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<document id="D2F5E79E0A722088EB1CF1D3347FA17D" ID-DOI="10.5252/zoosystema2024v46a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="10803267" ID-ZooBank="1B0779F3-FAE4-4105-8130-A26DE519721E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710145347001" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, M, Diego de, Bento, edeiros &amp; Ferreira, Rodrigo Lopes" docDate="2024" docId="03C687BFC619FFE41038F48D69DBF851" docLanguage="en" docName="zoosystema.2024.46.5.pdf" docOrigin="Zoosystema 46 (5)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Ctenorillo jequitinhonha Cardoso &amp; Ferreira 2024, n. sp." docType="treatment" docUuid="5CF8D003-88B7-465B-84EF-040FEC4FB7F2" docUuidSource="ZooBank" docVersion="2" lastPageNumber="127" masterDocId="FFFFFFC7C606FFC6136FF25C6C6BFFBA" masterDocTitle="Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation" masterLastPageNumber="132" masterPageNumber="95" pageNumber="124" updateTime="1710145921992" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2C2CD7A0D4233106D862911F3838BFDD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="602A3B514BFA48645856D243F9C189FF">
<mods:title id="181A0A4B75B472B29CEAD2ED2E270AA3">Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation</mods:title>
</mods:titleInfo>
<mods:name id="75CBEBCDFD7B7146414EBF12796B1240" type="personal">
<mods:role id="77AC8733958EF4DB39B6C64423B6D9FC">
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</mods:role>
<mods:namePart id="5BBD9E314CB36E70DE89D7AE940C079F">Cardoso, Giovanna Monticelli</mods:namePart>
<mods:affiliation id="C2AD489B026FA03E92B5CC7D546E367B">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) gmcardoso. bio @ gmail. com (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="39D1AB40E21186FFB9E77C9F01940C0B" type="email">bio@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="49C96D21F73A9356D9282525BD7B87FA" type="personal">
<mods:role id="FA6FDD3E320B909F1908BAE43C0627F7">
<mods:roleTerm id="C2905E49EDB01987DDFAE330DD237E6F">Author</mods:roleTerm>
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<mods:namePart id="921C5C715B53D7D498A0B9327BF70EFA">M, Diego de</mods:namePart>
</mods:name>
<mods:name id="F060D5A63BB480399DBB53DE706C45FF" type="personal">
<mods:role id="16006DEE8A03EE636796C5C1BCE7C867">
<mods:roleTerm id="60B8E2A6BAC760AA3D0C7D1826116759">Author</mods:roleTerm>
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<mods:namePart id="7D2A70C9F9F39617DC9A7FFEFEDABB58">Bento, edeiros</mods:namePart>
</mods:name>
<mods:name id="292F6B9463617C947408FD8EC73EC93F" type="personal">
<mods:role id="7F7CD22489613088DFA11253586D1AF2">
<mods:roleTerm id="53912BE9BD194A3FE6106B653D4964C3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4338550574D3D8C3E842A8FF027927DC">Ferreira, Rodrigo Lopes</mods:namePart>
<mods:affiliation id="C91D2CF1DC89F51546FEFE7DBD71C137">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) drops @ ufla. br</mods:affiliation>
<mods:nameIdentifier id="CA314B09153F8BF491FC9303C631FBE8" type="email">drops@ufla.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CDEDF5C9E7FAD7976EB99345A3A38CD0">text</mods:typeOfResource>
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<mods:title id="07B0B1DFF3D6370DC4FB5F0A5F9F13F2">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="F94CBB9A87BC2F6768B38F9F44E63186">
<mods:date id="F171BCB3F543770B75904653CFE76541">2024</mods:date>
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<mods:number id="ACCBC5E0CE57DC23E79CC3EB2CE0C717">2024-03-05</mods:number>
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<mods:number id="84C69552AFE559235E37514E33886337">46</mods:number>
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<mods:number id="1F9F6F1783B988DBBDA49BF352D370D0">5</mods:number>
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<mods:start id="85710DDC76974B6E54D438C4476F9395">95</mods:start>
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<mods:url id="B7DEB01FE0F23B6F029682987C71412A">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf</mods:url>
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<mods:classification id="0A6B7FDE987239E8C6C8A23B20DD54F4">journal article</mods:classification>
<mods:identifier id="B293127F04503B2988C3068CBF6BEF3E" type="DOI">10.5252/zoosystema2024v46a5</mods:identifier>
<mods:identifier id="B04431812FC927E9D2B7B85A06776E28" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="3A9D189648DD8326FB0944AFC11E8788" type="Zenodo-Dep">10803267</mods:identifier>
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<treatment id="03C687BFC619FFE41038F48D69DBF851" LSID="urn:lsid:zoobank.org:act:5CF8D003-88B7-465B-84EF-040FEC4FB7F2" httpUri="http://treatment.plazi.org/id/03C687BFC619FFE41038F48D69DBF851" lastPageId="34" lastPageNumber="127" pageId="31" pageNumber="124">
<subSubSection id="C3756522C619FFD91038F48D69EDF956" box="[855,1414,1745,1772]" pageId="31" pageNumber="124" type="nomenclature">
<paragraph id="8BD036A9C619FFD91038F48D69EDF956" blockId="31.[855,1414,1745,1804]" box="[855,1414,1745,1772]" pageId="31" pageNumber="124">
<heading id="D09881C5C619FFD91038F48D69EDF956" box="[855,1414,1745,1772]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="124" reason="2">
