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<document id="032B45546A2D3F3061F7A77D21396E27" ID-CLB-Dataset="20153" ID-DOI="10.11646/zootaxa.5116.4.3" ID-GBIF-Dataset="599081e4-2417-4e10-aa2e-bb7f8acb83a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="6375321" ID-ZooBank="DD221377-2CA5-479C-85C5-99640EA5E287" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647940542378" checkinUser="plazi" docAuthor="Huang, Hao &amp; Chen, Chang-Chin" docDate="2022" docId="03C687BAFFF8FFAA8581BA186D0555FE" docLanguage="en" docName="zootaxa.5116.4.3.pdf" docOrigin="Zootaxa 5116 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xizangia qiuae Huang &amp; Chen 2022, new species" docType="treatment" docVersion="17" lastPageNumber="538" masterDocId="FFFFFFC2FFFAFFBF8516BC536E12567F" masterDocTitle="Rediscovery of the myrmecophilous Lucanid genus Xizangia Zhang, 1988 from southwest China (Coleoptera: Scarabaeoidea: Lucanidae)" masterLastPageNumber="549" masterPageNumber="517" pageNumber="519" updateTime="1699318422324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F10C4B1FBBD895A3F09F03667D0404F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC48A9BFDBE71FCB20D3D86953949CAC">
<mods:title id="AA1510919BC68FCA972163F764A8C18B">Rediscovery of the myrmecophilous Lucanid genus Xizangia Zhang, 1988 from southwest China (Coleoptera: Scarabaeoidea: Lucanidae)</mods:title>
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<mods:name id="D5C3160361A0A00EF75351878591E94B" type="personal">
<mods:role id="DDFD0811D6B0FCAF4DC42B857CD68E94">
<mods:roleTerm id="2D3E9B4673AD6829620291662E259395">Author</mods:roleTerm>
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<mods:namePart id="3233EAD512D52093E187F51AE552D963">Huang, Hao</mods:namePart>
<mods:affiliation id="B302359D4F7745CE8BCBEF56F34E292B">503, Unit 1, # 1 Dongtinghu Road, Qingdao, P. R. China.</mods:affiliation>
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<mods:roleTerm id="6F4F84ECA362C797920324F7B50F5ADE">Author</mods:roleTerm>
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<mods:namePart id="AD29EA7C15305DA30B8F4F432FC58FFF">Chen, Chang-Chin</mods:namePart>
<mods:affiliation id="622FBBDB597750B430EC2302E269380C">Technical Center, Tianjin New Wei San Industrial Company, Limited, P. R. China.</mods:affiliation>
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<mods:typeOfResource id="FCF5057762E1D725179763FCBDAB93CE">text</mods:typeOfResource>
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<mods:title id="EEA9063F6B02EE06AB14689331569312">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4FAED4DFC39DB5EE7FF21776084E1175">
<mods:date id="7FDD4BC1E2A17BFDF349B056374719B9">2022</mods:date>
<mods:detail id="D1983EB72A570D66CF3020D897B39005" type="pubDate">
<mods:number id="FC09FCC26F41B4503B0CF8E7C7AF984B">2022-03-22</mods:number>
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<mods:number id="668E236B4E634821A07FF6A16D9ECA06">5116</mods:number>
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<mods:number id="FBEDA6D5E3CA49E7A558F02D43075E77">4</mods:number>
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<mods:classification id="AF4421677C7DA23B56FD9FFA035A52F7">journal article</mods:classification>
<mods:identifier id="73FC474532C456572444C96E9A5DE118" type="CLB-Dataset">20153</mods:identifier>
<mods:identifier id="3BA8D580A21EEEC4CBB62189E5214173" type="DOI">10.11646/zootaxa.5116.4.3</mods:identifier>
<mods:identifier id="32A530574072FA5E18D94F18B8ABC4F3" type="GBIF-Dataset">599081e4-2417-4e10-aa2e-bb7f8acb83a4</mods:identifier>
<mods:identifier id="BED35A41C4D459C4C42966424F782A59" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C687BAFFF8FFAA8581BA186D0555FE" ID-DOI="http://doi.org/10.5281/zenodo.7509778" ID-Zenodo-Dep="7509778" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE" httpUri="http://treatment.plazi.org/id/03C687BAFFF8FFAA8581BA186D0555FE" lastPageId="21" lastPageNumber="538" pageId="2" pageNumber="519">
<subSubSection id="C3756527FFF8FFBD8581BA186CA3501B" box="[151,689,1610,1637]" pageId="2" pageNumber="519" type="nomenclature">
<paragraph id="8BD036ACFFF8FFBD8581BA186CA3501B" blockId="2.[151,689,1610,1673]" box="[151,689,1610,1637]" pageId="2" pageNumber="519">
<heading id="D09881C0FFF8FFBD8581BA186CA3501B" bold="true" box="[151,689,1610,1637]" fontSize="11" level="1" pageId="2" pageNumber="519" reason="1">
<emphasis id="B91BEABEFFF8FFBD8581BA186CA3501B" bold="true" box="[151,689,1610,1637]" pageId="2" pageNumber="519">
<taxonomicName id="4C6F4D2FFFF8FFBD8581BA186C06501B" ID-CoL="8W9HR" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[151,532,1610,1637]" class="Insecta" family="Lucanidae" genus="Xizangia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF8FFBD8581BA186F5F501A" bold="true" box="[151,333,1611,1637]" italics="true" pageId="2" pageNumber="519">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF8FFBD8734BA186CA3501B" box="[546,689,1610,1637]" pageId="2" pageNumber="519" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF8FFBD8581BA3C6C7050F6" box="[151,610,1646,1673]" pageId="2" pageNumber="519" type="description">
<paragraph id="8BD036ACFFF8FFBD8581BA3C6C7050F6" blockId="2.[151,689,1610,1673]" box="[151,610,1646,1673]" pageId="2" pageNumber="519">
(
<figureCitation id="13542A29FFF8FFBD8589BA3C6F1950F6" box="[159,267,1647,1673]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–8" captionStartId-0="3.[151,264,1965,1989]" captionStartId-1="4.[151,264,1943,1967]" captionTargetBox-0="[193,1390,214,1907]" captionTargetBox-1="[249,1364,212,1873]" captionTargetId-0="figure-19@3.[151,1435,181,1942]" captionTargetId-1="figure-21@4.[202,1386,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–2. Xizangia qiuae Huang &amp; Chen, new species. 1, habitus in dorsal view; 2, habitus in ventral view (F4—female paratype from Mangdong)." captionText-1="FIGURES 3–8. Xizangia qiuae Huang &amp; Chen, new species, habitus and details (F4). 3, entire body in lateral view; 4, top of last antennomere; 5, part of head in ventral view; 6, left antenna in ventral view; 7, intercoxal process of prosternum; 8, part of head in dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.6375325" figureDoi-1="http://doi.org/10.5281/zenodo.6375327" httpUri-0="https://zenodo.org/record/6375325/files/figure.png" httpUri-1="https://zenodo.org/record/6375327/files/figure.png" pageId="2" pageNumber="519">Figs. 1–8</figureCitation>
,
<figureCitation id="13542A29FFF8FFBD8401BA3D6F4D50F6" box="[279,351,1646,1673]" captionStart="FIGURES 15–19" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[204,1436,181,1896]" captionTargetId="figure-21@6.[266,1321,181,1885]" captionTargetPageId="6" captionText="FIGURES 15–19. Xizangia qiuae Huang &amp; Chen, new species, habitus in dorsal view (M1—male holotype, F1—female paratype, F2—female paratype, F3—female paratype and F4 —female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6375331" httpUri="https://zenodo.org/record/6375331/files/figure.png" pageId="2" pageNumber="519">15–19</figureCitation>
,
<figureCitation id="13542A29FFF8FFBD847DBA3C6FA650F6" box="[363,436,1647,1673]" captionStart-0="FIGURE 21" captionStart-1="FIGURES 22–26" captionStartId-0="6.[151,250,2017,2041]" captionStartId-1="7.[151,264,1874,1898]" captionTargetBox-0="[204,1436,181,1896]" captionTargetBox-1="[151,1280,181,1866]" captionTargetId-0="figure-21@6.[266,1321,181,1885]" captionTargetId-1="figure-19@7.[151,1164,181,1851]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 21. Xizangia qiuae Huang &amp; Chen, new species, legs (F2)." captionText-1="FIGURES 22–26. Xizangia qiuae Huang &amp; Chen, new species, habitus in ventral view (M1, F1, F2, F3 and F4)." figureDoi-0="http://doi.org/10.5281/zenodo.7510338" figureDoi-1="http://doi.org/10.5281/zenodo.6375333" httpUri-0="https://zenodo.org/record/7510338/files/figure.png" httpUri-1="https://zenodo.org/record/6375333/files/figure.png" pageId="2" pageNumber="519">21–26</figureCitation>
,
<figureCitation id="13542A29FFF8FFBD84A9BA3C6C1550F6" box="[447,519,1647,1673]" captionStart-0="FIGURE 28" captionStart-1="FIGURES 29–31" captionStartId-0="7.[151,250,1946,1970]" captionStartId-1="7.[151,264,1982,2006]" captionTargetBox-0="[151,1280,181,1866]" captionTargetBox-1="[151,1280,181,1866]" captionTargetId-0="figure-19@7.[151,1164,181,1851]" captionTargetId-1="figure-19@7.[151,1164,181,1851]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURE 28. Xizangia qiuae Huang &amp; Chen, new species, metatarsi in ventral view (F1)." captionText-1="FIGURES 29–31. Xizangia qiuae Huang &amp; Chen, new species, mandibles in ventral view (F3, F2 and F1)" figureDoi-0="http://doi.org/10.5281/zenodo.7510344" figureDoi-1="http://doi.org/10.5281/zenodo.7510346" httpUri-0="https://zenodo.org/record/7510344/files/figure.png" httpUri-1="https://zenodo.org/record/7510346/files/figure.png" pageId="2" pageNumber="519">28–31</figureCitation>
,
<figureCitation id="13542A29FFF8FFBD8705BA3D6C4850F6" box="[531,602,1646,1673]" captionStart-0="FIGURE 33" captionStart-1="FIGURE 34" captionStart-2="FIGURES 35–39" captionStart-3="FIGURE 40" captionStart-4="FIGURES 41–42" captionStart-5="FIGURE 43" captionStart-6="FIGURES 44–48" captionStart-7="FIGURES 49–52" captionStart-8="FIGURES 53–57" captionStart-9="FIGURE 58" captionStartId-0="8.[151,250,1943,1967]" captionStartId-1="8.[151,250,1979,2003]" captionStartId-2="9.[151,264,1943,1967]" captionStartId-3="13.[151,250,1944,1968]" captionStartId-4="13.[151,264,1980,2004]" captionStartId-5="14.[151,250,1992,2016]" captionStartId-6="15.[151,264,1943,1967]" captionStartId-7="16.[151,264,1943,1967]" captionStartId-8="17.[151,264,1943,1967]" captionStartId-9="18.[151,250,1895,1919]" captionTargetBox-0="[174,1436,181,1932]" captionTargetBox-1="[174,1436,181,1932]" captionTargetBox-2="[230,1358,181,1920]" captionTargetBox-3="[151,1436,181,1948]" captionTargetBox-4="[151,1436,181,1948]" captionTargetBox-5="[151,1435,181,1969]" captionTargetBox-6="[194,1393,181,1920]" captionTargetBox-7="[203,1385,181,1920]" captionTargetBox-8="[194,1393,181,1920]" captionTargetBox-9="[151,1435,181,1871]" captionTargetId-0="figure-21@8.[230,1358,181,1920]" captionTargetId-1="figure-21@8.[230,1358,181,1920]" captionTargetId-2="figure-19@9.[230,1358,181,1920]" captionTargetId-3="figure-19@13.[151,1435,181,1920]" captionTargetId-4="figure-19@13.[151,1435,181,1920]" captionTargetId-5="figure-21@14.[151,1435,181,1969]" captionTargetId-6="figure-19@15.[194,1393,181,1920]" captionTargetId-7="figure-21@16.[203,1385,181,1920]" captionTargetId-8="figure-19@17.[194,1393,181,1920]" captionTargetId-9="figure-21@18.[151,1435,181,1871]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="13" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionTargetPageId-7="16" captionTargetPageId-8="17" captionTargetPageId-9="18" captionText-0="FIGURE 33. Diagnostic characters of Xizangia species: legs." captionText-1="FIGURE 34. Xizangia qiuae Huang &amp; Chen, new species, mandibles in dorsal, inner lateral and ventral views: left half — male mandibles; right half — female mandibles." captionText-2="FIGURES 35–39. Xizangia qiuae Huang &amp; Chen, new species, characters. 