand from the upper eulittoral (
<date id="FFD1100EFFA5FF58FCEEE12FFC1D0A53" box="[784,922,1296,1322]" pageId="6" pageNumber="7" value="1992-10-19">19/10/1992</date>
), same locality, fine sand with shell gravel, upper eulittoral (
<date id="FFD1100EFFA5FF58FEA5E107FE610A2B" box="[347,486,1336,1362]" pageId="6" pageNumber="7" value="1992-10-19">19/10/1992</date>
). Kanamai (
<collectingCountry id="F378765EFFA5FF58FD8BE107FD4F0A2B" box="[629,712,1336,1362]" name="Kenya" pageId="6" pageNumber="7">Kenya</collectingCountry>
): tide pool, coarse shell gravel with very fine sand, mid-eulittoral (
<date id="FFD1100EFFA5FF58FF21E15FFEEC0A03" box="[223,363,1376,1402]" pageId="6" pageNumber="7" value="1992-10-20">20/10/1992</date>
).
<collectingCountry id="F378765EFFA5FF58FE7FE15FFD950A03" box="[385,530,1376,1402]" name="Réunion" pageId="6" pageNumber="7">La Réunion</collectingCountry>
: Cap la Houssay, surf zone, coarse sand with pieces of coral (
<date id="FFD1100EFFA5FF58FAFDE15FFA080A03" box="[1283,1423,1376,1402]" pageId="6" pageNumber="7" value="1992-10-30">30/10/1992</date>
); Sabine les Bains, very coarse sand from a tide pool (
<date id="FFD1100EFFA5FF58FCA6E1B7FC640ADB" box="[856,995,1416,1442]" pageId="6" pageNumber="7" value="1992-11-02">02/11/1992</date>
). South-west Sulawesi (
<collectingCountry id="F378765EFFA5FF58FAE9E1B7FA160ADB" box="[1303,1425,1416,1442]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
): Samolana, Kudingareng Keke, Galesong, Kajangan, all in coral sand from the eulittoral up to 20 deep, all between
<date id="FFD1100EFFA5FF58FEFEE1E7FE020A8B" box="[256,389,1496,1522]" pageId="6" pageNumber="7" value="1984-09-30">30/09/1984</date>
and
<date id="FFD1100EFFA5FF58FE40E1E7FDC00A8B" box="[446,583,1496,1522]" pageId="6" pageNumber="7" value="1984-10-22">22/10/1984</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756545FFA5FF58FF3BE1C0FB3F093B" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BD036CEFFA5FF58FF3BE1C0FB3F093B" blockId="6.[151,1437,895,2002]" pageId="6" pageNumber="7">
<emphasis id="B91BEADCFFA5FF58FF3BE1C0FEB50960" bold="true" box="[197,306,1535,1561]" pageId="6" pageNumber="7">Material</emphasis>
. Four whole mounts from
<collectingCountry id="F378765EFFA5FF58FD8CE23FFD5B0963" box="[626,732,1536,1562]" name="Tanzania" pageId="6" pageNumber="7">Zanzibar</collectingCountry>
, four whole mounts from the
<collectingCountry id="F378765EFFA5FF58FBC0E23FFB390963" box="[1086,1214,1536,1562]" name="Seychelles" pageId="6" pageNumber="7">Seychelles</collectingCountry>
, six whole mounts from
<collectingCountry id="F378765EFFA5FF58FF29E217FEA0093B" box="[215,295,1576,1602]" name="Kenya" pageId="6" pageNumber="7">Kenya</collectingCountry>
, four whole mounts from
<collectingCountry id="F378765EFFA5FF58FDA9E217FD67093B" box="[599,736,1576,1602]" name="Réunion" pageId="6" pageNumber="7">La Réunion</collectingCountry>
and four whole mounts from
<collectingCountry id="F378765EFFA5FF58FBC1E217FB32093B" box="[1087,1205,1576,1602]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756545FFA5FF56FF3BE270FB010E2B" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BD036CEFFA5FF58FF3BE270FF5E08AB" blockId="6.[151,1437,895,2002]" pageId="6" pageNumber="7">
<emphasis id="B91BEADCFFA5FF58FF3BE270FEB00910" bold="true" box="[197,311,1615,1641]" pageId="6" pageNumber="7">Remarks</emphasis>
. The pioneering researches of
<bibRefCitation id="EFFE4B3FFFA5FF58FD5BE26FFCCB0913" author="Reuter" box="[677,844,1616,1642]" pageId="6" pageNumber="26" refString="Reuter, M. (1961) Untersuchungen uber rassenbildung bei Gyratrix hermaphroditus (Turbellaria Neorhabdocoela). Acta Zoologica Fennica, 100, 101 - 122." type="journal article" year="1961">Reuter (1961)</bibRefCitation>
,
<bibRefCitation id="EFFE4B3FFFA5FF58FCA4E26FFBA10913" author="Heitkamp" box="[858,1062,1616,1642]" pageId="6" pageNumber="25" refString="Heitkamp, U. (1978) Speciationsprozesse bei Gyratrix hermaphroditus Ehrenberg, 1831 (Turbellaria, Kalyptorhynchia). Zoomorphology, 90, 227 - 251." type="journal article" year="1978">Heitkamp (1978)</bibRefCitation>