<taxonomicName id="4C6F4D2AC619FFD91038F48D692BF956" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[855,1344,1745,1772]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="31" pageNumber="124" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC619FFD91038F48D680FF956" bold="true" box="[855,1124,1745,1772]" italics="true" pageId="31" pageNumber="124">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C619FFD91624F48E69EDF956" box="[1355,1414,1746,1772]" pageId="31" pageNumber="124" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3756522C619FFD91088F4AE69FFF8FE" pageId="31" pageNumber="124" type="description">
<paragraph id="8BD036A9C619FFD91088F4AE689EF8B6" blockId="31.[855,1414,1745,1804]" box="[999,1269,1778,1804]" pageId="31" pageNumber="124">
(
<figureCitation id="13542A2CC619FFD91081F4AE680DF8B6" box="[1006,1126,1778,1804]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="32.[133,144,1381,1399]" captionStartId-1="33.[132,143,1368,1386]" captionStartId-2="34.[132,143,1501,1519]" captionTargetBox-0="[131,1457,214,1328]" captionTargetBox-1="[132,1455,216,1317]" captionTargetBox-2="[132,1455,219,1454]" captionTargetId-0="figure-345@32.[698,1267,215,772]" captionTargetId-1="figure-299@33.[132,1455,216,1317]" captionTargetId-2="figure-256@34.[132,1455,219,1464]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionText-0="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." captionText-1="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." captionText-2="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi-0="http://doi.org/10.5281/zenodo.10803319" figureDoi-1="http://doi.org/10.5281/zenodo.10803321" figureDoi-2="http://doi.org/10.5281/zenodo.10803323" httpUri-0="https://zenodo.org/record/10803319/files/figure.png" httpUri-1="https://zenodo.org/record/10803321/files/figure.png" httpUri-2="https://zenodo.org/record/10803323/files/figure.png" pageId="31" pageNumber="124">Figs 24-26</figureCitation>
;
<figureCitation id="13542A2CC619FFD9171EF4AE68F2F8B6" box="[1137,1177,1778,1804]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="31" pageNumber="124">27J</figureCitation>
;
<figureCitation id="13542A2CC619FFD917CAF4AE6887F8B6" box="[1189,1260,1778,1804]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="31" pageNumber="124">28J, K</figureCitation>
)
</paragraph>
<paragraph id="8BD036A9C619FFD91027F56E69FFF8FE" blockId="31.[840,1428,1842,1860]" box="[840,1428,1842,1860]" pageId="31" pageNumber="124">
<uri id="FFFE3AABC619FFD91027F56E69FFF8FE" box="[840,1428,1842,1860]" pageId="31" pageNumber="124">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3756522C619FFD91042F53B689DF852" pageId="31" pageNumber="124" type="materials_examined">
<paragraph id="8BD036A9C619FFD91042F53B689DF852" blockId="31.[811,1456,1894,2024]" pageId="31" pageNumber="124">
<materialsCitation id="3B073CF4C619FFD91042F53B68FFF83A" box="[813,1172,1895,1920]" collectionCode="TYPE, MATERIAL" country="Brazil" location="Brazil" pageId="31" pageNumber="124" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED7EAE6CC619FFD9100BF5306FA9F8C7" box="[868,962,1900,1917]" pageId="31" pageNumber="124">MATERIAL</collectionCode>
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. —
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<typeStatus id="54D4880BC619FFD91081F53B6824F8C5" box="[1006,1103,1895,1919]" pageId="31" pageNumber="124" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F3787639C619FFD91735F53468FFF83A" box="[1114,1172,1896,1920]" name="Brazil" pageId="31" pageNumber="124">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C619FFD917C4F53B69C7F809" collectingDate="2017-04-07" collectionCode="R" collectorName="L. Ferreira" country="Iceland" latitude="-16.4175" location="Jequitinhonha" longLatPrecision="21" longitude="-40.923058" municipality="Toca Juparana cave" pageId="31" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<specimenCount id="9D69FD20C619FFD917C4F53B68A5F8C4" box="[1195,1230,1895,1919]" count="1" pageId="31" pageNumber="124" type="male">1♂</specimenCount>
(
<quantity id="4C979B4CC619FFD917B2F53B6971F8C4" box="[1245,1306,1895,1919]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="124" unit="mm" value="7.0">7 mm</quantity>
, parts in slide),
<collectingRegion id="49ABF84BC619FFD91042F5DD6FB6F822" box="[813,989,1921,1945]" country="Brazil" name="Minas Gerais" pageId="31" pageNumber="124">Minas Gerais state</collectingRegion>
,
<location id="8EB06072C619FFD9108AF5DD6805F823" LSID="urn:lsid:plazi:treatment:03C687BFC619FFE41038F48D69DBF851:8EB06072C619FFD9108AF5DD6805F823" box="[997,1134,1921,1945]" country="Iceland" latitude="-16.4175" longLatPrecision="21" longitude="-40.923058" municipality="Toca Juparana cave" name="Jequitinhonha" pageId="31" pageNumber="124" stateProvince="Minas Gerais">Jequitinhonha</location>
municipality,
<collectingMunicipality id="6BB4ACD3C619FFD9179BF5DD69C0F822" box="[1268,1451,1921,1944]" pageId="31" pageNumber="124">Toca Juparanã cave</collectingMunicipality>
;
<geoCoordinate id="EE5B506EC619FFD91044F5C76FFBF809" box="[811,912,1947,1971]" degrees="16" direction="south" minutes="25" orientation="latitude" pageId="31" pageNumber="124" precision="15" seconds="3" value="-16.4175">16°25’3”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC619FFD910F6F5C7687EF809" box="[921,1045,1947,1971]" degrees="40" direction="west" minutes="55" orientation="longitude" pageId="31" pageNumber="124" precision="15" seconds="23" value="-40.923058">40°55’23”W</geoCoordinate>
;
<collectionCode id="ED7EAE6CC619FFD91770F5C06859F809" box="[1055,1074,1948,1971]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="124">R</collectionCode>
.