35, hindwing; 36, female antenna in various views; 37, maxilla and labium; 38, clypeus, labrum and mentum in anterior view; 39, clypeus, labrum and mentum in ventral view." captionText-3="FIGURE 40. Xizangia qiuae Huang &amp; Chen, new species, tarsi and claws." captionText-4="FIGURES 41–42. Xizangia qiuae Huang &amp; Chen, new species, female genitalia." captionText-5="FIGURE 43. Xizangia qiuae Huang &amp; Chen, new species, male genitalia (M1—holotype)." captionText-6="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." captionText-7="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." captionText-8="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." captionText-9="FIGURE 58. Xizangia qiuae Huang &amp; Chen, new species, pupa in dorsal, lateral and ventral views." figureDoi-0="http://doi.org/10.5281/zenodo.6375335" figureDoi-1="http://doi.org/10.5281/zenodo.7510494" figureDoi-2="http://doi.org/10.5281/zenodo.6375337" figureDoi-3="http://doi.org/10.5281/zenodo.6375339" figureDoi-4="http://doi.org/10.5281/zenodo.6377198" figureDoi-5="http://doi.org/10.5281/zenodo.6375343" figureDoi-6="http://doi.org/10.5281/zenodo.6375345" figureDoi-7="http://doi.org/10.5281/zenodo.6375349" figureDoi-8="http://doi.org/10.5281/zenodo.6375351" figureDoi-9="http://doi.org/10.5281/zenodo.6375353" httpUri-0="https://zenodo.org/record/6375335/files/figure.png" httpUri-1="https://zenodo.org/record/7510494/files/figure.png" httpUri-2="https://zenodo.org/record/6375337/files/figure.png" httpUri-3="https://zenodo.org/record/6375339/files/figure.png" httpUri-4="https://zenodo.org/record/6377198/files/figure.png" httpUri-5="https://zenodo.org/record/6375343/files/figure.png" httpUri-6="https://zenodo.org/record/6375345/files/figure.png" httpUri-7="https://zenodo.org/record/6375349/files/figure.png" httpUri-8="https://zenodo.org/record/6375351/files/figure.png" httpUri-9="https://zenodo.org/record/6375353/files/figure.png" pageId="2" pageNumber="519">33–58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF8FFBD8581BAE46B1951B1" pageId="2" pageNumber="519" type="materials_examined">
<paragraph id="8BD036ACFFF8FFBD8581BAE46A9D50AE" blockId="2.[151,1437,1718,1998]" box="[151,1167,1718,1745]" pageId="2" pageNumber="519">
<emphasis id="B91BEABEFFF8FFBD8581BAE46F2450AF" bold="true" box="[151,310,1718,1745]" pageId="2" pageNumber="519">
<typeStatus id="54D4880EFFF8FFBD8581BAE46EC050AE" box="[151,210,1719,1745]" pageId="2" pageNumber="519">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F378763CFFF8FFBD842ABAE46F9050AE" box="[316,386,1719,1745]" name="China" pageId="2" pageNumber="519">China</collectingCountry>
,
<collectingRegion id="49ABF84EFFF8FFBD849ABAE46C5D50AE" box="[396,591,1719,1745]" country="China" name="Yunnan" pageId="2" pageNumber="519">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BB4ACD6FFF8FFBD874CBAE46D9750AE" box="[602,901,1719,1745]" pageId="2" pageNumber="519">Xishuangbanna Prefecture</collectingMunicipality>
,
<collectingCounty id="62B14E20FFF8FFBD8686BAE46A5350AE" box="[912,1089,1719,1745]" pageId="2" pageNumber="519">Mengla County</collectingCounty>
,
<location id="8EB06077FFF8FFBD815DBAE46A9950AE" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE:8EB06077FFF8FFBD815DBAE46A9950AE" box="[1099,1163,1719,1745]" country="China" county="Mengla County" municipality="Xishuangbanna Prefecture" name="Yiwu" pageId="2" pageNumber="519" stateProvince="Yunnan">Yiwu</location>
.
</paragraph>
<paragraph id="8BD036ACFFF8FFBD85D1BA886B1951B1" blockId="2.[151,1437,1718,1998]" pageId="2" pageNumber="519">
<emphasis id="B91BEABEFFF8FFBD85D1BA886F68508B" bold="true" box="[199,378,1754,1781]" pageId="2" pageNumber="519">Type material.</emphasis>
<materialsCitation id="3B073CF1FFF8FFBD8493BA886B0A5141" ID-GBIF-Occurrence="3712351304" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Yiwu" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B91BEABEFFF8FFBD8493BA886FE0508A" bold="true" box="[389,498,1755,1781]" pageId="2" pageNumber="519">
<typeStatus id="54D4880EFFF8FFBD8493BA886FE0508A" box="[389,498,1755,1781]" pageId="2" pageNumber="519" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13542A29FFF8FFBD8710BA886C7A508A" box="[518,616,1754,1781]" captionStart="FIGURES 15–19" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[204,1436,181,1896]" captionTargetId="figure-21@6.[266,1321,181,1885]" captionTargetPageId="6" captionText="FIGURES 15–19. Xizangia qiuae Huang &amp; Chen, new species, habitus in dorsal view (M1—male holotype, F1—female paratype, F2—female paratype, F3—female paratype and F4 —female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6375331" httpUri="https://zenodo.org/record/6375331/files/figure.png" pageId="2" pageNumber="519">Figs. 15</figureCitation>
,
<figureCitation id="13542A29FFF8FFBD876FBA886C8B508A" box="[633,665,1755,1781]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[151,1280,181,1866]" captionTargetId="figure-19@7.[151,1164,181,1851]" captionTargetPageId="7" captionText="FIGURES 22–26. Xizangia qiuae Huang &amp; Chen, new species, habitus in ventral view (M1, F1, F2, F3 and F4)." figureDoi="http://doi.org/10.5281/zenodo.6375333" httpUri="https://zenodo.org/record/6375333/files/figure.png" pageId="2" pageNumber="519">22</figureCitation>
):
<emphasis id="B91BEABEFFF8FFBD87A4BA896DC7508A" bold="true" box="[690,981,1754,1781]" pageId="2" pageNumber="519">
<collectingCountry id="F378763CFFF8FFBD87A4BA896D06508B" box="[690,788,1754,1780]" name="China" pageId="2" pageNumber="519">CHINA</collectingCountry>
:
<collectingRegion id="49ABF84EFFF8FFBD8635BA886DC2508A" box="[803,976,1755,1781]" country="China" name="Yunnan" pageId="2" pageNumber="519">Yunnan Prov.</collectingRegion>
:
</emphasis>
<specimenCount id="9D69FD25FFF8FFBD86F7BA886DEA508A" box="[993,1016,1755,1781]" pageId="2" pageNumber="519" type="male">♂</specimenCount>
,
<collectingMunicipality id="6BB4ACD6FFF8FFBD811EBA886AA4508A" box="[1032,1206,1755,1781]" pageId="2" pageNumber="519">Xishuangbanna</collectingMunicipality>
prefecture,
<collectingCounty id="62B14E20FFF8FFBD8051BA886EF95166" pageId="2" pageNumber="519">Mengla County</collectingCounty>
,
<location id="8EB06077FFF8FFBD85EEBAAC6F2A5166" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE:8EB06077FFF8FFBD85EEBAAC6F2A5166" box="[248,312,1791,1817]" country="China" county="Mengla County" municipality="Xishuangbanna" name="Yiwu" pageId="2" pageNumber="519" stateProvince="Yunnan">Yiwu</location>
,
<quantity id="4C979B49FFF8FFBD8453BAAC6F875166" box="[325,405,1791,1817]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="519" unit="m" value="1600.0">
<elevation id="0042D19FFFF8FFBD8453BAAC6F875166" box="[325,405,1791,1817]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="519" unit="m" value="1600.0">1600m</elevation>
</quantity>
, third instar larva collected in an ant nest on
<date id="FFD1106CFFF8FFBD86B1BAAC6A1C5166" box="[935,1038,1791,1817]" pageId="2" pageNumber="519" value="2017-02-01">
<collectingDate id="EF95E984FFF8FFBD86B1BAAC6A1C5166" box="[935,1038,1791,1817]" pageId="2" pageNumber="519" value="2017-02-01">1.II.2017</collectingDate>
</date>
by
<location id="8EB06077FFF8FFBD8128BAAC6AC15166" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE:8EB06077FFF8FFBD8128BAAC6AC15166" box="[1086,1235,1791,1817]" country="China" county="Mengla County" municipality="Xishuangbanna" name="Jian-Yue Qiu" pageId="2" pageNumber="519" stateProvince="Yunnan">
<collectorName id="269A537AFFF8FFBD8128BAAC6AC15166" box="[1086,1235,1791,1817]" pageId="2" pageNumber="519">Jian-Yue Qiu</collectorName>
</location>
and
<location id="8EB06077FFF8FFBD8019BAAC6B785166" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE:8EB06077FFF8FFBD8019BAAC6B785166" box="[1295,1386,1791,1817]" country="China" county="Mengla County" municipality="Xishuangbanna" name="Hao Xu" pageId="2" pageNumber="519" stateProvince="Yunnan">
<collectorName id="269A537AFFF8FFBD8019BAAC6B785166" box="[1295,1386,1791,1817]" pageId="2" pageNumber="519">Hao Xu</collectorName>
</location>
and emerged in the laboratory on
<date id="FFD1106CFFF8FFBD84FCBB706CD35142" box="[490,705,1827,1853]" pageId="2" pageNumber="519" value="2017-04-30" valueMax="2017-05-28" valueMin="2017-04-30">30.IV. –28.V. 2017</date>
(Mianyang Normal University, Mianyang, Sichuan)
</materialsCitation>
.
<materialsCitation id="3B073CF1FFF8FFBD8032BB706AFA511E" ID-GBIF-Occurrence="4011264304" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Yunnan Prov." municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="16" specimenCount-female="10" specimenCount-male="6" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B91BEABEFFF8FFBD8032BB706B8E5142" bold="true" box="[1316,1436,1827,1853]" pageId="2" pageNumber="519">
<typeStatus id="54D4880EFFF8FFBD8032BB706B8E5142" box="[1316,1436,1827,1853]" pageId="2" pageNumber="519" type="paratype">Paratypes</typeStatus>
</emphasis>
(9 ♂♂ &amp; 14 ♀♀ in total):
<emphasis id="B91BEABEFFF8FFBD84D6BB156CC2511E" bold="true" box="[448,720,1862,1889]" pageId="2" pageNumber="519">
<collectingCountry id="F378763CFFF8FFBD84D6BB156C30511F" box="[448,546,1862,1888]" name="China" pageId="2" pageNumber="519">CHINA</collectingCountry>
:
<collectingRegion id="49ABF84EFFF8FFBD873BBB146CC2511E" box="[557,720,1863,1889]" country="China" name="Yunnan" pageId="2" pageNumber="519">Yunnan Prov.</collectingRegion>
</emphasis>
:
<specimenCount id="9D69FD25FFF8FFBD87C9BB146D0C511E" box="[735,798,1863,1889]" pageId="2" pageNumber="519" type="male">6 ♂♂</specimenCount>
&amp;
<specimenCount id="9D69FD25FFF8FFBD8654BB146D82511E" box="[834,912,1863,1889]" pageId="2" pageNumber="519" type="female">10 ♀♀</specimenCount>
, same data as for the holotype
</materialsCitation>
;
<materialsCitation id="3B073CF1FFF8FFBD81E5BB146FF951FA" ID-GBIF-Occurrence="4011264307" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D69FD25FFF8FFBD81E5BB146B20511E" box="[1267,1330,1863,1889]" pageId="2" pageNumber="519" type="male">3 ♂♂</specimenCount>
&amp;
<specimenCount id="9D69FD25FFF8FFBD8040BB146B84511E" box="[1366,1430,1863,1889]" pageId="2" pageNumber="519" type="female">3 ♀♀</specimenCount>
, same data as for the holotype,
</materialsCitation>
<materialsCitation id="3B073CF1FFF8FFBD84E5BB386A9F51FA" ID-GBIF-Occurrence="4011264302" box="[499,1165,1899,1925]" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="paratype">
collected as adults (
<specimenCount id="9D69FD25FFF8FFBD87C2BB386D0651FA" box="[724,788,1899,1925]" pageId="2" pageNumber="519">3 ♀♀</specimenCount>
in
<location id="8EB06077FFF8FFBD862CBB386A9B51FA" LSID="urn:lsid:plazi:treatment:03C687BAFFF8FFAA8581BA186D0555FE:8EB06077FFF8FFBD862CBB386A9B51FA" box="[826,1161,1899,1925]" country="China" name="Mianyang Normal University" pageId="2" pageNumber="519" stateProvince="Yunnan">Mianyang Normal University</location>
,
</materialsCitation>
<materialsCitation id="3B073CF1FFF8FFBD8183BB386EE951D6" ID-GBIF-Occurrence="4011264305" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D69FD25FFF8FFBD8183BB386AD251FA" box="[1173,1216,1899,1925]" pageId="2" pageNumber="519">1 ♂</specimenCount>
&amp;
<specimenCount id="9D69FD25FFF8FFBD81F0BB386B3451FA" box="[1254,1318,1899,1925]" pageId="2" pageNumber="519">2 ♀♀</specimenCount>
in coll.
