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<document id="1EC8119BBAD3DB1D4C0CCDF9DA5011A6" ID-CLB-Dataset="22059" ID-DOI="10.5852/ejt.2017.350" ID-GBIF-Dataset="52e4ae01-f51f-4c4a-8a14-9fb5b17788d6" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832682" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831660737" checkinUser="carolina" docAuthor="Fuhrmann, Juares &amp; Vaz-De-Mello, Fernando Z." docDate="2017" docId="03C687BD4608324FFDDEF8DADBDBD3C4" docLanguage="en" docName="ejt-2017-350.pdf.imf" docOrigin="European Journal of Taxonomy 350" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Calodactylus Blanchard 1850" docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFFFFFC5461D3259FFB9FFA0DF34D174" masterDocTitle="Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification" masterLastPageNumber="71" masterPageNumber="1" pageNumber="22" updateTime="1698840167045" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F9FE79CD59305E853CD076F8C494F9D1">Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification</mods:title>
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<mods:namePart id="52A2011B3983A9A8E7CCF262DB62186C">Fuhrmann, Juares</mods:namePart>
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<mods:namePart id="75EC4C8E3DC86BA20EBF2127A88DD0A8">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<treatment id="03C687BD4608324FFDDEF8DADBDBD3C4" ID-DOI="http://doi.org/10.5281/zenodo.3851134" ID-GBIF-Taxon="164167392" ID-Zenodo-Dep="3851134" LSID="urn:lsid:plazi:treatment:03C687BD4608324FFDDEF8DADBDBD3C4" httpUri="http://treatment.plazi.org/id/03C687BD4608324FFDDEF8DADBDBD3C4" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C37565204608324CFDDEF8DADCF9D6E1" box="[615,973,1914,1941]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BD036AB4608324CFDDEF8DADCF9D6E1" blockId="21.[189,973,1914,1978]" box="[615,973,1914,1941]" pageId="21" pageNumber="22">
<heading id="D09881C74608324CFDDEF8DADCF9D6E1" box="[615,973,1914,1941]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="4C6F4D284608324CFDDEF8DADCF9D6E1" ID-CoL="3FTG" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[615,973,1914,1941]" class="Insecta" family="Scarabaeidae" genus="Calodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB94608324CFDDEF8DADC37D6E1" bold="true" box="[615,771,1914,1941]" italics="true" pageId="21" pageNumber="22">Calodactylus</emphasis>
<bibRefCitation id="EFFE4B5A4608324CFCB2F8DADCF9D6E1" author="Blanchard" box="[779,973,1914,1941]" firstAuthor="Blanchard" pageId="21" pageNumber="22" refId="ref27119" refString="Blanchard C. E. 1850. Classe des insectes. Ordre des coleopteres. In: Milne-Edwards H., Blanchard C. E. &amp; Lucas H. (eds) Catalogue de la collection entomologique du Museum d'Histoire naturelle de Paris. Gide et Baudry, Paris." type="book" year="1850">Blanchard, 1850</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37565204608324CFF04F800DC9ED69F" pageId="21" pageNumber="22" type="type_taxon">
<paragraph id="8BD036AB4608324CFF04F800DE6CD6CE" blockId="21.[189,973,1914,1978]" box="[189,344,1952,1978]" pageId="21" pageNumber="22">
<heading id="D09881C74608324CFF04F800DE6CD6CE" bold="true" box="[189,344,1952,1978]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B91BEAB94608324CFF04F800DE6CD6CE" bold="true" box="[189,344,1952,1978]" pageId="21" pageNumber="22">
<typeStatus id="54D488094608324CFF04F800DFCED6CE" box="[189,250,1952,1978]" pageId="21" pageNumber="22">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB4608324CFF04F871DC9ED69F" blockId="21.[189,938,2000,2027]" box="[189,938,2000,2027]" pageId="21" pageNumber="22">
<taxonomicName id="4C6F4D284608324CFF04F871DD4ED69F" ID-CoL="PXV8" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[189,634,2000,2027]" class="Insecta" family="Scarabaeidae" genus="Calodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B91BEAB94608324CFF04F871DE85D69F" box="[189,433,2001,2027]" italics="true" pageId="21" pageNumber="22">Calodactylus tibialis</emphasis>