<collectorName id="269A537FC619FFD91759F5C068F2F809" box="[1078,1177,1947,1971]" pageId="31" pageNumber="124">L. Ferreira</collectorName>
leg.,
<date id="FFD11069C619FFD917A5F5C76944F809" box="[1226,1327,1947,1971]" pageId="31" pageNumber="124" value="2017-04-07">
<collectingDate id="EF95E981C619FFD917A5F5C76944F809" box="[1226,1327,1947,1971]" pageId="31" pageNumber="124" value="2017-04-07">7.IV.2017</collectingDate>
</date>
;
<collectingCountry id="F3787639C619FFD91658F5C76905F809" box="[1335,1390,1947,1971]" name="Iceland" pageId="31" pageNumber="124">ISLA</collectingCountry>
96804
</materialsCitation>
.
<materialsCitation id="3B073CF4C619FFD91044F5EA6FBBF875" box="[811,976,1974,1999]" country="Brazil" location="Brazil" pageId="31" pageNumber="124" specimenCount="1" typeStatus="paratype">
<emphasis id="B91BEABBC619FFD91044F5EA6FBBF875" bold="true" box="[811,976,1974,1999]" pageId="31" pageNumber="124">
<typeStatus id="54D4880BC619FFD91044F5EA6FE6F874" box="[811,909,1974,1998]" pageId="31" pageNumber="124" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3787639C619FFD910F9F5EB6FBBF875" box="[918,976,1975,1999]" name="Brazil" pageId="31" pageNumber="124">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C619FFD9108AF5EA69C4F874" box="[997,1455,1974,1998]" collectingDate="2017-04-07" collectionCode="R" collectorName="L. Ferreira" country="Iceland" latitude="-16.4175" location="Jequitinhonha" longLatPrecision="21" longitude="-40.923058" municipality="Toca Juparana cave" pageId="31" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D69FD20C619FFD9108AF5EA6861F877" box="[997,1034,1974,1998]" count="1" pageId="31" pageNumber="124" type="male">1 ♂</specimenCount>
in slide; same data as holotype;
<collectingCountry id="F3787639C619FFD91652F5EA6918F874" box="[1341,1395,1974,1998]" name="Iceland" pageId="31" pageNumber="124">ISLA</collectingCountry>
96806
</materialsCitation>
•
<materialsCitation id="3B073CF4C619FFD91052F58C6899F852" box="[829,1266,2000,2024]" collectingDate="2017-04-07" collectionCode="R" collectorName="L. Ferreira" country="Iceland" latitude="-16.4175" location="Jequitinhonha" longLatPrecision="21" longitude="-40.923058" municipality="Toca Juparana cave" pageId="31" pageNumber="124" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D69FD20C619FFD91052F58C6F0AF85D" box="[829,865,2000,2024]" count="3" pageId="31" pageNumber="124" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D69FD20C619FFD91002F58C6FE5F85D" box="[877,910,2000,2024]" count="8" pageId="31" pageNumber="124" type="female">8 ♀</specimenCount>
; same data as holotype;
<collectingCountry id="F3787639C619FFD91713F58C68DFF852" box="[1148,1204,2000,2024]" name="Iceland" pageId="31" pageNumber="124">ISLA</collectingCountry>
96805
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF106621C626FFE613EAF7396F7FFA1E" ID-DOI="http://doi.org/10.5281/zenodo.10803319" ID-Zenodo-Dep="10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125" startId="32.[133,144,1381,1399]" targetBox="[131,1457,214,1328]" targetPageId="32" targetType="figure">
<paragraph id="8BD036A9C626FFE613EAF7396F7FFA1E" blockId="32.[133,1457,1381,1444]" pageId="32" pageNumber="125">
<smallCapsWord id="8D36A075C626FFE613EAF7396CF6FACC" baselines="1394,1395" box="[133,157,1381,1399]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="32" pageNumber="125">FIG</smallCapsWord>
. 24. —
<taxonomicName id="4C6F4D2AC626FFE6138DF7396E38FACD" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[226,595,1381,1399]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="32" pageNumber="125" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC626FFE6138DF7396DC2FACD" box="[226,425,1381,1399]" italics="true" pageId="32" pageNumber="125">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C626FFE61134F7396EE6FACD" box="[603,653,1381,1399]" pageId="32" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
paratype (male, 7 mm, ISLA96806):
<emphasis id="B91BEABBC626FFE610ABF7396FBAFACD" bold="true" box="[964,977,1381,1399]" pageId="32" pageNumber="125">A</emphasis>
, habitus, lateral view;
<emphasis id="B91BEABBC626FFE617FEF73968F5FACD" bold="true" box="[1169,1182,1381,1399]" pageId="32" pageNumber="125">B</emphasis>
, disposition of dorsal tubercles;
<emphasis id="B91BEABBC626FFE613EAF7276CF8FA37" bold="true" box="[133,147,1403,1421]" pageId="32" pageNumber="125">C</emphasis>
, cephalon and epimeron 1, frontal view;
<emphasis id="B91BEABBC626FFE6128DF7206D9BFA34" bold="true" box="[482,496,1404,1422]" pageId="32" pageNumber="125">D</emphasis>
, epimera 1-3 ventral view;
<emphasis id="B91BEABBC626FFE611A1F7206EB1FA34" bold="true" box="[718,730,1404,1422]" pageId="32" pageNumber="125">E</emphasis>
, pleonites 3-5, uropods and telson, dorsal view;
<emphasis id="B91BEABBC626FFE61703F720681CFA34" bold="true" box="[1132,1143,1404,1422]" pageId="32" pageNumber="125">F</emphasis>
, antennula;
<emphasis id="B91BEABBC626FFE617B2F7276880FA37" bold="true" box="[1245,1259,1403,1421]" pageId="32" pageNumber="125">G</emphasis>