<collectorName id="269A537AFFF8FFBD8090BB386EE551D6" pageId="2" pageNumber="519">S. Kakizoe</collectorName>
,
</materialsCitation>
<materialsCitation id="3B073CF1FFF8FFBD8414BBDC6C7C51D6" ID-GBIF-Occurrence="4011264301" box="[258,622,1935,1961]" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D69FD25FFF8FFBD8414BBDC6F5251D6" box="[258,320,1935,1961]" pageId="2" pageNumber="519" type="male">4 ♂♂</specimenCount>
&amp;
<specimenCount id="9D69FD25FFF8FFBD8475BBDC6FB351D6" box="[355,417,1935,1961]" pageId="2" pageNumber="519" type="female">4 ♀♀</specimenCount>
in coll.
<collectorName id="269A537AFFF8FFBD84EDBBDC6C7851D6" box="[507,618,1935,1961]" pageId="2" pageNumber="519">H. Huang</collectorName>
,
</materialsCitation>
<materialsCitation id="3B073CF1FFF8FFBD8763BBDC6DEF51D5" ID-GBIF-Occurrence="4011264306" box="[629,1021,1935,1962]" collectingDate="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D69FD25FFF8FFBD8763BBDC6CA151D6" box="[629,691,1935,1961]" pageId="2" pageNumber="519" type="male">4 ♂♂</specimenCount>
&amp;
<specimenCount id="9D69FD25FFF8FFBD87C0BBDC6D0651D6" box="[726,788,1935,1961]" pageId="2" pageNumber="519" type="female">4 ♀♀</specimenCount>
in coll.
<collectorName id="269A537AFFF8FFBD8678BBDC6DE351D5" box="[878,1009,1935,1962]" pageId="2" pageNumber="519">C.-C. Chen</collectorName>
).
</materialsCitation>
<materialsCitation id="3B073CF1FFF8FFBD8115BBDC6B1951B1" ID-GBIF-Occurrence="4011264303" collectingDate="2017-02-01" collectingDateMax="2017-05-28" collectingDateMin="2017-02-01" country="China" county="Mengla County" elevation="1600" location="Mianyang Normal University" municipality="Xishuangbanna" pageId="2" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D69FD25FFF8FFBD8115BBDC6A3E51D6" box="[1027,1068,1935,1961]" pageId="2" pageNumber="519" type="female">1 ♀</specimenCount>
, Dehong prefecture, Longchuan, Mangdong,
<quantity id="4C979B49FFF8FFBD8409BBE06F7D51B2" box="[287,367,1971,1997]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="2" pageNumber="519" unit="m" value="1770.0">1770m</quantity>
,
<date id="FFD1106CFFF8FFBD846CBBE06FFC51B2" box="[378,494,1971,1997]" pageId="2" pageNumber="519" value="2016-06-02">2.VI.2016</date>
, legit Yu-Tang Wang, using a flight intercept trap (in coll. C.-C. Chen).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF106624FFF9FFBC8581BBFE6FB85195" ID-DOI="http://doi.org/10.5281/zenodo.6375325" ID-Zenodo-Dep="6375325" httpUri="https://zenodo.org/record/6375325/files/figure.png" pageId="3" pageNumber="520" startId="3.[151,264,1965,1989]" targetBox="[193,1390,214,1907]" targetPageId="3">
<paragraph id="8BD036ACFFF9FFBC8581BBFE6FB85195" blockId="3.[151,1436,1965,2026]" pageId="3" pageNumber="520">
<emphasis id="B91BEABEFFF9FFBC8581BBFE6F2951BA" bold="true" box="[151,315,1965,1989]" pageId="3" pageNumber="520">FIGURES 1–2.</emphasis>
<taxonomicName id="4C6F4D2FFFF9FFBC8457BBFD6C6E51B9" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[321,636,1966,1990]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="520" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF9FFBC8457BBFD6FCB51BA" box="[321,473,1966,1989]" italics="true" pageId="3" pageNumber="520">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF9FFBC8790BBFD6D1151B9" box="[646,771,1966,1990]" pageId="3" pageNumber="520" rank="species">new species</taxonomicNameLabel>
. 1, habitus in dorsal view; 2, habitus in ventral view (F4—female paratype from Mangdong).
</paragraph>
</caption>
<caption id="DF106624FFFEFFBB8581BBC46F715187" ID-DOI="http://doi.org/10.5281/zenodo.6375327" ID-Zenodo-Dep="6375327" httpUri="https://zenodo.org/record/6375327/files/figure.png" pageId="4" pageNumber="521" startId="4.[151,264,1943,1967]" targetBox="[249,1364,212,1873]" targetPageId="4">
<paragraph id="8BD036ACFFFEFFBB8581BBC46F715187" blockId="4.[151,1436,1943,2040]" pageId="4" pageNumber="521">
<emphasis id="B91BEABEFFFEFFBB8581BBC46F2951D0" bold="true" box="[151,315,1943,1967]" pageId="4" pageNumber="521">FIGURES 3–8.</emphasis>
<taxonomicName id="4C6F4D2FFFFEFFBB8454BBCB6C9251CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[322,640,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="521" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFEFFBB8454BBCB6FC951D0" box="[322,475,1944,1967]" italics="true" pageId="4" pageNumber="521">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFEFFBB879DBBCB6D1A51CF" box="[651,776,1944,1968]" pageId="4" pageNumber="521" rank="species">new species</taxonomicNameLabel>
, habitus and details (F4). 3, entire body in lateral view; 4, top of last antennomere; 5, part of head in ventral view; 6, left antenna in ventral view; 7, intercoxal process of prosternum; 8, part of head in dorsal view.
</paragraph>
</caption>
<caption id="DF106624FFFFFFBA8581BBC46F005187" ID-DOI="http://doi.org/10.5281/zenodo.6375329" ID-Zenodo-Dep="6375329" httpUri="https://zenodo.org/record/6375329/files/figure.png" pageId="5" pageNumber="522" startId="5.[151,264,1943,1967]" targetBox="[202,1386,181,1891]" targetPageId="5">
<paragraph id="8BD036ACFFFFFFBA8581BBC46F005187" blockId="5.[151,1437,1943,2040]" pageId="5" pageNumber="522">
<emphasis id="B91BEABEFFFFFFBA8581BBC46F5551D0" bold="true" box="[151,327,1943,1967]" pageId="5" pageNumber="522">FIGURES 9–14.</emphasis>
<taxonomicName id="4C6F4D2FFFFFFFBA8458BBCB6CAC51CF" authority="Zhang, 1988" authorityName="Zhang" authorityYear="1988" box="[334,702,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="522" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFFFFFBA8458BBCB6C2751CF" box="[334,565,1944,1968]" italics="true" pageId="5" pageNumber="522">Xizangia cryptonychus</emphasis>
<bibRefCitation id="EFFE4B5DFFFFFFBA872ABBCB6CAC51CF" author="Zhang, Y. - W." box="[572,702,1944,1968]" pageId="5" pageNumber="522" pagination="233 - 237" refId="ref10547" refString="Zhang, Y. - W. (1988) Coleoptera: Trogidae. In: F. - S. Huang, F. - S. (Ed.), Insects of Mt. Namjagbarwa Region of Xizang. Science Press, Beijing, pp. 233 - 237. [in Chinese]" type="book chapter" year="1988">Zhang, 1988</bibRefCitation>
</taxonomicName>
, habitus and details (female paratype). 9, entire body in lateral view; 10, top of last antennomere; 11, part of head in ventral view; 12, antenna; 13, intercoxal process of prosternum; 14, part of head in dorsal view.
</paragraph>
</caption>
<caption id="DF106624FFFCFFB98581BB266DC151CE" ID-DOI="http://doi.org/10.5281/zenodo.6375331" ID-Zenodo-Dep="6375331" httpUri="https://zenodo.org/record/6375331/files/figure.png" pageId="6" pageNumber="523" startId="6.[151,264,1909,1933]" targetBox="[204,1436,181,1896]" targetPageId="6">
<paragraph id="8BD036ACFFFCFFB98581BB266DC151CE" blockId="6.[151,1436,1909,1969]" pageId="6" pageNumber="523">
<emphasis id="B91BEABEFFFCFFB98581BB266F4A51F2" bold="true" box="[151,344,1909,1933]" pageId="6" pageNumber="523">FIGURES 15–19.</emphasis>
<taxonomicName id="4C6F4D2FFFFCFFB98477BB256CB951F2" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[353,683,1909,1933]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="523" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFCFFB98477BB256FEF51F2" box="[353,509,1910,1933]" italics="true" pageId="6" pageNumber="523">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFCFFB987AFBB266D2B51F2" box="[697,825,1909,1933]" pageId="6" pageNumber="523" rank="species">new species</taxonomicNameLabel>
, habitus in dorsal view (M1—male holotype, F1—female paratype, F2—female paratype, F3—female paratype and F4 —female paratype).
</paragraph>
</caption>
<caption id="DF106624FFFCFFB98581BBEE6A5351AA" ID-DOI="http://doi.org/10.5281/zenodo.6379417" ID-Zenodo-Dep="6379417" box="[151,1089,1981,2005]" httpUri="https://zenodo.org/record/6379417/files/figure.png" pageId="6" pageNumber="523" startId="6.[151,250,1981,2005]" targetBox="[204,1436,181,1896]" targetPageId="6">
<paragraph id="8BD036ACFFFCFFB98581BBEE6A5351AA" blockId="6.[151,1089,1981,2005]" box="[151,1089,1981,2005]" pageId="6" pageNumber="523">
<emphasis id="B91BEABEFFFCFFB98581BBEE6F3251AA" bold="true" box="[151,288,1981,2005]" pageId="6" pageNumber="523">FIGURE 20.</emphasis>
<taxonomicName id="4C6F4D2FFFFCFFB98430BBED6C8451AA" authority="Zhang, 1988" authorityName="Zhang" authorityYear="1988" box="[294,662,1981,2005]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="523" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFFCFFB98430BBED6C1F51AA" box="[294,525,1981,2005]" italics="true" pageId="6" pageNumber="523">Xizangia cryptonychus</emphasis>
<bibRefCitation id="EFFE4B5DFFFCFFB98702BBEE6C8451AA" author="Zhang, Y. - W." box="[532,662,1981,2005]" pageId="6" pageNumber="523" pagination="233 - 237" refId="ref10547" refString="Zhang, Y. - W. (1988) Coleoptera: Trogidae. In: F. - S. Huang, F. - S. (Ed.), Insects of Mt. Namjagbarwa Region of Xizang. Science Press, Beijing, pp. 233 - 237. [in Chinese]" type="book chapter" year="1988">Zhang, 1988</bibRefCitation>
</taxonomicName>
, habitus in dorsal view (female
<typeStatus id="54D4880EFFFCFFB986CCBBEE6A2451AA" box="[986,1078,1981,2005]" pageId="6" pageNumber="523" type="paratype">paratype</typeStatus>
).