, LHardy (1986) and especially Curini-Galletti &amp; Puccinelli (1989,1990, 1994, 1998),
<bibRefCitation id="EFFE4B3FFFA5FF58FCD6E247FB5709EB" author="Puccinelli" box="[808,1232,1656,1682]" pageId="6" pageNumber="26" refString="Puccinelli, I &amp; Curini-Galletti, M. C. (1987) Chromosomal evolution and speciation in marine populations of Gyratrix hermaphroditus sensu lato (Platyhelminthes: Kalyptorhynchia) and in other species of the Gyratricinae. Transactions of the American Microscopical Society, 106, 311 - 320." type="journal article" year="1987">Puccinelli &amp; Curini-Galletti (1987)</bibRefCitation>
, Puccinelli
<emphasis id="B91BEADCFFA5FF58FAA0E247FA1B09E8" box="[1374,1436,1656,1681]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(1990) and, more recently, Timoshkin
<emphasis id="B91BEADCFFA5FF58FDA4E29FFD1209C0" box="[602,661,1696,1721]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2004) have shown that this “species” is actually a large complex of cryptic species. These species differ from each other in karyotype and detailed construction and dimensions of the hard parts in the male system. The organisation of the female system can also differ, some of the species are eyeless and their colour can vary (see
<bibRefCitation id="EFFE4B3FFFA5FF58FD72E327FC96084B" author="Graff" box="[652,785,1816,1842]" pageId="6" pageNumber="25" refString="Graff, L. von (1905) Marine Turbellarien Orotavas und der Kusten Europas. Ergebnisse einiger, mit Unterstutzung der kais. Akademie der Wissenschaften in Wien (aus dem Legate Wedl) in den Jahren 1902 und 1903 unternommen Studienreise. II. Rhabdocoela. Zeitschrift fur Wissenschaftlichen Zoologie, 83, 68 - 148." type="journal article" year="1905">Graff 1905</bibRefCitation>
,
<bibRefCitation id="EFFE4B3FFFA5FF58FCE1E327FCDB084B" author="Graff" box="[799,860,1816,1842]" pageId="6" pageNumber="25" refString="Graff, L. von (1911) Acoela, Rhabdocoela und Alloeocoela des Ostens der Vereinigten Staaten von Amerika. Mit Nachtragen zu den &quot; Marinen Turbellarien Orotavas und der Kusten Europas &quot;. Zeitschrift fur Wissenschaftlichen Zoologie, 99, 1 - 108." type="journal article" year="1911">1911</bibRefCitation>
;
<bibRefCitation id="EFFE4B3FFFA5FF58FC97E327FB22084B" author="Artois" box="[873,1189,1816,1842]" pageId="6" pageNumber="25" refString="Artois, T. J. &amp; Schockaert, E. R. (2001) Interstitial fauna of the Galapagos: Duplacrorhynchinae, Macrorhynchinae, Polycystidinae, Gyratricinae (Platyhelminthes Polycystididae). Tropical Zoology, 14, 63 - 85." type="journal article" year="2001">Artois &amp; Schockaert 2001</bibRefCitation>
). The morphological and karyological diversity is not yet fully understood on a worldwide base, and the complex has not formally been split up. It could well be that the species complex is not monophyletic, and that some of the species are more closely related to
<taxonomicName id="4C6F4D4DFFA5FF58FE42E3AFFCC008D3" authority="Meixner, 1929" authorityName="Meixner" authorityYear="1929" box="[444,839,1936,1962]" class="Rhabditophora" family="Polycystididae" genus="Gyratrix" kingdom="Animalia" order="Rhabdocoela" pageId="6" pageNumber="7" phylum="Platyhelminthes" rank="species" species="proavus">
<emphasis id="B91BEADCFFA5FF58FE42E3AFFD0D08D0" box="[444,650,1936,1961]" italics="true" pageId="6" pageNumber="7">Gyratrix proavus</emphasis>
<bibRefCitation id="EFFE4B3FFFA5FF58FD68E3AFFCC008D3" author="Meixner" box="[662,839,1936,1962]" pageId="6" pageNumber="26" refString="Meixner, J. (1929) Morphologisch-okologische Studien an neuen Turbellarien aus dem Meeressande der Kieler Bucht. Zeitschrift fur Morphologie und Okologie der Tiere, 14, 765 - 791." type="journal article" year="1929">Meixner, 1929</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="4C6F4D4DFFA5FF58FC89E3AFFF5208AB" authority="Karling &amp; Schockaert, 1977" authorityName="Karling &amp; Schockaert" authorityYear="1977" class="Rhabditophora" family="Polycystididae" genus="Gyratrix" kingdom="Animalia" order="Rhabdocoela" pageId="6" pageNumber="7" phylum="Platyhelminthes" rank="species" species="proaviformis">