, antenna;
<emphasis id="B91BEABBC626FFE6162DF720693BFA34" bold="true" box="[1346,1360,1404,1422]" pageId="32" pageNumber="125">H</emphasis>
, right mandible;
<emphasis id="B91BEABBC626FFE613D8F7CE6CD6FA1E" bold="true" box="[183,189,1426,1444]" pageId="32" pageNumber="125">I</emphasis>
, left mandible;
<emphasis id="B91BEABBC626FFE61252F7CE6D2CFA1E" bold="true" box="[317,327,1426,1444]" pageId="32" pageNumber="125">J</emphasis>
, maxillula;
<emphasis id="B91BEABBC626FFE612CAF7CE6DD9FA1E" bold="true" box="[421,434,1426,1444]" pageId="32" pageNumber="125">K</emphasis>
, maxilla;
<emphasis id="B91BEABBC626FFE6116EF7CE6E67FA1E" bold="true" box="[513,524,1426,1444]" pageId="32" pageNumber="125">L</emphasis>
, maxilliped. Scale bar: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3756522C626FFE613EBF7AF6F63F9CE" pageId="32" pageNumber="125" type="etymology">
<paragraph id="8BD036A9C626FFE613EBF7AF6F63F9CE" blockId="32.[132,776,1522,1652]" pageId="32" pageNumber="125">
<smallCapsWord id="8D36A075C626FFE613EBF7AF6C96F9B3" baselines="1540,1541" box="[132,253,1523,1546]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="32" pageNumber="125">ETYMOLOGY</smallCapsWord>
. — The new species name is a noun in apposition that refers to the city and the Jequitinhonha River, one of the most important rivers in the north of
<collectingRegion id="49ABF84BC626FFE612F1F47B6E3AF985" box="[414,593,1575,1599]" country="Brazil" name="Minas Gerais" pageId="32" pageNumber="125">Minas Gerais state</collectingRegion>
; the
<typeStatus id="54D4880BC626FFE61111F4746ECCF9FA" box="[638,679,1576,1600]" pageId="32" pageNumber="125">type</typeStatus>
locality is within the drainage basin of the Jequitinhonha river. The word Jequitinhonha is of indigenous origin and means “wide river full of fish”.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C626FFE613EBF4D76E40F89C" pageId="32" pageNumber="125" type="diagnosis">
<paragraph id="8BD036A9C626FFE613EBF4D76E40F89C" blockId="32.[131,777,1674,1830]" pageId="32" pageNumber="125">
<smallCapsWord id="8D36A075C626FFE613EBF4D76C9AF91B" baselines="1692,1693" box="[132,241,1675,1698]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="32" pageNumber="125">DIAGNOSIS</smallCapsWord>
.— Dorsum covered with round tubercles with three rows on cephalon (4, 4, 4); three rows on pereonite 1 (2, 4, 7); two rows on pereonites 2-7 (6 + 4); and two paramedian tubercles on pleonites 3-5 and on telson. Pleopod 1 exopod with protruding distal portion, triangular, outer margin sinuous; endopod distal portion straight, twice longer than exopod.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C626FFE613EBF5616F63F8EE" box="[132,776,1852,1876]" pageId="32" pageNumber="125" type="distribution">
<paragraph id="8BD036A9C626FFE613EBF5616F63F8EE" blockId="32.[132,776,1852,1876]" box="[132,776,1852,1876]" pageId="32" pageNumber="125">
<smallCapsWord id="8D36A075C626FFE613EBF5616D7AF8E9" baselines="1870,1871" box="[132,273,1853,1876]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="32" pageNumber="125">DISTRIBUTION</smallCapsWord>
. — Jequitinhonha municipality,
<collectingRegion id="49ABF84BC626FFE6113FF5606F6FF8EE" box="[592,772,1852,1876]" country="Brazil" name="Minas Gerais" pageId="32" pageNumber="125">Minas Gerais state</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C626FFE413EBF52E686DF896" lastPageId="34" lastPageNumber="127" pageId="32" pageNumber="125" type="description">
<paragraph id="8BD036A9C626FFE613EBF52E6D7EF830" blockId="32.[131,775,1906,2027]" box="[132,277,1906,1932]" pageId="32" pageNumber="125">
<smallCapsWord id="8D36A075C626FFE613EBF52E6D7EF830" baselines="1925,1925" box="[132,277,1906,1932]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="32" pageNumber="125">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C626FFE713EBF5CD6D06F956" blockId="32.[131,775,1906,2027]" lastBlockId="33.[132,777,1491,1772]" lastPageId="33" lastPageNumber="126" pageId="32" pageNumber="125">
Maximum size:
<specimenCount id="9D69FD20C626FFE61256F5C96D3BF811" box="[313,336,1941,1963]" count="1" pageId="32" pageNumber="125" type="male">♂</specimenCount>
,
<quantity id="4C979B4CC626FFE61230F5CE6DCDF816" box="[351,422,1938,1964]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="125" unit="mm" value="7.0">7 mm</quantity>
,
<specimenCount id="9D69FD20C626FFE612DDF5C96DADF811" box="[434,454,1941,1963]" count="1" pageId="32" pageNumber="125" type="female">♀</specimenCount>
,
<quantity id="4C979B4CC626FFE612BAF5CD6E77F816" box="[469,540,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="32" pageNumber="125" unit="mm" value="9.0">9 mm</quantity>
. Color grey with lateral margins depigmented (
<figureCitation id="13542A2CC626FFE612DDF5ED6E50F871" box="[434,571,1969,1995]" captionStart="FIG" captionStartId="34.[132,143,1501,1519]" captionTargetBox="[132,1455,219,1454]" captionTargetId="figure-256@34.[132,1455,219,1464]" captionTargetPageId="34" captionText="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi="http://doi.org/10.5281/zenodo.10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="32" pageNumber="125">Fig. 26C, D</figureCitation>