</paragraph>
</caption>
<caption id="DF106624FFFCFFB98581BBB26D415186" ID-DOI="http://doi.org/10.5281/zenodo.7510338" ID-Zenodo-Dep="7510338" box="[151,851,2017,2041]" httpUri="https://zenodo.org/record/7510338/files/figure.png" pageId="6" pageNumber="523" startId="6.[151,250,2017,2041]" targetBox="[204,1436,181,1896]" targetPageId="6">
<paragraph id="8BD036ACFFFCFFB98581BBB26D415186" blockId="6.[151,851,2017,2041]" box="[151,851,2017,2041]" pageId="6" pageNumber="523">
<emphasis id="B91BEABEFFFCFFB98581BBB26F325186" bold="true" box="[151,288,2017,2041]" pageId="6" pageNumber="523">FIGURE 21.</emphasis>
<taxonomicName id="4C6F4D2FFFFCFFB98430BBB16C735186" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[294,609,2017,2041]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="523" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFCFFB98430BBB16FAC5186" box="[294,446,2018,2041]" italics="true" pageId="6" pageNumber="523">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFCFFB9877DBBB26CFA5186" box="[619,744,2017,2041]" pageId="6" pageNumber="523" rank="species">new species</taxonomicNameLabel>
, legs (F2).
</paragraph>
</caption>
<caption id="DF106624FFFDFFB88581BB016B035114" ID-DOI="http://doi.org/10.5281/zenodo.6375333" ID-Zenodo-Dep="6375333" box="[151,1297,1874,1899]" httpUri="https://zenodo.org/record/6375333/files/figure.png" pageId="7" pageNumber="524" startId="7.[151,264,1874,1898]" targetBox="[151,1280,181,1866]" targetPageId="7">
<paragraph id="8BD036ACFFFDFFB88581BB016B035114" blockId="7.[151,1297,1874,1899]" box="[151,1297,1874,1899]" pageId="7" pageNumber="524">
<emphasis id="B91BEABEFFFDFFB88581BB016F465114" bold="true" box="[151,340,1874,1899]" pageId="7" pageNumber="524">FIGURES 22–26.</emphasis>
<taxonomicName id="4C6F4D2FFFFDFFB8844CBB006C875114" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[346,661,1875,1899]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="524" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFDFFB8844CBB006FE15115" box="[346,499,1875,1898]" italics="true" pageId="7" pageNumber="524">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFDFFB88789BB006D0E5114" box="[671,796,1875,1899]" pageId="7" pageNumber="524" rank="species">new species</taxonomicNameLabel>
, habitus in ventral view (M1, F1, F2, F3 and F4).
</paragraph>
</caption>
<caption id="DF106624FFFDFFB88581BB256A5B51F0" ID-DOI="http://doi.org/10.5281/zenodo.6379421" ID-Zenodo-Dep="6379421" box="[151,1097,1910,1935]" httpUri="https://zenodo.org/record/6379421/files/figure.png" pageId="7" pageNumber="524" startId="7.[151,250,1910,1934]" targetBox="[151,1280,181,1866]" targetPageId="7">
<paragraph id="8BD036ACFFFDFFB88581BB256A5B51F0" blockId="7.[151,1097,1910,1935]" box="[151,1097,1910,1935]" pageId="7" pageNumber="524">
<emphasis id="B91BEABEFFFDFFB88581BB256F3251F0" bold="true" box="[151,288,1910,1935]" pageId="7" pageNumber="524">FIGURE 27.</emphasis>
<taxonomicName id="4C6F4D2FFFFDFFB88430BB246C8451F0" authority="Zhang, 1988" authorityName="Zhang" authorityYear="1988" box="[294,662,1911,1935]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="524" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFFDFFB88430BB246C1F51F0" box="[294,525,1911,1935]" italics="true" pageId="7" pageNumber="524">Xizangia cryptonychus</emphasis>
<bibRefCitation id="EFFE4B5DFFFDFFB88702BB246C8451F0" author="Zhang, Y. - W." box="[532,662,1911,1935]" pageId="7" pageNumber="524" pagination="233 - 237" refId="ref10547" refString="Zhang, Y. - W. (1988) Coleoptera: Trogidae. In: F. - S. Huang, F. - S. (Ed.), Insects of Mt. Namjagbarwa Region of Xizang. Science Press, Beijing, pp. 233 - 237. [in Chinese]" type="book chapter" year="1988">Zhang, 1988</bibRefCitation>
</taxonomicName>
, habitus in ventral view (female
<typeStatus id="54D4880EFFFDFFB886F4BB246A2C51F0" box="[994,1086,1911,1935]" pageId="7" pageNumber="524" type="paratype">paratype</typeStatus>
).
</paragraph>
</caption>
<caption id="DF106624FFFDFFB88581BBC96A3751CC" ID-DOI="http://doi.org/10.5281/zenodo.7510344" ID-Zenodo-Dep="7510344" box="[151,1061,1946,1971]" httpUri="https://zenodo.org/record/7510344/files/figure.png" pageId="7" pageNumber="524" startId="7.[151,250,1946,1970]" targetBox="[151,1280,181,1866]" targetPageId="7">
<paragraph id="8BD036ACFFFDFFB88581BBC96A3751CC" blockId="7.[151,1061,1946,1971]" box="[151,1061,1946,1971]" pageId="7" pageNumber="524">
<emphasis id="B91BEABEFFFDFFB88581BBC96F3251CC" bold="true" box="[151,288,1946,1971]" pageId="7" pageNumber="524">FIGURE 28.</emphasis>
<taxonomicName id="4C6F4D2FFFFDFFB88430BBC86C7351CC" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[294,609,1947,1971]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="524" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFDFFB88430BBC86FAC51CD" box="[294,446,1947,1970]" italics="true" pageId="7" pageNumber="524">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFDFFB8877DBBC86CFA51CC" box="[619,744,1947,1971]" pageId="7" pageNumber="524" rank="species">new species</taxonomicNameLabel>
, metatarsi in ventral view (F1).
</paragraph>
</caption>
<caption id="DF106624FFFDFFB88581BBED6AC651A8" ID-DOI="http://doi.org/10.5281/zenodo.7510346" ID-Zenodo-Dep="7510346" box="[151,1236,1982,2007]" httpUri="https://zenodo.org/record/7510346/files/figure.png" pageId="7" pageNumber="524" startId="7.[151,264,1982,2006]" targetBox="[151,1280,181,1866]" targetPageId="7">
<paragraph id="8BD036ACFFFDFFB88581BBED6AC651A8" blockId="7.[151,1236,1982,2007]" box="[151,1236,1982,2007]" pageId="7" pageNumber="524">
<emphasis id="B91BEABEFFFDFFB88581BBED6F4651A8" bold="true" box="[151,340,1982,2007]" pageId="7" pageNumber="524">FIGURES 29–31.</emphasis>
<taxonomicName id="4C6F4D2FFFFDFFB8844CBBEC6C8751A8" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[346,661,1983,2007]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="524" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFFDFFB8844CBBEC6FE151A9" box="[346,499,1983,2006]" italics="true" pageId="7" pageNumber="524">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFFDFFB88789BBEC6D0E51A8" box="[671,796,1983,2007]" pageId="7" pageNumber="524" rank="species">new species</taxonomicNameLabel>
, mandibles in ventral view (F3, F2 and F1)
</paragraph>
</caption>
<paragraph id="8BD036ACFFFDFFB881C2BBEC6ACA51A8" blockId="7.[1236,1240,1983,2007]" box="[1236,1240,1983,2007]" pageId="7" pageNumber="524">.</paragraph>
<caption id="DF106624FFFDFFB88581BBB16A7A5184" ID-DOI="http://doi.org/10.5281/zenodo.7510350" ID-Zenodo-Dep="7510350" box="[151,1128,2018,2043]" httpUri="https://zenodo.org/record/7510350/files/figure.png" pageId="7" pageNumber="524" startId="7.[151,250,2018,2042]" targetBox="[151,1280,181,1866]" targetPageId="7">
<paragraph id="8BD036ACFFFDFFB88581BBB16A7A5184" blockId="7.[151,1128,2018,2043]" box="[151,1128,2018,2043]" pageId="7" pageNumber="524">
<emphasis id="B91BEABEFFFDFFB88581BBB16F325184" bold="true" box="[151,288,2018,2043]" pageId="7" pageNumber="524">FIGURE 32.</emphasis>
<taxonomicName id="4C6F4D2FFFFDFFB88430BBB06C845184" authority="Zhang, 1988" authorityName="Zhang" authorityYear="1988" box="[294,662,2019,2043]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="524" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFFDFFB88430BBB06C1F5184" box="[294,525,2019,2043]" italics="true" pageId="7" pageNumber="524">Xizangia cryptonychus</emphasis>
<bibRefCitation id="EFFE4B5DFFFDFFB88702BBB06C845184" author="Zhang, Y. - W." box="[532,662,2019,2043]" pageId="7" pageNumber="524" pagination="233 - 237" refId="ref10547" refString="Zhang, Y. - W. (1988) Coleoptera: Trogidae. In: F. - S. Huang, F. - S. (Ed.), Insects of Mt. Namjagbarwa Region of Xizang. Science Press, Beijing, pp. 233 - 237. [in Chinese]" type="book chapter" year="1988">Zhang, 1988</bibRefCitation>
</taxonomicName>
, mandibles in ventral view (female
<typeStatus id="54D4880EFFFDFFB88117BBB06A4F5184" box="[1025,1117,2019,2043]" pageId="7" pageNumber="524" type="paratype">paratype</typeStatus>
).
</paragraph>
</caption>
<caption id="DF106624FFF2FFB78581BBC46D1B51CF" ID-DOI="http://doi.org/10.5281/zenodo.6375335" ID-Zenodo-Dep="6375335" box="[151,777,1943,1968]" httpUri="https://zenodo.org/record/6375335/files/figure.png" pageId="8" pageNumber="525" startId="8.[151,250,1943,1967]" targetBox="[174,1436,181,1932]" targetPageId="8">
<paragraph id="8BD036ACFFF2FFB78581BBC46D1B51CF" blockId="8.[151,777,1943,1968]" box="[151,777,1943,1968]" pageId="8" pageNumber="525">
<emphasis id="B91BEABEFFF2FFB78581BBC46F3251D0" bold="true" box="[151,288,1943,1967]" pageId="8" pageNumber="525">FIGURE 33.</emphasis>
Diagnostic characters of
<taxonomicName id="4C6F4D2FFFF2FFB78735BBCB6CC251CF" box="[547,720,1944,1968]" pageId="8" pageNumber="525">
<emphasis id="B91BEABEFFF2FFB78735BBCB6C6F51D0" box="[547,637,1944,1967]" italics="true" pageId="8" pageNumber="525">Xizangia</emphasis>
species
</taxonomicName>
: legs.
</paragraph>
</caption>
<caption id="DF106624FFF2FFB78581BBE86C595187" ID-DOI="http://doi.org/10.5281/zenodo.7510494" ID-Zenodo-Dep="7510494" httpUri="https://zenodo.org/record/7510494/files/figure.png" pageId="8" pageNumber="525" startId="8.[151,250,1979,2003]" targetBox="[174,1436,181,1932]" targetPageId="8">
<paragraph id="8BD036ACFFF2FFB78581BBE86C595187" blockId="8.[151,1436,1979,2040]" pageId="8" pageNumber="525">
<emphasis id="B91BEABEFFF2FFB78581BBE86F0D51AC" bold="true" box="[151,287,1979,2003]" pageId="8" pageNumber="525">FIGURE 34.</emphasis>
<taxonomicName id="4C6F4D2FFFF2FFB78433BBEF6C4E51AB" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[293,604,1980,2004]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="8" pageNumber="525" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF2FFB78433BBEF6FAE51AC" box="[293,444,1980,2003]" italics="true" pageId="8" pageNumber="525">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF2FFB78770BBEF6CF051AB" box="[614,738,1980,2004]" pageId="8" pageNumber="525" rank="species">new species</taxonomicNameLabel>
, mandibles in dorsal, inner lateral and ventral views: left half — male mandibles; right half — female mandibles.