<bibRefCitation id="EFFE4B5A4608324CFE01F871DD4ED69F" author="Blanchard" box="[440,634,2000,2027]" firstAuthor="Blanchard" pageId="21" pageNumber="22" refId="ref27119" refString="Blanchard C. E. 1850. Classe des insectes. Ordre des coleopteres. In: Milne-Edwards H., Blanchard C. E. &amp; Lucas H. (eds) Catalogue de la collection entomologique du Museum d'Histoire naturelle de Paris. Gide et Baudry, Paris." type="book" year="1850">Blanchard, 1850</bibRefCitation>
</taxonomicName>
(designation: monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C3756520460B324FFF04FEABDD6BD0D7" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BD036AB460B324FFF04FEABDE02D051" blockId="22.[189,310,267,293]" box="[189,310,267,293]" pageId="22" pageNumber="23">
<heading id="D09881C7460B324FFF04FEABDE02D051" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B91BEAB9460B324FFF04FEABDE02D051" bold="true" box="[189,310,267,293]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB460B324FFF04FE9DDD6BD0D7" blockId="22.[189,1398,316,419]" pageId="22" pageNumber="23">
Prosternum with an anterior longitudinal carina; protibia with two teeth and with a spur; meso- and metatibiae medially enlarged (
<figureCitation id="13542A2E460B324FFD9DFEC2DDB4D009" box="[548,640,354,381]" captionStart="Fig" captionStartId="23.[189,232,1929,1955]" captionTargetBox="[240,1381,823,1878]" captionTargetId="figure@23.[189,1398,815,1886]" captionTargetPageId="23" captionText="Fig. 7. Calodactylus tibialis Blanchard, 1850. A. Male habitus, lateral (without some appendages). B. Male abdomen lateroventral detail. CD. Aedeagus (lateral, parameres apex). E. Metatibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832696" httpUri="https://zenodo.org/record/3832696/files/figure.png" pageId="22" pageNumber="23">Fig. 7E</figureCitation>
); meso- and metatarsal claws simple; male ventrite VI with two parallel and setose carina (
<figureCitation id="13542A2E460B324FFE4CFE28DD66D0D7" box="[501,594,392,419]" captionStart="Fig" captionStartId="23.[189,232,1929,1955]" captionTargetBox="[240,1381,823,1878]" captionTargetId="figure@23.[189,1398,815,1886]" captionTargetPageId="23" captionText="Fig. 7. Calodactylus tibialis Blanchard, 1850. A. Male habitus, lateral (without some appendages). B. Male abdomen lateroventral detail. CD. Aedeagus (lateral, parameres apex). E. Metatibia. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832696" httpUri="https://zenodo.org/record/3832696/files/figure.png" pageId="22" pageNumber="23">Fig. 7B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756520460B324FFF04FE6DDBDBD3C4" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BD036AB460B324FFF04FE6DDE1BD093" blockId="22.[189,303,461,487]" box="[189,303,461,487]" pageId="22" pageNumber="23">
<heading id="D09881C7460B324FFF04FE6DDE1BD093" bold="true" box="[189,303,461,487]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B91BEAB9460B324FFF04FE6DDE1BD093" bold="true" box="[189,303,461,487]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB460B324FFF04FE5EDBDBD3C4" blockId="22.[189,1399,510,688]" pageId="22" pageNumber="23">
The genera
<taxonomicName id="4C6F4D28460B324FFEF1FE5EDED7D36C" authorityName="Blanchard" authorityYear="1850" box="[328,483,510,536]" class="Insecta" family="Scarabaeidae" genus="Calodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFEF1FE5EDED7D36C" box="[328,483,510,536]" italics="true" pageId="22" pageNumber="23">Calodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D28460B324FFDA5FE5FDBC6D36C" authority="LePeletier de Saint-Fargeau &amp; Audinet-Serville, 1828" authorityName="LePeletier de Saint-Fargeau &amp; Audinet-Serville" authorityYear="1828" box="[540,1266,510,536]" class="Insecta" family="Scarabaeidae" genus="Dasyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFDA5FE5FDD46D36C" box="[540,626,511,536]" italics="true" pageId="22" pageNumber="23">Dasyus</emphasis>