). Dorsum covered with round tubercles (
<figureCitation id="13542A2CC626FFE612EDF58D6E60F851" box="[386,523,2001,2027]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Fig. 24A, B</figureCitation>
): cephalon with three rows (4, 4, and 4 from front to back of vertex); pereonite 1 with
<quantity id="4C979B4CC626FFE61007F44F6FCEF996" box="[872,933,1554,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="32" pageNumber="125" unit="in" value="13.0">13 in</quantity>
three rows (2, 4 and 7); pereonites 2-7 with 10 (6+ 4); pleonites 3-5 and telson with two paramedian tubercles. Dorsal cuticle (
<figureCitation id="13542A2CC626FFE61724F40E68A1F9D6" box="[1099,1226,1618,1644]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="32" pageNumber="125">Fig. 28J, K</figureCitation>
) with short triangular scale setae, pereonites 1-7 bearing one line of
<emphasis id="B91BEABBC626FFE61607F42E6FEAF916" italics="true" pageId="32" pageNumber="125">noduli lateralis</emphasis>
per side on outer surface of posterior tubercle of second tubercle. Cephalon with frontal shield protruding above vertex; eye consisting of 16 ommatidia (
<figureCitation id="13542A2CC626FFE61650F48E69C1F956" box="[1343,1450,1746,1772]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Figs 24C</figureCitation>
;
<figureCitation id="13542A2CC626FFE61042F4AE6F33F8B6" box="[813,856,1778,1804]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="32" pageNumber="125">28J</figureCitation>
). Pereonite 1 with posterior margin sinuous at sides; inner lobe of schisma rounded, extending beyond posterior margin of outer lobe; pereonite 2 with triangular ventral tooth reaching epimeron posterior margin (
<figureCitation id="13542A2CC626FFE6165DF50D69C8F8D6" box="[1330,1443,1873,1900]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Fig. 24D</figureCitation>
). Pereonites 2-7 (
<figureCitation id="13542A2CC626FFE610B4F52D6856F831" box="[987,1085,1905,1931]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Fig. 24A</figureCitation>
) with wide quadrangular epimera slightly bent outwards. Telson (
<figureCitation id="13542A2CC626FFE61712F5CD68B6F811" box="[1149,1245,1937,1963]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Fig. 24E</figureCitation>
) hourglass-shaped, proximal portion broader than distal portion. Antennula (
<figureCitation id="13542A2CC626FFE6105AF58D6FF7F851" box="[821,924,2001,2027]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="32" pageNumber="125">Fig. 24F</figureCitation>
) of three articles, second article shorter than first and third, third article with seven apical aesthetascs. Antenna (
<figureCitation id="13542A2CC627FFE71395F7AF6D0CF9B7" box="[250,359,1523,1549]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="33" pageNumber="126">Fig. 24G</figureCitation>
) short and stout, flagellum shorter than fifth article of peduncle; second article of flagellum about three times as long as first. Buccal pieces as
<taxonomicName id="4C6F4D2AC627FFE711DEF46F6DE3F9D6" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="33" pageNumber="126" phylum="Arthropoda" rank="species" species="pelado" status="n. sp.">
<emphasis id="B91BEABBC627FFE711DEF46F6CF5F9D6" italics="true" pageId="33" pageNumber="126">C. pelado</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C627FFE712FAF40F6DBFF9D7" box="[405,468,1619,1645]" pageId="33" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13542A2CC627FFE7128AF40E6E5CF9D6" box="[485,567,1618,1644]" captionStart="FIG" captionStartId="32.[133,144,1381,1399]" captionTargetBox="[131,1457,214,1328]" captionTargetId="figure-345@32.[698,1267,215,772]" captionTargetPageId="32" captionText="FIG. 24. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. paratype (male, 7 mm, ISLA96806): A, habitus, lateral view; B, disposition of dorsal tubercles; C, cephalon and epimeron 1, frontal view; D, epimera 1-3 ventral view; E, pleonites 3-5, uropods and telson, dorsal view;F, antennula; G, antenna; H, right man- dible; I, left mandible; J, maxillula; K, maxilla; L, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803319" httpUri="https://zenodo.org/record/10803319/files/figure.png" pageId="33" pageNumber="126">Fig. 24</figureCitation>
H-L). All pleopod exopods with monospiracular covered lungs. Uropod (
<figureCitation id="13542A2CC627FFE713E3F4CE6C9BF917" box="[140,240,1682,1709]" captionStart="FIG" captionStartId="33.[132,143,1368,1386]" captionTargetBox="[132,1455,216,1317]" captionTargetId="figure-299@33.[132,1455,216,1317]" captionTargetPageId="33" captionText="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126">Fig. 25A</figureCitation>
) protopod flattened, enlarged on basal portion; exopod outward, short, inserted dorsally close to medial margin of protopod.