</paragraph>
</caption>
<caption id="DF106624FFF3FFB68581BBC46B6C51AB" ID-DOI="http://doi.org/10.5281/zenodo.6375337" ID-Zenodo-Dep="6375337" httpUri="https://zenodo.org/record/6375337/files/figure.png" pageId="9" pageNumber="526" startId="9.[151,264,1943,1967]" targetBox="[230,1358,181,1920]" targetPageId="9">
<paragraph id="8BD036ACFFF3FFB68581BBC46B6C51AB" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="526">
<emphasis id="B91BEABEFFF3FFB68581BBC46F4651D0" bold="true" box="[151,340,1943,1967]" pageId="9" pageNumber="526">FIGURES 35–39.</emphasis>
<taxonomicName id="4C6F4D2FFFF3FFB6844CBBCB6C8651CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[346,660,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="9" pageNumber="526" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF3FFB6844CBBCB6FE051D0" box="[346,498,1944,1967]" italics="true" pageId="9" pageNumber="526">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF3FFB68788BBCB6D0851CF" box="[670,794,1944,1968]" pageId="9" pageNumber="526" rank="species">new species</taxonomicNameLabel>
, characters. 35, hindwing; 36, female antenna in various views; 37, maxilla and labium; 38, clypeus, labrum and mentum in anterior view; 39, clypeus, labrum and mentum in ventral view.
</paragraph>
</caption>
<subSubSection id="C3756527FFF0FFB48581BCC46A2A55FE" lastPageId="11" lastPageNumber="528" pageId="10" pageNumber="527" type="description">
<paragraph id="8BD036ACFFF0FFB58581BCC46F8A56CF" blockId="10.[151,1437,150,2013]" box="[151,408,150,177]" pageId="10" pageNumber="527">
<heading id="D09881C0FFF0FFB58581BCC46F8A56CF" bold="true" box="[151,408,150,177]" fontSize="11" level="1" pageId="10" pageNumber="527" reason="1">
<emphasis id="B91BEABEFFF0FFB58581BCC46F8A56CF" bold="true" box="[151,408,150,177]" pageId="10" pageNumber="527">
<typeStatus id="54D4880EFFF0FFB58581BCC46F1656CE" box="[151,260,151,177]" pageId="10" pageNumber="527" type="holotype">Holotype</typeStatus>
description.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036ACFFF0FFB585D1BCE86D275762" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="527">Measurement. Body length, measured from the anterior margin of head to the posterior margin of elytra: 6.68 mm; body thickness: 1.35 mm; head width: 2.53 mm; pronotum width: 2.85 mm; elytra width: 2.90 mm; body thickness/body length: 0.20; elytra width/body length: 0.43.</paragraph>
<paragraph id="8BD036ACFFF0FFB585D1BD746D805076" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="527">
Body rectangular, strongly compressed dorso-ventrally; color dark reddish brown. Head: Transverse, surface punctate and glabrous; clypeus oblong, defined posteriorly by a transversal depression and laterally by a pair of longitudinal depressions; anterior margin broadly rounded, not emarginated, at most with a pair of weak indentations at lateral corners of clypeus, and with mandibles invisible from above; ocular canthus very large, dilated. Labrum small, triangular in anterior view and trapezoidal in ventral view, completely fused with clypeus (without suture), raised from the ventral surface of clypeus, extending downwards to fix the inner sides of mandibles. Mandibles small and slender, sharply pointed at apex; left mandible with a ventral tooth, right mandible with a medial tooth raised from dorsal surface. Maxilla: galea very small and brush-like at apex, lacinia small and free at apex, without a hook, maxillary palpus with 4 palpomeres, with terminal palpomere 3 times as long as wide. Labium consisting of a ligula and separated palpus on anterior portion of dorsal surface of mentum; ligula stick-like, longer than wide, rounded at apex and not bilobed, extending ventrally between bases of palpal insertions; palpal insertions separated by the ventral extension of ligula which is as wide as basal palpifer; labial palpus with 3 palpomeres (not counting basal palpifer).
<collectingRegion id="49ABF84EFFF0FFB58453BE846FB6548E" box="[325,420,727,753]" country="Myanmar" name="Chin" pageId="10" pageNumber="527">Mentum</collectingRegion>
in ventral view more or less trapezoidal, with both anterior and posterior margins concave; mentum in anterior view half-rounded with outer anterior margin convex; surface sparsely punctate, glabrous. Submentum expanded at anterior part, sparsely setose, not clearly separated from impunctate, glabrous gula. Eye twice as long as wide in dorsal view. Antenna with ten antennomeres; both antennomeres 1 and 2 geniculate; antennomeres 3–7 being gradually widened distally and without hairy sensorial areas, antennomeres 6 and 7 being more pointed at inner apex, antennomeres 5–7 with a pair of setae at inner apex; club distinctly formed by the last 3 antennomeres (8–10), antennomere 8 with a small hairy sensorial area at inner apex, antennomere 9 being mostly hairy except for distal marginal area, antennomere 10 being completely hairy on external surfaces (glabrous only at proximal side); 2 small, round holes present longitudinally (transversely in
<taxonomicName id="4C6F4D2FFFF0FFB586EBBFA46B06526D" box="[1021,1300,1015,1042]" pageId="10" pageNumber="527">
<emphasis id="B91BEABEFFF0FFB586EBBFA46AA8526E" box="[1021,1210,1015,1041]" italics="true" pageId="10" pageNumber="527">Penichrolucanus</emphasis>
species
</taxonomicName>
) in the midline on top of antennomere 10. Pronotum: Transverse, slightly wider in posterior half than in anterior half, base convex, anterior margin almost straight; surface punctate and glabrous, each side with superficial longitudinal, curved wrinkles and with a small laterodiscal depression. Intercoxal process of prosternum narrow, convex dorsally, expanding posteriorly, with all margins not ridged, not high above mesosternum, and associated with a pair of additional tubercles at posterior corners. Mesocoxae separated by flat, clearly ridged anterior projection of metasternum. Metacoxae remote from mesocoxae, not intruded by posterior part of metasternum. Scutellum: Ogival. Elytra: Longer than head and prothorax combined, parallel-sided, with 6 thin striae, interstriae superficially punctate; intervals between striae very finely punctate.Abdomen with five visible abdominal ventrites; first projected anteriorly, forming an intercoxal process of hind legs, somewhat triangular in shape; all ventrites finely punctate and glabrous, with lateral wrinkles as in metasternum. Legs: Short; all tibiae straight, protibia with three continuous triangular teeth in addition to the well-developed apical fork, meso—and metatibiae with two spines on external margins; protarsus with tarsomeres 1–4 short, subequal in length, fused together, tarsomere 5 longer than others combined; meso—and metatarsi with 3 segments, and with last tarsomere more depressed than in protarsus; a pair of claws hidden in terminal cavity of last tarsomere, arolium absent.
</paragraph>
<paragraph id="8BD036ACFFF0FFB585D1BA406C85500A" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="527">
Hindwing (terminology follows
<bibRefCitation id="EFFE4B5DFFF0FFB5872CBA406DD25052" author="Kukalova-Peck, J. &amp; Lawrence, J. F." box="[570,960,1555,1581]" pageId="10" pageNumber="527" pagination="181 - 258" refId="ref9893" refString="Kukalova-Peck, J. &amp; Lawrence, J. F. (1993) Evolution of the hindwing in Coleoptera. Canadian Entomologist, 125, 181 - 258. https: // doi. org / 10.4039 / Ent 125181 - 2" type="journal article" year="1993">Kukalová-Peck &amp; Lawrence 1993</bibRefCitation>
) 7.4 mm long; vein RP absent, vein MP
<subScript id="17EB34E9FFF0FFB58082BA766B8E504C" attach="left" box="[1428,1436,1573,1587]" fontSize="6" pageId="10" pageNumber="527">3</subScript>
absent, vein RA
<subScript id="17EB34E9FFF0FFB5845ABA1A6F465028" attach="left" box="[332,340,1609,1623]" fontSize="6" pageId="10" pageNumber="527">4</subScript>
remote from costa of hindwing, vein RP
<subScript id="17EB34E9FFF0FFB58636BA1A6D3A5028" attach="left" box="[800,808,1609,1623]" fontSize="6" pageId="10" pageNumber="527">1</subScript>
with basal part present but with apical part absent, vein AA
<subScript id="17EB34E9FFF0FFB585D6BA3E6ECB5004" attach="left" box="[192,217,1645,1659]" fontSize="6" pageId="10" pageNumber="527">1+2</subScript>
reaching vein CuA to form a close cell.
</paragraph>
<paragraph id="8BD036ACFFF0FFB485D1BA2C6DB556AA" blockId="10.[151,1437,150,2013]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="528" pageId="10" pageNumber="527">
Last tergite without colourless stripe. Last sternite without colourless area, and with posterior margin in a smooth curve at middle. Paired pleurites of 9th abdominal segment large and close, contracted at medial part. Ventral plate of 9th abdominal segment expanded at terminal end, strongly sclerotized, without lateral processes, and with no longitudinal carinae. Male genitalia with a pair of struts articulating near the proximal end of median lobe and enclosed by basal piece; basal piece with no dorsal plates, and with ventral surface largely colourless and membranous at caudal part, forming a large triangular excavation on the pigmented part; median lobe with a dorsal crossbar at base and with lateral bridge between base of median lobe and dorsal crossbar; dorsal crossbar of median lobe with dorsal margin connected by membrane to an extra plate (not found in other
<taxonomicName id="4C6F4D2FFFF0FFB58174BB286AC551EA" baseAuthorityName="sensu Holloway" baseAuthorityYear="1968" box="[1122,1239,1915,1941]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">Lucaninae</taxonomicName>
tribes) extending proximally into basal piece; median lobe simple, oblong, depressed dorso-ventrally, even in width, extending proximally and rounded caudally in ventral view, with ventral surface evenly pigmented, and with dorsal surface membranous and not expansible; permanently everted internal sac depressed and nearly even in width throughout, neither contracted nor expanded at apex, and clothed with microsetae.
