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<document id="A5EB9FE012C30DEE7186C566B8CF8BF7" ID-CLB-Dataset="22059" ID-DOI="10.5852/ejt.2017.350" ID-GBIF-Dataset="52e4ae01-f51f-4c4a-8a14-9fb5b17788d6" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832682" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831660737" checkinUser="carolina" docAuthor="Fuhrmann, Juares &amp; Vaz-De-Mello, Fernando Z." docDate="2017" docId="03C687BD46073243FDC9FAEFDC9BD69F" docLanguage="en" docName="ejt-2017-350.pdf.imf" docOrigin="European Journal of Taxonomy 350" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chariodactylus Moser 1919" docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FFFFFFC5461D3259FFB9FFA0DF34D174" masterDocTitle="Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification" masterLastPageNumber="71" masterPageNumber="1" pageNumber="27" updateTime="1698840167045" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4133C1EFB1ED2C28AA2DC28D77778D0C">Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification</mods:title>
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<mods:namePart id="950F440CA6FA4C74D1A05E02FFCB1685">Fuhrmann, Juares</mods:namePart>
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<mods:namePart id="6BB9A50ED56900B14F92B639581FD3B2">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="18B707D18B2B881AE5E4082B6ABEF44A">European Journal of Taxonomy</mods:title>
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<mods:date id="A2EB7DBDAB8211DC3781B7DE9B44106D">2017</mods:date>
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<treatment id="03C687BD46073243FDC9FAEFDC9BD69F" ID-DOI="http://doi.org/10.5281/zenodo.3851128" ID-GBIF-Taxon="164167397" ID-Zenodo-Dep="3851128" LSID="urn:lsid:plazi:treatment:03C687BD46073243FDC9FAEFDC9BD69F" httpUri="http://treatment.plazi.org/id/03C687BD46073243FDC9FAEFDC9BD69F" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection id="C375652046073243FDC9FAEFDCF7D41E" box="[624,963,1359,1386]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BD036AB46073243FDC9FAEFDCF7D41E" blockId="26.[189,963,1359,1422]" box="[624,963,1359,1386]" pageId="26" pageNumber="27">
<heading id="D09881C746073243FDC9FAEFDCF7D41E" box="[624,963,1359,1386]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="4C6F4D2846073243FDC9FAEFDCF7D41E" ID-CoL="3M9R" authority="Moser, 1919" authorityName="Moser" authorityYear="1919" box="[624,963,1359,1386]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FDC9FAEFDC1CD41E" bold="true" box="[624,808,1359,1386]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
<bibRefCitation id="EFFE4B5A46073243FC89FAF0DCF7D41E" author="Moser" box="[816,963,1360,1386]" firstAuthor="Moser" pageId="26" pageNumber="27" pagination="3 - 64" refId="ref29954" refString="Moser J. 1919 a. Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." type="journal article" year="1919">Moser, 1919</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375652046073243FF04FAD4DB27D4CA" pageId="26" pageNumber="27" type="type_taxon">
<paragraph id="8BD036AB46073243FF04FAD4DE6CD4FA" blockId="26.[189,963,1359,1422]" box="[189,344,1396,1422]" pageId="26" pageNumber="27">
<heading id="D09881C746073243FF04FAD4DE6CD4FA" bold="true" box="[189,344,1396,1422]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEAB946073243FF04FAD4DE6CD4FA" bold="true" box="[189,344,1396,1422]" pageId="26" pageNumber="27">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB46073243FF04FA04DB27D4CA" blockId="26.[189,1043,1444,1470]" box="[189,1043,1444,1470]" pageId="26" pageNumber="27">
<taxonomicName id="4C6F4D2846073243FF04FA04DDA1D4CA" ID-CoL="TNWY" authority="Moser, 1919" authorityName="Moser" authorityYear="1919" box="[189,661,1444,1470]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chacoensis">
<emphasis id="B91BEAB946073243FF04FA04DECED4CA" box="[189,506,1444,1470]" italics="true" pageId="26" pageNumber="27">Chariodactylus chacoensis</emphasis>
<bibRefCitation id="EFFE4B5A46073243FDB8FA04DDA1D4CA" author="Moser" box="[513,661,1444,1470]" firstAuthor="Moser" pageId="26" pageNumber="27" pagination="3 - 64" refId="ref29954" refString="Moser J. 1919 a. Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." type="journal article" year="1919">Moser, 1919</bibRefCitation>