</paragraph>
<caption id="DF106621C627FFE713EBF7046FC8FA3B" ID-DOI="http://doi.org/10.5281/zenodo.10803321" ID-Zenodo-Dep="10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126" startId="33.[132,143,1368,1386]" targetBox="[132,1455,216,1317]" targetPageId="33" targetType="figure">
<paragraph id="8BD036A9C627FFE713EBF7046FC8FA3B" blockId="33.[132,1455,1368,1409]" pageId="33" pageNumber="126">
<smallCapsWord id="8D36A075C627FFE713EBF7046CF6FAD3" baselines="1381,1382" box="[132,157,1368,1386]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="33" pageNumber="126">FIG</smallCapsWord>
. 25. —
<taxonomicName id="4C6F4D2AC627FFE713B0F7046E21FAD0" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[223,586,1368,1386]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="33" pageNumber="126" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC627FFE713B0F7046DCEFAD0" box="[223,421,1368,1386]" italics="true" pageId="33" pageNumber="126">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C627FFE7113DF7046EEEFAD0" box="[594,645,1368,1386]" pageId="33" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
, paratype (male, 7 mm, ISLA96806):
<emphasis id="B91BEABBC627FFE710D7F7046FAEFAD0" bold="true" box="[952,965,1368,1386]" pageId="33" pageNumber="126">A</emphasis>
, uropod,
<emphasis id="B91BEABBC627FFE71779F7046848FAD0" bold="true" box="[1046,1059,1368,1386]" pageId="33" pageNumber="126">B</emphasis>
, pereopod 1;
<emphasis id="B91BEABBC627FFE717F6F70468CCFAD0" bold="true" box="[1177,1191,1368,1386]" pageId="33" pageNumber="126">C</emphasis>
, pereopod 7;
<emphasis id="B91BEABBC627FFE71672F7046940FAD0" bold="true" box="[1309,1323,1368,1386]" pageId="33" pageNumber="126">D</emphasis>
, pleopod 1 and genital papilla;
<emphasis id="B91BEABBC627FFE7126DF7336D65FA3B" bold="true" box="[258,270,1391,1409]" pageId="33" pageNumber="126">E</emphasis>
, pleopod 2;
<emphasis id="B91BEABBC627FFE71218F7336DE9FA3B" bold="true" box="[375,386,1391,1409]" pageId="33" pageNumber="126">F</emphasis>
, pleopod 3 exopod;
<emphasis id="B91BEABBC627FFE7115FF7326E55FA3A" bold="true" box="[560,574,1390,1408]" pageId="33" pageNumber="126">G</emphasis>
, pleopod 4 exopod;
<emphasis id="B91BEABBC627FFE71184F7336E92FA3B" bold="true" box="[747,761,1391,1409]" pageId="33" pageNumber="126">H</emphasis>
, pleopod 5 exopod.
</paragraph>
</caption>
<paragraph id="8BD036A9C627FFE713EBF54E6CD2F896" blockId="33.[131,777,1810,2028]" box="[132,185,1810,1836]" pageId="33" pageNumber="126">
<emphasis id="B91BEABBC627FFE713EBF54E6CD2F896" box="[132,185,1810,1836]" italics="true" pageId="33" pageNumber="126">Male</emphasis>
</paragraph>
<paragraph id="8BD036A9C627FFE713EBF56D6D33F856" blockId="33.[131,777,1810,2028]" pageId="33" pageNumber="126">
Pereopods without particular modifications (
<figureCitation id="13542A2CC627FFE7111DF56D6E92F8F6" box="[626,761,1841,1868]" captionStart="FIG" captionStartId="33.[132,143,1368,1386]" captionTargetBox="[132,1455,216,1317]" captionTargetId="figure-299@33.[132,1455,216,1317]" captionTargetPageId="33" captionText="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126">Fig. 25B, C</figureCitation>
). Pleopod 1 exopod (
<figureCitation id="13542A2CC627FFE7120EF50D6DA3F8D6" box="[353,456,1873,1900]" captionStart="FIG" captionStartId="33.[132,143,1368,1386]" captionTargetBox="[132,1455,216,1317]" captionTargetId="figure-299@33.[132,1455,216,1317]" captionTargetPageId="33" captionText="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126">Fig. 25D</figureCitation>
) as wide as long, protruding distal portion, triangular, outer margin sinuous; endopod twice longer than exopod, distal portion straight. Pleopod 2 endopod as long as exopod (
<figureCitation id="13542A2CC627FFE712C2F5ED6E67F876" box="[429,524,1969,1996]" captionStart="FIG" captionStartId="33.[132,143,1368,1386]" captionTargetBox="[132,1455,216,1317]" captionTargetId="figure-299@33.[132,1455,216,1317]" captionTargetPageId="33" captionText="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126">Fig. 25E</figureCitation>
). Pleopods 3-5 exopods as in
<figureCitation id="13542A2CC627FFE713D4F58D6D48F856" box="[187,291,2001,2028]" captionStart="FIG" captionStartId="33.[132,143,1368,1386]" captionTargetBox="[132,1455,216,1317]" captionTargetId="figure-299@33.[132,1455,216,1317]" captionTargetPageId="33" captionText="FIG. 25. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp., paratype (male, 7 mm, ISLA96806): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803321" httpUri="https://zenodo.org/record/10803321/files/figure.png" pageId="33" pageNumber="126">Figure 25</figureCitation>
F-H.