<emphasis id="B91BEADCFFA5FF58FC89E3AFFBF908D0" box="[887,1150,1936,1961]" italics="true" pageId="6" pageNumber="7">Gyratrix proaviformis</emphasis>
<bibRefCitation id="EFFE4B3FFFA5FF58FB74E3AFFF5208AB" author="Karling" pageId="6" pageNumber="26" refString="Karling, T. G. &amp; Schockaert, E. R. (1977) Anatomy and systematics of some Polycystididae (Turbellaria, Kalyptorhynchia) from the Pacific and S. Atlantic. Zoologica Scripta, 6, 5 - 19." type="journal article" year="1977">Karling &amp; Schockaert, 1977</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="DF106646FFA4FF59FF69E31CFD0E08BD" httpUri="https://zenodo.org/record/183375/files/figure.png" pageId="7" pageNumber="8" targetBox="[207,1381,232,1801]" targetPageId="7">
<paragraph id="8BD036CEFFA4FF59FF69E31CFD0E08BD" blockId="7.[151,1436,1827,1988]" pageId="7" pageNumber="8">
<emphasis id="B91BEADCFFA4FF59FF69E31CFE9C0842" bold="true" box="[151,283,1827,1851]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
<taxonomicName id="4C6F4D4DFFA4FF59FEDBE31CFDC70843" box="[293,576,1827,1850]" class="Rhabditophora" family="Polycystididae" genus="Paraustrorhynchus" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="8" phylum="Platyhelminthes" rank="species" species="neleae" status="sp. nov.">
<emphasis id="B91BEADCFFA4FF59FEDBE31CFDC70843" box="[293,576,1827,1850]" italics="true" pageId="7" pageNumber="8">Paraustrorhynchus neleae</emphasis>
</taxonomicName>
<emphasis id="B91BEADCFFA4FF59FDB7E31CFD000842" bold="true" box="[585,647,1827,1851]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A22857A7FFA4FF59FDB7E31CFD000842" box="[585,647,1827,1851]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A, Habitus (from a live animal). B, H, Prostate stylet type II (from the holotype). C, I, Prostate stylet type III (from the holotype).
<emphasis id="B91BEADCFFA4FF59FCE1E379FC070824" box="[799,896,1862,1885]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C6F4D4DFFA4FF59FCE1E379FCFB0824" box="[799,892,1862,1885]" class="Rhabditophora" family="Polycystididae" genus="Paraustrorhynchus" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="8" phylum="Platyhelminthes" rank="species" species="elixus">P. elixus</taxonomicName>
.
</emphasis>
D, Prostate stylet type III (from a specimen from Kenya). E, Prostate stylet type II (from the same specimen from Kenya). F, Prostate stylet type III (from a specimen from Brazil). G, Prostate stylet type II (from the same specimen from Brazil). J, Prostate stylet type III (from a specimen from New Caledonia). Arrows explained in the text.
</paragraph>
</caption>
<paragraph id="8BD036CEFFABFF56FF38E4A7FB010E2B" blockId="8.[151,1436,152,338]" pageId="8" pageNumber="9">
The measurements of the stylet, sheath and stalk of the new material are given in
<tableCitation id="C6ED0375FFABFF56FB5EE4A7FB7D0FCB" box="[1184,1274,152,178]" captionStart="TABLE 3" captionStartId="8.[151,239,391,415]" captionTargetBox="[159,1245,488,724]" captionTargetPageId="8" captionText="TABLE 3. Mean length (and SD) of stylet, sheath and stalk in G. hermaphroditus from different localities in the Indian Ocean." httpUri="http://table.plazi.org/id/DF106646FFABFF56FF69E5B8FF650EBF" pageId="8" pageNumber="9" tableUuid="DF106646FFABFF56FF69E5B8FF650EBF">Table 3</tableCitation>
. They all fall within the range known from the rest of the world. These measurements once again illustrate the huge variation occurring within this species complex. No clear delimitation of cryptic species can be made, probably because of the confounding effect of the existence of multiple sympatric cryptic species, which is known to occur in a very high degree in the species complex (Curini-Galletti &amp; Pucinelli 1990).
</paragraph>
</subSubSection>
</treatment>
</document>