</taxonomicName>
(designation:
<bibRefCitation id="EFFE4B5A46073243FC85FA04DB32D4CA" author="Evans" box="[828,1030,1444,1470]" firstAuthor="Evans" pageId="26" pageNumber="27" pagination="1 - 458" refId="ref28076" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458." type="journal article" year="2003">Evans 2003: 247</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C375652046073243FF04FA46DD2DD7B6" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BD036AB46073243FF04FA46DE02D774" blockId="26.[189,310,1510,1536]" box="[189,310,1510,1536]" pageId="26" pageNumber="27">
<heading id="D09881C746073243FF04FA46DE02D774" bold="true" box="[189,310,1510,1536]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEAB946073243FF04FA46DE02D774" bold="true" box="[189,310,1510,1536]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB46073243FF04F9B6DD2DD7B6" blockId="26.[189,1399,1558,1731]" pageId="26" pageNumber="27">
Clypeus and frons gradually raised over the lateral area of head (
<figureCitation id="13542A2E46073243FC78F9B6DB1AD745" box="[961,1070,1558,1585]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11A</figureCitation>
); clypeal anteroventral area strongly reduced (similar to
<figureCitation id="13542A2E46073243FDBEF99BDD46D721" box="[519,626,1595,1621]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11H</figureCitation>
); posterior ocular area exposed (
<figureCitation id="13542A2E46073243FC54F99BDB6CD721" box="[1005,1112,1595,1621]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11A</figureCitation>
), eye large; pronotum as long as wide, with anterior and posterior margins not beaded; prosternum with two anterior sulci (similar to
<figureCitation id="13542A2E46073243FF63F924DE70D7EA" box="[218,324,1668,1694]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11H</figureCitation>
); protibia with two external teeth and without spur (
<figureCitation id="13542A2E46073243FC27F924DB3DD7EA" box="[926,1033,1668,1694]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11A</figureCitation>
); abdomen with intersegmental membrane VII–VIII exposed.
</paragraph>
</subSubSection>
<subSubSection id="C375652046073243FF04F94BDC9BD69F" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BD036AB46073243FF04F94BDE1BD671" blockId="26.[189,303,1771,1797]" box="[189,303,1771,1797]" pageId="26" pageNumber="27">
<heading id="D09881C746073243FF04F94BDE1BD671" bold="true" box="[189,303,1771,1797]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEAB946073243FF04F94BDE1BD671" bold="true" box="[189,303,1771,1797]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036AB46073243FF04F8BADC9BD69F" blockId="26.[189,1399,1818,2027]" pageId="26" pageNumber="27">
The genera
<taxonomicName id="4C6F4D2846073243FEF5F8BBDD35D641" authorityName="Moser" authorityYear="1919" box="[332,513,1819,1845]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FEF5F8BBDD35D641" box="[332,513,1819,1845]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D2846073243FDABF8BBDDF5D641" authorityName="Dejean" authorityYear="1821" box="[530,705,1819,1845]" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FDABF8BBDDF5D641" box="[530,705,1819,1845]" italics="true" pageId="26" pageNumber="27">Macrodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D2846073243FD47F8BBDB77D641" authority="Moser, 1919" authorityName="Moser" authorityYear="1919" box="[766,1091,1819,1845]" class="Insecta" family="Scarabaeidae" genus="Manodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FD47F8BBDC90D641" box="[766,932,1819,1845]" italics="true" pageId="26" pageNumber="27">Manodactylus</emphasis>
<bibRefCitation id="EFFE4B5A46073243FC17F8BBDB77D641" author="Moser" box="[942,1091,1819,1845]" firstAuthor="Moser" pageId="26" pageNumber="27" pagination="3 - 64" refId="ref29954" refString="Moser J. 1919 a. Beitrag zur Kenntnis der Melolonthiden (Col.). (IX). Stettiner Entomologische Zeitung 80: 3 - 64." type="journal article" year="1919">Moser, 1919</bibRefCitation>
</taxonomicName>
are similar to each other.