</paragraph>
<paragraph id="8BD036ACFFF1FFB48581BD506F435762" blockId="11.[151,1436,259,393]" box="[151,337,259,285]" pageId="11" pageNumber="528">
<heading id="D09881C0FFF1FFB48581BD506F435762" bold="true" box="[151,337,259,285]" fontSize="11" level="1" pageId="11" pageNumber="528" reason="1">
<emphasis id="B91BEABEFFF1FFB48581BD506F435762" bold="true" box="[151,337,259,285]" pageId="11" pageNumber="528">
Male
<typeStatus id="54D4880EFFF1FFB485CDBD506F435762" box="[219,337,259,285]" pageId="11" pageNumber="528" type="paratype">paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BD746C18573E" blockId="11.[151,1436,259,393]" box="[199,522,295,321]" pageId="11" pageNumber="528">Body length: 6.60–7.30 mm.</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BD186FEA57F6" blockId="11.[151,1436,259,393]" pageId="11" pageNumber="528">Variation.Anterior margin of head a little variable in the detailed shape. Meso—and metatibiae sometimes with additional one or two denticles.</paragraph>
<paragraph id="8BD036ACFFF1FFB48581BDE46C8457AD" blockId="11.[151,1437,439,1293]" box="[151,662,439,466]" pageId="11" pageNumber="528">
<emphasis id="B91BEABEFFF1FFB48581BDE46F7957AE" bold="true" box="[151,363,439,465]" pageId="11" pageNumber="528">
Female
<typeStatus id="54D4880EFFF1FFB485E3BDE46F7957AE" box="[245,363,439,465]" pageId="11" pageNumber="528" type="paratype">paratypes</typeStatus>
</emphasis>
(
<figureCitation id="13542A29FFF1FFB4846DBDE46FF457AE" box="[379,486,439,465]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1965,1989]" captionTargetBox="[193,1390,214,1907]" captionTargetId="figure-19@3.[151,1435,181,1942]" captionTargetPageId="3" captionText="FIGURES 1–2. Xizangia qiuae Huang &amp; Chen, new species. 1, habitus in dorsal view; 2, habitus in ventral view (F4—female paratype from Mangdong)." figureDoi="http://doi.org/10.5281/zenodo.6375325" httpUri="https://zenodo.org/record/6375325/files/figure.png" pageId="11" pageNumber="528">Figs. 1–2</figureCitation>
,
<figureCitation id="13542A29FFF1FFB484E4BDE46C2957AE" box="[498,571,439,465]" captionStart="FIGURES 15–19" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[204,1436,181,1896]" captionTargetId="figure-21@6.[266,1321,181,1885]" captionTargetPageId="6" captionText="FIGURES 15–19. Xizangia qiuae Huang &amp; Chen, new species, habitus in dorsal view (M1—male holotype, F1—female paratype, F2—female paratype, F3—female paratype and F4 —female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6375331" httpUri="https://zenodo.org/record/6375331/files/figure.png" pageId="11" pageNumber="528">16–19</figureCitation>
,
<figureCitation id="13542A29FFF1FFB48750BDE46C9C57AD" box="[582,654,439,466]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[151,1280,181,1866]" captionTargetId="figure-19@7.[151,1164,181,1851]" captionTargetPageId="7" captionText="FIGURES 22–26. Xizangia qiuae Huang &amp; Chen, new species, habitus in ventral view (M1, F1, F2, F3 and F4)." figureDoi="http://doi.org/10.5281/zenodo.6375333" httpUri="https://zenodo.org/record/6375333/files/figure.png" pageId="11" pageNumber="528">23–26</figureCitation>
)
</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BD886C18578A" blockId="11.[151,1437,439,1293]" box="[199,522,474,501]" pageId="11" pageNumber="528">Body length: 6.70–7.50 mm.</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BDAC6D2A541D" blockId="11.[151,1437,439,1293]" pageId="11" pageNumber="528">Sexual dimorphism only found in length of ventral tooth of left mandible and length of last abdominal ventrite. Ventral tooth of left mandible longer than in male. Last ventrite more than 1.5 times longer than penultimate ventrite (less than 1.5 times as long as penultimate ventrite in male).</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BE386DDF54FA" blockId="11.[151,1437,439,1293]" box="[199,973,619,645]" pageId="11" pageNumber="528">Hindwing as in male, with no difference in size, shape, and venation.</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BEDC6F845546" blockId="11.[151,1437,439,1293]" pageId="11" pageNumber="528">
Female genitalia (
<figureCitation id="13542A29FFF1FFB4848EBEDC6C3454D6" box="[408,550,655,681]" captionStart="FIGURES 41–42" captionStartId="13.[151,264,1980,2004]" captionTargetBox="[151,1436,181,1948]" captionTargetId="figure-19@13.[151,1435,181,1920]" captionTargetPageId="13" captionText="FIGURES 41–42. Xizangia qiuae Huang &amp; Chen, new species, female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6377198" httpUri="https://zenodo.org/record/6377198/files/figure.png" pageId="11" pageNumber="528">Figs. 41–42</figureCitation>
). Hemisternites well sclerotized and setose near apex, with styli sclerotized, non-setose, hawk-beak-like and pointed outwards; spermatheca tube-like and rounded at apex, pigmented at apical half but colourless at basal half; spermathecal duct not defined as the membranous tube connected to spermatheca expansible and better explained as bursa copulatrix; spermathecal gland and its duct originated from spermatheca near base, rather short.
</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1BF106A2A55FE" blockId="11.[151,1437,439,1293]" pageId="11" pageNumber="528">
The unique female specimen from Mangdong, SW.
<collectingRegion id="49ABF84EFFF1FFB48638BF106D955522" box="[814,903,835,861]" country="China" name="Yunnan" pageId="11" pageNumber="528">Yunnan</collectingRegion>
, another locality, does not show any reliable difference from the specimens from Xishuangbanna, SC.
<collectingRegion id="49ABF84EFFF1FFB4860CBF346D6155FE" box="[794,883,871,897]" country="China" name="Yunnan" pageId="11" pageNumber="528">Yunnan</collectingRegion>
, the
<typeStatus id="54D4880EFFF1FFB486B1BF346DC555FE" box="[935,983,871,897]" pageId="11" pageNumber="528">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF1FFB485D1BFD96F7C5296" pageId="11" pageNumber="528" type="diagnosis">
<paragraph id="8BD036ACFFF1FFB485D1BFD96AD6524A" blockId="11.[151,1437,439,1293]" pageId="11" pageNumber="528">
<emphasis id="B91BEABEFFF1FFB485D1BFD96F5055DB" bold="true" box="[199,322,906,932]" pageId="11" pageNumber="528">Diagnosis.</emphasis>
This new species is distinguishable from
<taxonomicName id="4C6F4D2FFFF1FFB4860ABFD86A0F55DA" authorityName="Zhang" authorityYear="1988" box="[796,1053,907,933]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="11" pageNumber="528" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFF1FFB4860ABFD86A0F55DA" box="[796,1053,907,933]" italics="true" pageId="11" pageNumber="528">Xizangia cryptonychus</emphasis>
</taxonomicName>
(
<figureCitation id="13542A29FFF1FFB4813BBFD86ABA55DA" box="[1069,1192,907,933]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[202,1386,181,1891]" captionTargetId="figure-19@5.[202,1386,181,1920]" captionTargetPageId="5" captionText="FIGURES 9–14. Xizangia cryptonychus Zhang, 1988, habitus and details (female paratype). 9, entire body in lateral view; 10, top of last antennomere; 11, part of head in ventral view; 12, antenna; 13, intercoxal process of prosternum; 14, part of head in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6375329" httpUri="https://zenodo.org/record/6375329/files/figure.png" pageId="11" pageNumber="528">Figs. 9–14</figureCitation>
,
<figureCitation id="13542A29FFF1FFB481A2BFD86AC155DA" box="[1204,1235,907,933]" captionStart="FIGURE 20" captionStartId="6.[151,250,1981,2005]" captionTargetBox="[204,1436,181,1896]" captionTargetId="figure-21@6.[266,1321,181,1885]" captionTargetPageId="6" captionText="FIGURE 20. Xizangia cryptonychus Zhang, 1988, habitus in dorsal view (female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6379417" httpUri="https://zenodo.org/record/6379417/files/figure.png" pageId="11" pageNumber="528">20</figureCitation>
,
<figureCitation id="13542A29FFF1FFB481C9BFD86AEE55DA" box="[1247,1276,907,933]" captionStart="FIGURE 27" captionStartId="7.[151,250,1910,1934]" captionTargetBox="[151,1280,181,1866]" captionTargetId="figure-19@7.[151,1164,181,1851]" captionTargetPageId="7" captionText="FIGURE 27. Xizangia cryptonychus Zhang, 1988, habitus in ventral view (female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6379421" httpUri="https://zenodo.org/record/6379421/files/figure.png" pageId="11" pageNumber="528">27</figureCitation>
) from Motuo, SE
<collectingRegion id="49ABF84EFFF1FFB485A9BFFC6EEB55B6" box="[191,249,943,969]" country="China" name="Xizang" pageId="11" pageNumber="528">Tibet</collectingRegion>
in female by having a shorter ventral tooth of left mandible (
<figureCitation id="13542A29FFF1FFB486BBBFFC6A2455B6" box="[941,1078,943,969]" captionStart-0="FIGURES 29–31" captionStart-1="FIGURE 32" captionStartId-0="7.[151,264,1982,2006]" captionStartId-1="7.[151,250,2018,2042]" captionTargetBox-0="[151,1280,181,1866]" captionTargetBox-1="[151,1280,181,1866]" captionTargetId-0="figure-19@7.[151,1164,181,1851]" captionTargetId-1="figure-19@7.[151,1164,181,1851]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURES 29–31. Xizangia qiuae Huang &amp; Chen, new species, mandibles in ventral view (F3, F2 and F1)" captionText-1="FIGURE 32. Xizangia cryptonychus Zhang, 1988, mandibles in ventral view (female paratype)." figureDoi-0="http://doi.org/10.5281/zenodo.7510346" figureDoi-1="http://doi.org/10.5281/zenodo.7510350" httpUri-0="https://zenodo.org/record/7510346/files/figure.png" httpUri-1="https://zenodo.org/record/7510350/files/figure.png" pageId="11" pageNumber="528">Figs. 29–32</figureCitation>
), a shorter inner spur at apex of protibia (
<figureCitation id="13542A29FFF1FFB485EBBF806F425592" box="[253,336,979,1005]" captionStart="FIGURE 33" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[174,1436,181,1932]" captionTargetId="figure-21@8.[230,1358,181,1920]" captionTargetPageId="8" captionText="FIGURE 33. Diagnostic characters of Xizangia species: legs." figureDoi="http://doi.org/10.5281/zenodo.6375335" httpUri="https://zenodo.org/record/6375335/files/figure.png" pageId="11" pageNumber="528">Fig. 33</figureCitation>
), the shorter last tarsomeres in all legs (
<figureCitation id="13542A29FFF1FFB4861DBF806D4C5592" box="[779,862,979,1005]" captionStart="FIGURE 33" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[174,1436,181,1932]" captionTargetId="figure-21@8.[230,1358,181,1920]" captionTargetPageId="8" captionText="FIGURE 33. Diagnostic characters of Xizangia species: legs." figureDoi="http://doi.org/10.5281/zenodo.6375335" httpUri="https://zenodo.org/record/6375335/files/figure.png" pageId="11" pageNumber="528">Fig. 33</figureCitation>
), the fewer external spines in all tibiae, and the less convex intervals between striae on the elytra. It can be supposed that these differences should also be present in male as no sexual dimorphism of these characters is found in the known species of
<taxonomicName id="4C6F4D2FFFF1FFB486E8B8486AD2524A" box="[1022,1216,1051,1077]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="519" phylum="Arthropoda" rank="tribe" tribe="Penichrolucanini">Penichrolucanini</taxonomicName>
.
</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1B86C6F7C5296" blockId="11.[151,1437,439,1293]" pageId="11" pageNumber="528">
Only the
<specimenCount id="9D69FD25FFF1FFB4843CB86C6FB75226" box="[298,421,1087,1113]" pageId="11" pageNumber="528" type="female">two female</specimenCount>
type specimens are known for
<taxonomicName id="4C6F4D2FFFF1FFB487E5B86C6DE25226" authorityName="Zhang" authorityYear="1988" box="[755,1008,1087,1113]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="11" pageNumber="528" phylum="Foraminifera" rank="species" species="cryptonychus">
<emphasis id="B91BEABEFFF1FFB487E5B86C6DE25226" box="[755,1008,1087,1113]" italics="true" pageId="11" pageNumber="528">Xizangia cryptonychus</emphasis>
</taxonomicName>
, both deposited in Institute of Zoology, Chinese Academy of Science; the
<typeStatus id="54D4880EFFF1FFB48709B8306C935202" box="[543,641,1123,1149]" pageId="11" pageNumber="528" type="holotype">holotype</typeStatus>
remains untraceable (M.-Y. Lin &amp; M. Bai, personal communications, 2018). The last abdominal segments of the only available female
<typeStatus id="54D4880EFFF1FFB4869DB8D46DF952DE" box="[907,1003,1159,1185]" pageId="11" pageNumber="528" type="paratype">paratype</typeStatus>
are missing, most probably dissected by its author during the first research, therefore a comparison of genital structures between the two species is not possible at present.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF1FFB485D1B8A06B9D5372" box="[199,1423,1267,1293]" pageId="11" pageNumber="528" type="etymology">
<paragraph id="8BD036ACFFF1FFB485D1B8A06B9D5372" blockId="11.[151,1437,439,1293]" box="[199,1423,1267,1293]" pageId="11" pageNumber="528">
<emphasis id="B91BEABEFFF1FFB485D1B8A06F5C5372" bold="true" box="[199,334,1267,1293]" pageId="11" pageNumber="528">Etymology.</emphasis>
The new species is named in honor of Dr. Qiu Jian-Yue, one of the collectors of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF1FFB38581B9686DB05522" lastPageId="12" lastPageNumber="529" pageId="11" pageNumber="528" type="description">
<paragraph id="8BD036ACFFF1FFB48581B9686C0C532A" blockId="11.[151,1437,1338,2013]" box="[151,542,1338,1365]" pageId="11" pageNumber="528">
<emphasis id="B91BEABEFFF1FFB48581B9686F92532A" bold="true" box="[151,384,1339,1365]" pageId="11" pageNumber="528">Larval morphology</emphasis>
(
<figureCitation id="13542A29FFF1FFB48499B9686C04532A" box="[399,534,1338,1365]" captionStart-0="FIGURES 44–48" captionStart-1="FIGURES 49–52" captionStart-2="FIGURES 53–57" captionStartId-0="15.[151,264,1943,1967]" captionStartId-1="16.[151,264,1943,1967]" captionStartId-2="17.[151,264,1943,1967]" captionTargetBox-0="[194,1393,181,1920]" captionTargetBox-1="[203,1385,181,1920]" captionTargetBox-2="[194,1393,181,1920]" captionTargetId-0="figure-19@15.[194,1393,181,1920]" captionTargetId-1="figure-21@16.[203,1385,181,1920]" captionTargetId-2="figure-19@17.[194,1393,181,1920]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." captionText-1="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." captionText-2="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." figureDoi-0="http://doi.org/10.5281/zenodo.6375345" figureDoi-1="http://doi.org/10.5281/zenodo.6375349" figureDoi-2="http://doi.org/10.5281/zenodo.6375351" httpUri-0="https://zenodo.org/record/6375345/files/figure.png" httpUri-1="https://zenodo.org/record/6375349/files/figure.png" httpUri-2="https://zenodo.org/record/6375351/files/figure.png" pageId="11" pageNumber="528">Figs. 44–57</figureCitation>
)
</paragraph>
<paragraph id="8BD036ACFFF1FFB485D1B90C6AC85306" blockId="11.[151,1437,1338,2013]" box="[199,1242,1374,1401]" pageId="11" pageNumber="528">
A detailed description of the third instar larva of
<taxonomicName id="4C6F4D2FFFF1FFB487FCB90C6D805307" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[746,914,1375,1400]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="11" pageNumber="528" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF1FFB487FCB90C6D805307" box="[746,914,1375,1400]" italics="true" pageId="11" pageNumber="528">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFF1FFB486B7B90C6A385307" bold="true" box="[929,1066,1374,1401]" pageId="11" pageNumber="528">
<taxonomicNameLabel id="A22857C5FFF1FFB486B7B90C6A385307" box="[929,1066,1374,1401]" pageId="11" pageNumber="528" rank="species">new species</taxonomicNameLabel>
</emphasis>
is given below.