</paragraph>
<paragraph id="8BD036A9C627FFE71042F78F6FE6FA51" blockId="33.[811,1457,1491,2027]" box="[813,909,1491,1517]" pageId="33" pageNumber="126">
<smallCapsWord id="8D36A075C627FFE71042F78F6FE6FA51" baselines="1510,1510" box="[813,909,1491,1517]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitat" pageId="33" pageNumber="126">HABITAT</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C627FFE71042F7AF68E8F896" blockId="33.[811,1457,1491,2027]" pageId="33" pageNumber="126">
Specimens of
<taxonomicName id="4C6F4D2AC627FFE710A7F7AF690EF9B7" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[968,1381,1523,1549]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="33" pageNumber="126" phylum="Arthropoda" rank="species" species="jequitinhonha" status="n. sp.">
<emphasis id="B91BEABBC627FFE710A7F7AF68EBF9B7" box="[968,1152,1523,1549]" italics="true" pageId="33" pageNumber="126">C. jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C627FFE7161DF7AF69DBF9B7" box="[1394,1456,1523,1549]" pageId="33" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
were found in Toca Juparanã, a granite cave located in Jequitinhonha municipality, in northeastern
<collectingRegion id="49ABF84BC627FFE717B4F46E69C0F9F7" box="[1243,1451,1586,1613]" country="Brazil" name="Minas Gerais" pageId="33" pageNumber="126">Minas Gerais state</collectingRegion>
,
<collectingCountry id="F3787639C627FFE71041F40E6F1BF9D6" box="[814,880,1618,1644]" name="Brazil" pageId="33" pageNumber="126">Brazil</collectingCountry>
. The average annual temperature is around 25°C, with the minimum temperature of approximately 15°C and the maximum of 38°C. The rainy season lasts from October to April, with December being the wettest month (
<quantity id="4C979B4CC627FFE7165AF4EE69F8F977" box="[1333,1427,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="33" pageNumber="126" unit="mm" value="140.0">140 mm</quantity>
of precipitation). The dry season lasts from May to September, with July being the driest month (average of
<quantity id="4C979B4CC627FFE71674F4AE6936F8B6" box="[1307,1373,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="126" unit="mm" value="3.0">3 mm</quantity>
of precipitation) (
<bibRefCitation id="EFFE4B58C627FFE710C2F54F6818F896" author="ALVARES C. A. &amp; STAPE J. L. &amp; SENTELHAS P. C. &amp; MORAES G. &amp; DE &amp; LEONARDO J. &amp; SPAROVEK G." box="[941,1139,1810,1839]" pageId="33" pageNumber="126" pagination="711 - 728" refId="ref31163" refString="ALVARES C. A., STAPE J. L., SENTELHAS P. C., MORAES G. DE, LEONARDO J. &amp; SPAROVEK G. 2013. - Koppen's climate classification map for Brazil. Meteorologische Zeitschrift 22 (6): 711 - 728. https: // doi. org / 10.1127 / 0941 - 2948 / 2013 / 0507" type="journal article" year="2013">
Álvares
<emphasis id="B91BEABBC627FFE7176EF54E685EF896" box="[1025,1077,1810,1836]" italics="true" pageId="33" pageNumber="126">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036A9C627FFE4102CF56D6E95F8D6" blockId="33.[811,1457,1491,2027]" lastBlockId="34.[130,776,1618,2028]" lastPageId="34" lastPageNumber="127" pageId="33" pageNumber="126">
The Toca Juparanã cave is a granitic talus cave that was formed by water removal of smaller sediments between large collapsed granitic blocks. The cave has two entrances of irregular shape, delineated by the contact of boulders (
<figureCitation id="13542A2CC627FFE71629F5CD69CFF811" box="[1350,1444,1937,1963]" captionStart="FIG" captionStartId="34.[132,143,1501,1519]" captionTargetBox="[132,1455,219,1454]" captionTargetId="figure-256@34.[132,1455,219,1464]" captionTargetPageId="34" captionText="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi="http://doi.org/10.5281/zenodo.10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="33" pageNumber="126">Fig. 26A</figureCitation>
). The main chamber of the cave has an irregular floor made up of collapsed boulders (
<figureCitation id="13542A2CC627FFE7174BF58D68EDF851" box="[1060,1158,2001,2027]" captionStart="FIG" captionStartId="34.[132,143,1501,1519]" captionTargetBox="[132,1455,219,1454]" captionTargetId="figure-256@34.[132,1455,219,1464]" captionTargetPageId="34" captionText="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi="http://doi.org/10.5281/zenodo.10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="33" pageNumber="126">Fig. 26B</figureCitation>
) and is euphotic/disphotic due to the presence of two entrances located on either side of the chamber. There is a descending aphotic conduit from this chamber, which is crossed by a small hypogenic drainage. Specimens of
<taxonomicName id="4C6F4D2AC624FFE4127EF4EE6EFAF976" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[273,657,1714,1740]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="jequitinhonha" status="n. sp.">
<emphasis id="B91BEABBC624FFE4127EF4EE6DD5F976" box="[273,446,1714,1740]" italics="true" pageId="34" pageNumber="127">C. jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE411F5F4EF6EB9F977" box="[666,722,1715,1741]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
were found under fallen rocks in this conduit trespassed by the small drainage, although the specimens were not observed close to the mainstream passage. The only visible organic resource in this area was plant debris brought by water, indicating that the isopods were likely feeding on this resource (
<figureCitation id="13542A2CC624FFE4110DF50D6E9BF8D6" box="[610,752,1873,1900]" captionStart="FIG" captionStartId="34.[132,143,1501,1519]" captionTargetBox="[132,1455,219,1454]" captionTargetId="figure-256@34.[132,1455,219,1464]" captionTargetPageId="34" captionText="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi="http://doi.org/10.5281/zenodo.10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="34" pageNumber="127">Fig. 26 C, D</figureCitation>
).