<taxonomicName id="4C6F4D2846073243FF04F89FDE46D62D" authorityName="Moser" authorityYear="1919" box="[189,370,1855,1881]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FF04F89FDE46D62D" box="[189,370,1855,1881]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
</taxonomicName>
(
<figureCitation id="13542A2E46073243FE3DF89FDEC4D62D" box="[388,496,1855,1881]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11A</figureCitation>
) is distinguished as follows (opposition to 1,
<taxonomicName id="4C6F4D2846073243FBA3F89FDBFDD62D" authorityName="Dejean" authorityYear="1821" box="[1050,1225,1855,1881]" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FBA3F89FDBFDD62D" box="[1050,1225,1855,1881]" italics="true" pageId="26" pageNumber="27">Macrodactylus</emphasis>
</taxonomicName>
(
<figureCitation id="13542A2E46073243FB65F89FDA7CD62D" box="[1244,1352,1855,1881]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11C</figureCitation>
); 2,
<taxonomicName id="4C6F4D2846073243FF04F8C3DE57D609" authorityName="Moser" authorityYear="1919" box="[189,355,1891,1917]" class="Insecta" family="Scarabaeidae" genus="Manodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FF04F8C3DE57D609" box="[189,355,1891,1917]" italics="true" pageId="26" pageNumber="27">Manodactylus</emphasis>
</taxonomicName>
(
<figureCitation id="13542A2E46073243FECCF8C3DED4D60A" box="[373,480,1891,1918]" captionStart="Fig" captionStartId="28.[189,232,1890,1916]" captionTargetBox="[191,1398,542,1862]" captionTargetId="figure@28.[189,1398,539,1862]" captionTargetPageId="28" captionText="Fig. 11. ♂♂. A−C. Head–prothorax dorsal, tarsus rotated laterally to apex. D−G. Aedeagus (lateral, parameres apex). H. Head−prothorax, ventral. A, D−E. Chariodactylus chacoensis Moser, 1919. B, F−H. Manodactylus gaujoni Moser, 1919. C. Macrodactylus pumilio Burmeister, 1855. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832704" httpUri="https://zenodo.org/record/3832704/files/figure.png" pageId="26" pageNumber="27">Fig. 11B</figureCitation>
)): clypeus broad, trapezoid (1, narrow, trapezoid; 2, as
<taxonomicName id="4C6F4D2846073243FBC4F8C3DA06D609" authorityName="Moser" authorityYear="1919" box="[1149,1330,1891,1917]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FBC4F8C3DA06D609" box="[1149,1330,1891,1917]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
</taxonomicName>
); eye large (1, 2, small); pronotum wider than long (1, longer than large; 2, as
<taxonomicName id="4C6F4D2846073243FC4EF828DB98D6D6" authorityName="Moser" authorityYear="1919" box="[1015,1196,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FC4EF828DB98D6D6" box="[1015,1196,1928,1954]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
</taxonomicName>
). Future cladistic analysis and a taxonomic review of
<taxonomicName id="4C6F4D2846073243FDC0F80CDC1CD6B2" authorityName="Dejean" authorityYear="1821" box="[633,808,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Macrodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FDC0F80CDC1CD6B2" box="[633,808,1964,1990]" italics="true" pageId="26" pageNumber="27">Macrodactylus</emphasis>
</taxonomicName>
including
<taxonomicName id="4C6F4D2846073243FC15F80CDB55D6B2" authorityName="Moser" authorityYear="1919" box="[940,1121,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Chariodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FC15F80CDB55D6B2" box="[940,1121,1964,1990]" italics="true" pageId="26" pageNumber="27">Chariodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D2846073243FB18F80CDA73D6B2" authorityName="Moser" authorityYear="1919" box="[1185,1351,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Manodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAB946073243FB18F80CDA73D6B2" box="[1185,1351,1964,1990]" italics="true" pageId="26" pageNumber="27">Manodactylus</emphasis>
</taxonomicName>
are needed to confirm the possible synonymy between these genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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