<bibRefCitation id="EFFE4B5A460B324FFDC3FE5EDBC6D36C" author="LePeletier de Saint-Fargeau &amp; Audinet-Serville" box="[634,1266,510,536]" firstAuthor="LePeletier de Saint-Fargeau" pageId="22" pageNumber="23" pagination="345 - 832" refId="ref29420" refString="LePeletier de Saint-Fargeau A. L. M. &amp; Audinet-Serville J. G. 1828. Scarabe, Scarabaeus. In: Latreille, LePeletier de Saint Fargeau, Serville &amp; Guerin-Meneville. (eds). 1825 - 1828. Tome 10. Part 2. In: Latreille (ed.). Histoire naturelle. Entomologie, ou histoire naturelle de crustaces, des arachnides et des insectes. In: Encyclopedie methodique, ou par ordre de matieres; par une societe de gens de lettres, de savants et d'artistes; precedee d'un vocabulaire universel, servant de table pour tout l'ouvrage, ornee des portraits de MM. Diderot et d'Alembert, premiers editeurs de l'Encyclopedie: 346 - 382. Chez Mme Veuve Agasse, Paris, 832 p + [1 errata]. [1825: tome 10, part 1, 1 - 344; 1828: tome 10, part 2, 345 - 832]." type="book chapter" year="1828">LePeletier de Saint-Fargeau &amp; Audinet-Serville, 1828</bibRefCitation>
</taxonomicName>
are similar to each other. They can be distinguished by enlargements of the meso- and metatibiae,
<taxonomicName id="4C6F4D28460B324FFB65FD84DA43D34A" authorityName="Blanchard" authorityYear="1850" box="[1244,1399,548,574]" class="Insecta" family="Scarabaeidae" genus="Calodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFB65FD84DA43D34A" box="[1244,1399,548,574]" italics="true" pageId="22" pageNumber="23">Calodactylus</emphasis>
</taxonomicName>
have a medial enlargement whereas
<taxonomicName id="4C6F4D28460B324FFDDEFDEBDD89D310" authorityName="LePeletier de Saint-Fargeau &amp; Audinet-Serville" authorityYear="1828" box="[615,701,587,612]" class="Insecta" family="Scarabaeidae" genus="Dasyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFDDEFDEBDD89D310" box="[615,701,587,612]" italics="true" pageId="22" pageNumber="23">Dasyus</emphasis>
</taxonomicName>
have medial and distal areas of the same width. An evident characteristic of these genera is the simple meso- and metatarsal claws, which also occurs in male
<taxonomicName id="4C6F4D28460B324FFF04FD36DED3D3C4" authority="Arrow, 1913" authorityName="Arrow" authorityYear="1913" box="[189,487,662,688]" class="Insecta" family="Scarabaeidae" genus="Pectinosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFF04FD36DE61D3DB" box="[189,341,662,687]" italics="true" pageId="22" pageNumber="23">Pectinosoma</emphasis>
<bibRefCitation id="EFFE4B5A460B324FFEE3FD36DED3D3C4" author="Arrow" box="[346,487,662,688]" firstAuthor="Arrow" pageId="22" pageNumber="23" pagination="425 - 432" refId="ref26476" refString="Arrow G. J. 1913. Synopsis of the melolonthid genus Ancistrosoma, with descriptions of new species and an allied new genus. Annals and Magazine of Natural History (8) 12: 425 - 432. https: // doi. org / 10.1080 / 00222931308693420" type="journal article" year="1913">Arrow,1913</bibRefCitation>
</taxonomicName>
and some species of
<taxonomicName id="4C6F4D28460B324FFD58FD36DCB5D3C4" authorityName="Curtis" authorityYear="1835" box="[737,897,662,688]" class="Insecta" family="Scarabaeidae" genus="Ancistrosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFD58FD36DCB5D3C4" box="[737,897,662,688]" italics="true" pageId="22" pageNumber="23">Ancistrosoma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D28460B324FFC29FD36DCC6D3C4" authorityName="Burmeister" authorityYear="1855" box="[912,1010,662,688]" class="Insecta" family="Scarabaeidae" genus="Barybas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFC29FD36DCC6D3C4" box="[912,1010,662,688]" italics="true" pageId="22" pageNumber="23">Barybas</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D28460B324FFBB9FD36DB5CD3C4" authorityName="LePeletier de Saint-Fargeau &amp; Audinet-Serville" authorityYear="1828" box="[1024,1128,662,688]" class="Insecta" family="Scarabaeidae" genus="Ceraspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFBB9FD36DB5CD3C4" box="[1024,1128,662,688]" italics="true" pageId="22" pageNumber="23">Ceraspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D28460B324FFB1BFD36DBDCD3C4" authorityName="Blanchard" authorityYear="1850" box="[1186,1256,662,688]" class="Insecta" family="Scarabaeidae" genus="Faula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB9460B324FFB1BFD36DBDCD3C4" box="[1186,1256,662,688]" italics="true" pageId="22" pageNumber="23">Faula</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>