</paragraph>
<paragraph id="8BD036ACFFF1FFB385D1B9D06F59571A" blockId="11.[151,1437,1338,2013]" lastBlockId="12.[151,1437,151,1977]" lastPageId="12" lastPageNumber="529" pageId="11" pageNumber="528">
Antenna (
<figureCitation id="13542A29FFF1FFB48420B9D06F8653E2" box="[310,404,1411,1437]" captionStart="FIGURES 44–48" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@15.[194,1393,181,1920]" captionTargetPageId="15" captionText="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6375345" httpUri="https://zenodo.org/record/6375345/files/figure.png" pageId="11" pageNumber="528">Figs. 48</figureCitation>
,
<figureCitation id="13542A29FFF1FFB484B6B9D16FF953E2" box="[416,491,1410,1437]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@17.[194,1393,181,1920]" captionTargetPageId="17" captionText="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." figureDoi="http://doi.org/10.5281/zenodo.6375351" httpUri="https://zenodo.org/record/6375351/files/figure.png" pageId="11" pageNumber="528">53–54</figureCitation>
): Antenna 3-segmented (excluding antennifer). Basal antennomere subdivided, with an unpigmented ring near base, indicating it is formed by two segments, completely fused; it bearing very few sensory spots and small setae. Penultimate antennomere (antennomere 2) as long as and markedly wider than basal antennomere, with more sensory spots and small setae than basal antennomere, and with no large sensorium below last antennomere (antennomere 3). Last antennomere very small, with very few setae and sensory spots, and with no apical appendage. Epipharynx (
<figureCitation id="13542A29FFF1FFB4870ABA646C62502E" box="[540,624,1590,1617]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@17.[194,1393,181,1920]" captionTargetPageId="17" captionText="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." figureDoi="http://doi.org/10.5281/zenodo.6375351" httpUri="https://zenodo.org/record/6375351/files/figure.png" pageId="11" pageNumber="528">Fig. 56</figureCitation>
): Phoba divided into an anterior section (namely protophoba) and lateral sections (namely chaetoparia). Protophoba distinct and consisting of 11 short, blunt and densely arranged setae, and with about 3 sensory spots. Haptomerum hardly separated from protophoba, consisting of 2 sensory spots, without setae. Setae in chaetoparia very few and much sparser than in protophoba. Haptolachus consisting of 2 nesia, the median one being more elongate and bearing very small granules; all nesia without setae. Epitorma well sclerotized but very small and short. Pternotorma well developed and very long. Mandibles (
<figureCitation id="13542A29FFF1FFB48128BAB86A86517A" box="[1086,1172,1770,1797]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@17.[194,1393,181,1920]" captionTargetPageId="17" captionText="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." figureDoi="http://doi.org/10.5281/zenodo.6375351" httpUri="https://zenodo.org/record/6375351/files/figure.png" pageId="11" pageNumber="528">Fig. 55</figureCitation>
): Left mandible with 3 apical teeth at tip, a complicated mola at basal half, a ventral process at base and a minute inner tubercle along the inner edge between apical teeth and mola; apical teeth all blunt, with dorsal one shorter and ventral one longer; inner tubercle minute, not forming clear tooth; mola triangular in both dorsal and ventral views, somewhat square in inner lateral view, and with a deep furrow between dorsal and ventral surfaces. Right mandible with 2 apical teeth at tip, 2 simple molae at base and an inner tubercle between apical teeth and molae; apical teeth with dorsal one shorter and blunter than ventral one; inner tubercle minute; both basal and distal molae restricted to dorsal side, rather short; ventral process fully developed as in left mandible. Outer edge of both mandibles coarse and dentate, with very few minute setae. Mouthparts (
<figureCitation id="13542A29FFF6FFB384D4BCE86C0456AA" box="[450,534,186,213]" captionStart="FIGURES 53–57" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@17.[194,1393,181,1920]" captionTargetPageId="17" captionText="FIGURES 53–57. Xizangia qiuae Huang &amp; Chen, new species, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart." figureDoi="http://doi.org/10.5281/zenodo.6375351" httpUri="https://zenodo.org/record/6375351/files/figure.png" pageId="12" pageNumber="529">Fig. 57</figureCitation>
): Stipes without stridulatory teeth. Galea and lacinia separate and fixed to stipes. Galea with membranous subdivision. Lacinia single-pointed at apex. Maxillary palpus with 3 palpomeres excluding palpifer and with palpifer present. Ligula absent. Labial palpus with 2 palpomeres, rather straight and glabrous, without palpifer. Hypopharynx not markedly asymmetrical, with right dorsally pointed sclerotized projection visible in oblique view.
</paragraph>
<paragraph id="8BD036ACFFF6FFB385D1BD3C6D5854B2" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
Prothorax (
<figureCitation id="13542A29FFF6FFB38450BD3C6F8F57F6" box="[326,413,367,393]" captionStart="FIGURES 44–48" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@15.[194,1393,181,1920]" captionTargetPageId="15" captionText="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6375345" httpUri="https://zenodo.org/record/6375345/files/figure.png" pageId="12" pageNumber="529">Fig. 44</figureCitation>
): Pronotum without an anteriorly projecting lobe or an extensive pronotal plate. Precoxale and episternum hardly sclerotized. Mesocoxal stridulatory organ (pars stridens,
<figureCitation id="13542A29FFF6FFB38101BDC06A7957D2" box="[1047,1131,402,429]" captionStart="FIGURES 49–52" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." figureDoi="http://doi.org/10.5281/zenodo.6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="12" pageNumber="529">Fig. 50</figureCitation>
): Pars stridens on posterior surface of mesocoxa consisting of a main carina and a field of minute, rounded granules, all about equal in size, without well developed granules outside of the main row. Metatrochanteral stridulatory organ (plectrum,
<figureCitation id="13542A29FFF6FFB38073BD886EF15466" captionStart="FIGURES 49–52" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." figureDoi="http://doi.org/10.5281/zenodo.6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="12" pageNumber="529">Figs. 51–52</figureCitation>
): Plectrum on anterior surface of metatrochanter consisting of a single row of continuous granules which might be interrupted by a empty field; granules a little rounded or oblong, smaller in terminal part than in basal part. Tarsungulus: Tarsungulus of prothoracic leg (
<figureCitation id="13542A29FFF6FFB38798BE146CCD541E" box="[654,735,583,609]" captionStart="FIGURES 49–52" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." figureDoi="http://doi.org/10.5281/zenodo.6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="12" pageNumber="529">Fig. 49</figureCitation>
) 2 times longer and more gradually attenuated at apex than that of mesothoracic (
<figureCitation id="13542A29FFF6FFB3844CBE386FB954FA" box="[346,427,618,645]" captionStart="FIGURES 49–52" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." figureDoi="http://doi.org/10.5281/zenodo.6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="12" pageNumber="529">Fig. 50</figureCitation>
) and metathoracic legs (
<figureCitation id="13542A29FFF6FFB387AABE386D1C54FA" box="[700,782,618,645]" captionStart="FIGURES 49–52" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURES 49–52. Xizangia qiuae Huang &amp; Chen, new species, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg." figureDoi="http://doi.org/10.5281/zenodo.6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="12" pageNumber="529">Fig. 51</figureCitation>
), flanked by 2 shorter setae arising from the same position on each side. Tarsungulus of mesothoracic and metathoracic legs strongly and abruptly attenuated at apex, flanked by 2 longer setae arising from the same position on each side.