</paragraph>
<caption id="DF106621C624FFE413EBF7816DFEF9A6" ID-DOI="http://doi.org/10.5281/zenodo.10803323" ID-Zenodo-Dep="10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="34" pageNumber="127" startId="34.[132,143,1501,1519]" targetBox="[132,1455,219,1454]" targetPageId="34" targetType="figure">
<paragraph id="8BD036A9C624FFE413EBF7816DFEF9A6" blockId="34.[132,1457,1500,1564]" pageId="34" pageNumber="127">
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. 26. —
<taxonomicName id="4C6F4D2AC624FFE4138FF7806E20FA55" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[224,587,1500,1519]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC624FFE4138FF7806DCEFA55" box="[224,421,1500,1519]" italics="true" pageId="34" pageNumber="127">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE4113CF7816EEEFA55" box="[595,645,1501,1519]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B91BEABBC624FFE411E3F7816EF2FA55" bold="true" box="[652,665,1501,1519]" pageId="34" pageNumber="127">A</emphasis>
, detail of the Toca do Juparanã cave entrance;
<emphasis id="B91BEABBC624FFE41746F781685DFA55" bold="true" box="[1065,1078,1501,1519]" pageId="34" pageNumber="127">B</emphasis>
, inner portion of the Toca do Juparanã cave;
<emphasis id="B91BEABBC624FFE413EBF7AF6CF9F9BF" bold="true" box="[132,146,1523,1541]" pageId="34" pageNumber="127">C</emphasis>
,
<emphasis id="B91BEABBC624FFE413F2F7AF6CC0F9BF" bold="true" box="[157,171,1523,1541]" pageId="34" pageNumber="127">D</emphasis>
,
<taxonomicName id="4C6F4D2AC624FFE413D9F7AF6E49F9BF" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[182,546,1523,1541]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC624FFE413D9F7AF6D17F9BF" box="[182,380,1523,1541]" italics="true" pageId="34" pageNumber="127">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE41144F7AF6E30F9BF" box="[555,603,1523,1541]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
habitus in natural habitat;
<emphasis id="B91BEABBC624FFE41054F7AF6F2CF9BF" bold="true" box="[827,839,1523,1541]" pageId="34" pageNumber="127">E</emphasis>
, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen.
</paragraph>
</caption>
<paragraph id="8BD036A9C624FFE413F4F52D686DF896" blockId="34.[130,776,1618,2028]" lastBlockId="34.[813,1457,1618,1836]" pageId="34" pageNumber="127">
The vegetation surrounding the cave represents a well-preserved secondary forest, although many areas in the region are deforested for pastures and monocultures, especially near the Jequitinhonha River floodplain (
<figureCitation id="13542A2CC624FFE4128FF58D6E2AF851" box="[480,577,2001,2027]" captionStart="FIG" captionStartId="34.[132,143,1501,1519]" captionTargetBox="[132,1455,219,1454]" captionTargetId="figure-256@34.[132,1455,219,1464]" captionTargetPageId="34" captionText="FIG. 26. — Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: A, detail of the Toca do Juparanã cave entrance; B, inner portion of the Toca do Juparanã cave; C, D, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. habitus in natural habitat; E, landscape at the surroundings of the Toca do Juparanã cave, where the Jequitinhonha river can be seen." figureDoi="http://doi.org/10.5281/zenodo.10803323" httpUri="https://zenodo.org/record/10803323/files/figure.png" pageId="34" pageNumber="127">Fig. 26E</figureCitation>
). The cave receives no visitors and is unknown to most of the local population. As for other
<taxonomicName id="4C6F4D2AC624FFE410D8F42E6874F936" authorityName="Verhoeff" authorityYear="1942" box="[951,1055,1650,1676]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABBC624FFE410D8F42E6874F936" box="[951,1055,1650,1676]" italics="true" pageId="34" pageNumber="127">Ctenorillo</emphasis>
</taxonomicName>
herein described, no samplings were conducted outside caves in the area, thus
<taxonomicName id="4C6F4D2AC624FFE41793F4CF686BF976" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="jequitinhonha" status="n. sp.">
<emphasis id="B91BEABBC624FFE41793F4CF69DAF916" box="[1276,1457,1682,1709]" italics="true" pageId="34" pageNumber="127">C. jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE41765F4EF682FF977" box="[1034,1092,1715,1741]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
is likely to occur in external habitats. Therefore, it is advisable that further studies including external samplings be conducted to better understand the species distribution.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C624FFE41042F50E69DBF851" pageId="34" pageNumber="127" type="discussion">
<paragraph id="8BD036A9C624FFE41042F50E6FECF8D0" blockId="34.[811,1456,1874,2028]" box="[813,903,1874,1900]" pageId="34" pageNumber="127">
<smallCapsWord id="8D36A075C624FFE41042F50E6FECF8D0" baselines="1893,1893" box="[813,903,1874,1900]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remark" pageId="34" pageNumber="127">REMARK</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C624FFE41042F52D69DBF851" blockId="34.[811,1456,1874,2028]" pageId="34" pageNumber="127">
<taxonomicName id="4C6F4D2AC624FFE41042F52D6898F831" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[813,1267,1905,1931]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="jequitinhonha" status="sp. nov.">
<emphasis id="B91BEABBC624FFE41042F52D6848F831" box="[813,1059,1905,1931]" italics="true" pageId="34" pageNumber="127">Ctenorillo jequitinhonha</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE41794F52E6959F836" box="[1275,1330,1906,1932]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
differs from the Brazilian species by the round shape of tubercles, resembling
<taxonomicName id="4C6F4D2AC624FFE41004F5EE68D4F871" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; M &amp; Bento &amp; Ferreira" authorityYear="2024" box="[875,1215,1969,1996]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="34" pageNumber="127" phylum="Arthropoda" rank="species" species="iuiuensis" status="n. sp.">
<emphasis id="B91BEABBC624FFE41004F5EE6F8FF871" box="[875,996,1970,1996]" italics="true" pageId="34" pageNumber="127">C. iuiuensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C624FFE417A6F5EE696FF876" box="[1225,1284,1970,1996]" pageId="34" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
by the presence of two paramedian tubercles on pleonites 3-5 and on telson.
</paragraph>
</subSubSection>
</treatment>
</document>
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