</paragraph>
<paragraph id="8BD036ACFFF6FFB385D1BE846DB05522" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
Vestiture of thoracic and abdominal tergites (
<figureCitation id="13542A29FFF6FFB387D4BE846D5E548E" box="[706,844,726,753]" captionStart="FIGURES 44–48" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@15.[194,1393,181,1920]" captionTargetPageId="15" captionText="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6375345" httpUri="https://zenodo.org/record/6375345/files/figure.png" pageId="12" pageNumber="529">Figs. 44–45</figureCitation>
): All tergites sparsely clad with small setae; spinules absent from thoracic tergites and abdominal tergites 1 and 7–10, very few on abdominal tergites 2–6. Vestiture of abdominal sternite 10 (
<figureCitation id="13542A29FFF6FFB38488BF4C6FE65545" box="[414,500,799,826]" captionStart="FIGURES 44–48" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[194,1393,181,1920]" captionTargetId="figure-19@15.[194,1393,181,1920]" captionTargetPageId="15" captionText="FIGURES 44–48. Xizangia qiuae Huang &amp; Chen, new species, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6375345" httpUri="https://zenodo.org/record/6375345/files/figure.png" pageId="12" pageNumber="529">Fig. 46</figureCitation>
). Raster well developed as two lateral groups of posteromesally directed spinules. Anal sclerite. Dorsal lobe obsolete. Lateral lobes with very few setae.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFF6FFAA85D1BF346FFF573E" lastPageId="21" lastPageNumber="538" pageId="12" pageNumber="529" type="discussion">
<paragraph id="8BD036ACFFF6FFB385D1BF346F6B52BA" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
<emphasis id="B91BEABEFFF6FFB385D1BF346F0355FE" bold="true" box="[199,273,871,897]" pageId="12" pageNumber="529">Notes.</emphasis>
In
<bibRefCitation id="EFFE4B5DFFF6FFB38420BF346FC255FE" author="Emden, F." box="[310,464,870,897]" pageId="12" pageNumber="529" pagination="178 - 200" refId="ref9485" refString="Emden, F. (1935) Die Gattungsunterschiede der Hirschkaferlarven, ein Beitrag zum naturlichen System der Familie (Col. Lucan.). Stettiner Entomologische Zeitung, 93, 178 - 200. [in Germany]" type="journal article" year="1935">Emden (1935</bibRefCitation>
,
<bibRefCitation id="EFFE4B5DFFF6FFB384CABF356C0755FE" author="Emden, F." box="[476,533,870,897]" pageId="12" pageNumber="529" pagination="301 - 310" refId="ref9525" refString="Emden, F. (1951) The larvae of Dendezia and Figulus, with notes on some other larvae of Lucanidae. Revue de Zoologie et de Botanique Africaines, 46, 301 - 310." type="journal article" year="1951">1951</bibRefCitation>
) and
<bibRefCitation id="EFFE4B5DFFF6FFB38742BF346D0655FE" author="Lawrence, J. F." box="[596,788,871,897]" pageId="12" pageNumber="529" pagination="213 - 219" refId="ref9936" refString="Lawrence, J. F. (1981) Notes on larval Lucanidae (Coleoptera). Journal of the Australian Entomological Society, 20, 213 - 219. https: // doi. org / 10.1111 / j. 1440 - 6055.1981. tb 01035. x" type="journal article" year="1981">Lawrence (1981)</bibRefCitation>
works the number of inner teeth on the left mandible was used as a major character in larval classification, and a major split is based on the presence of at least one tooth along the inner edge of the left mandible between the two or three apical teeth and the mola, with the larvae possessing such inner teeth belonging to the subfamily
<taxonomicName id="4C6F4D2FFFF6FFB38793BF806A155591" authority="(sensu Holloway 1968)" baseAuthorityName="sensu Holloway" baseAuthorityYear="1968" box="[645,1031,979,1006]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">
Lucaninae (
<emphasis id="B91BEABEFFF6FFB3861CBF806D5A5593" box="[778,840,979,1004]" italics="true" pageId="12" pageNumber="529">sensu</emphasis>
<bibRefCitation id="EFFE4B5DFFF6FFB38646BF806DEC5591" author="Holloway, B. A." box="[848,1022,979,1006]" pageId="12" pageNumber="529" pagination="264 - 269" refId="ref9714" refString="Holloway, B. A. (1968) The relationships of Syndesus Macleay and Sinodendron Schneider (Coleoptera: Lucanidae). New Zealand Journal of Science, 11, 264 - 269." type="journal article" year="1968">Holloway 1968</bibRefCitation>
)
</taxonomicName>
whilst the group lacking inner teeth includes
<taxonomicName id="4C6F4D2FFFF6FFB385EABFA46C19526E" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[252,523,1015,1041]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB385EABFA46F41526E" box="[252,339,1015,1041]" italics="true" pageId="12" pageNumber="529">Aesalus</emphasis>
Fabricius, 1801
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFF6FFB38701BFA46D5F526E" authority="Hellwig, 1792" authorityName="Hellwig" authorityYear="1792" box="[535,845,1015,1041]" class="Insecta" family="Lucanidae" genus="Sinodendron" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB38701BFA46CB7526E" box="[535,677,1015,1041]" italics="true" pageId="12" pageNumber="529">Sinodendron</emphasis>
Hellwig, 1792
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFF6FFB3864ABFA46A92526E" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[860,1152,1015,1041]" class="Insecta" family="Lucanidae" genus="Ceruchus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB3864ABFA46DD5526E" box="[860,967,1015,1041]" italics="true" pageId="12" pageNumber="529">Ceruchus</emphasis>
MacLeay, 1819
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFF6FFB3819DBFA46B85526E" authority="LeConte, 1861" authorityName="LeConte" authorityYear="1861" box="[1163,1431,1015,1041]" class="Insecta" family="Lucanidae" genus="Nicagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB3819DBFA46AF5526F" box="[1163,1255,1015,1040]" italics="true" pageId="12" pageNumber="529">Nicagus</emphasis>
LeConte, 1861
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFF6FFB38581B8486FA8524A" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[151,442,1051,1077]" class="Insecta" family="Lucanidae" genus="Syndesus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB38581B8486EEC524A" box="[151,254,1051,1077]" italics="true" pageId="12" pageNumber="529">Syndesus</emphasis>
MacLeay, 1819
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFF6FFB384DEB8486D2A524A" authority="Westwood, 1838" authorityName="Westwood" authorityYear="1838" box="[456,824,1051,1077]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB384DEB8486C7C524A" box="[456,622,1051,1077]" italics="true" pageId="12" pageNumber="529">Ceratognathus</emphasis>
Westwood, 1838
</taxonomicName>
, and the subfamily
<taxonomicName id="4C6F4D2FFFF6FFB3810AB8486AA5524A" authorityName="Macleay" authorityYear="1819" box="[1052,1207,1051,1077]" class="Globothalamea" family="Orbitolinidae" kingdom="Chromista" order="Loftusiida" pageId="12" pageNumber="529" phylum="Foraminifera" rank="subFamily" subFamily="Lampriminae">Lampriminae</taxonomicName>
. However,
<taxonomicName id="4C6F4D2FFFF6FFB3802FB8486EC75227" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="12" pageNumber="529" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF6FFB3802FB8486EC75227" italics="true" pageId="12" pageNumber="529">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFF6FFB385F3B86C6F625227" bold="true" box="[229,368,1086,1113]" pageId="12" pageNumber="529">
<taxonomicNameLabel id="A22857C5FFF6FFB385F3B86C6F625227" box="[229,368,1086,1113]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
shows only minute inner tubercle between apical teeth and mola, without distinct excision in front of the tubercle, which cannot be distinguished from that of
<taxonomicName id="4C6F4D2FFFF6FFB3867AB8306B345202" authority="Araya &amp; Yoshitomi, 2003" authorityName="Araya &amp; Yoshitomi" authorityYear="2003" box="[876,1318,1123,1149]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B91BEABEFFF6FFB3867AB8306A125203" box="[876,1024,1123,1149]" italics="true" pageId="12" pageNumber="529">Aesalus satoi</emphasis>
Araya &amp; Yoshitomi, 2003
</taxonomicName>
(
<bibRefCitation id="EFFE4B5DFFF6FFB38023B8306EEA52DE" author="Huang, H. &amp; Bi, W. - X. &amp; Li, L. - Z." pageId="12" pageNumber="529" pagination="18 - 42" refId="ref9790" refString="Huang, H., Bi, W. - X. &amp; Li, L. - Z. (2009) Discovery of a second species of Aesalini from continental China, with description of the new species and its third instar larva (Coleoptera: Scarabaeoidea: Lucanidae). Zootaxa, 2069, 18 - 42. https: // doi. org / 10.11646 / zootaxa. 2069.1.2" type="journal article" year="2009">
Huang
<emphasis id="B91BEABEFFF6FFB3809EB8306EA652DE" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
2009
</bibRefCitation>
). Therefore, the presence or absence of inner teeth should not be used as a major character in classification of larval
<taxonomicName id="4C6F4D2FFFF6FFB385EAB8F86F6652BA" authorityName="Latreille" authorityYear="1804" box="[252,372,1195,1221]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
.
</paragraph>
<paragraph id="8BD036ACFFF6FFB385D1B89C6D275076" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
<taxonomicName id="4C6F4D2FFFF6FFB385D1B89C6F655297" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[199,375,1231,1256]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="12" pageNumber="529" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF6FFB385D1B89C6F655297" box="[199,375,1231,1256]" italics="true" pageId="12" pageNumber="529">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFF6FFB3849AB89C6C0F5297" bold="true" box="[396,541,1230,1257]" pageId="12" pageNumber="529">
<taxonomicNameLabel id="A22857C5FFF6FFB3849AB89C6C0F5297" box="[396,541,1230,1257]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
larva possesses the following distinct differences from
<taxonomicName id="4C6F4D2FFFF6FFB381D3B89C6F4D5372" authority="(Boisduval, 1835)" baseAuthorityName="Boisduval" baseAuthorityYear="1835" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="B91BEABEFFF6FFB381D3B89C6B8E5296" box="[1221,1436,1231,1257]" italics="true" pageId="12" pageNumber="529">Figulus foveicollis</emphasis>
(Boisduval, 1835)
</taxonomicName>
(larva described in
<bibRefCitation id="EFFE4B5DFFF6FFB38720B8A06CD65372" author="Emden, F." box="[566,708,1266,1293]" pageId="12" pageNumber="529" pagination="301 - 310" refId="ref9525" refString="Emden, F. (1951) The larvae of Dendezia and Figulus, with notes on some other larvae of Lucanidae. Revue de Zoologie et de Botanique Africaines, 46, 301 - 310." type="journal article" year="1951">Emden 1951</bibRefCitation>
) and
<taxonomicName id="4C6F4D2FFFF6FFB387E9B8A06A455372" authority="Westwood, 1855" authorityName="Westwood" authorityYear="1855" box="[767,1111,1266,1293]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="lilliputanus">
<emphasis id="B91BEABEFFF6FFB387E9B8A06D8B5372" box="[767,921,1267,1293]" italics="true" pageId="12" pageNumber="529">F. lilliputanus</emphasis>
Westwood, 1855
</taxonomicName>
(larva described in
<bibRefCitation id="EFFE4B5DFFF6FFB38038B8A06EC6534E" author="Lawrence, J. F." pageId="12" pageNumber="529" pagination="213 - 219" refId="ref9936" refString="Lawrence, J. F. (1981) Notes on larval Lucanidae (Coleoptera). Journal of the Australian Entomological Society, 20, 213 - 219. https: // doi. org / 10.1111 / j. 1440 - 6055.1981. tb 01035. x" type="journal article" year="1981">Lawrence 1981</bibRefCitation>
): antenna 3-segmented with basal antennomere subdivided by an unpigmented ring near base, not distinctly 4-segmented; left mandible without large inner tooth; haptomerum with very few sensory spots but without setae; chaetoparia with markedly sparser setae; haptolachus with left nesia present but with no right nesia; epitorma pointed, not obsolete; sternite 10 with distinct raster which is interrupted on median line by a smooth longitudinal strip; plectrum being a single row of the conjoined tubercles; par stridens with most part of main row of tubercles conjoined (as in
<taxonomicName id="4C6F4D2FFFF6FFB3844FB9986FD4539A" authorityName="Westwood" authorityYear="1834" box="[345,454,1483,1509]" class="Insecta" family="Lucanidae" genus="Cardanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB3844FB9986FD4539A" box="[345,454,1483,1509]" italics="true" pageId="12" pageNumber="529">Cardanus</emphasis>
</taxonomicName>
), not separated, and without granules outside of main row; the two setae of the claw inserted side by side (as in
<taxonomicName id="4C6F4D2FFFF6FFB384D0B9BC6C375076" authorityName="MacLeay" authorityYear="1819" box="[454,549,1519,1545]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB384D0B9BC6C375076" box="[454,549,1519,1545]" italics="true" pageId="12" pageNumber="529">Nigidius</emphasis>
</taxonomicName>
), not behind each other.
</paragraph>
<paragraph id="8BD036ACFFF6FFB385D1BA406B30509E" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
<taxonomicName id="4C6F4D2FFFF6FFB385D1BA406F625053" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[199,368,1555,1580]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="12" pageNumber="529" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF6FFB385D1BA406F625053" box="[199,368,1555,1580]" italics="true" pageId="12" pageNumber="529">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFF6FFB38468BA406C155053" bold="true" box="[382,519,1554,1581]" pageId="12" pageNumber="529">
<taxonomicNameLabel id="A22857C5FFF6FFB38468BA406C155053" box="[382,519,1554,1581]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
larva differs from the one of
<taxonomicName id="4C6F4D2FFFF6FFB38646BA406AD75052" authority="Harold, 1878" authorityName="Harold" authorityYear="1878" box="[848,1221,1555,1581]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="perforatus">
<emphasis id="B91BEABEFFF6FFB38646BA406A395052" box="[848,1067,1555,1581]" italics="true" pageId="12" pageNumber="529">Nigidius perforatus</emphasis>
Harold, 1878
</taxonomicName>
(larva described in
<bibRefCitation id="EFFE4B5DFFF6FFB38581BA646F35502E" author="Emden, F." box="[151,295,1590,1617]" pageId="12" pageNumber="529" pagination="178 - 200" refId="ref9485" refString="Emden, F. (1935) Die Gattungsunterschiede der Hirschkaferlarven, ein Beitrag zum naturlichen System der Familie (Col. Lucan.). Stettiner Entomologische Zeitung, 93, 178 - 200. [in Germany]" type="journal article" year="1935">Emden 1935</bibRefCitation>
,
<bibRefCitation id="EFFE4B5DFFF6FFB38426BA656F78502E" author="Emden, F." box="[304,362,1590,1617]" pageId="12" pageNumber="529" pagination="301 - 310" refId="ref9525" refString="Emden, F. (1951) The larvae of Dendezia and Figulus, with notes on some other larvae of Lucanidae. Revue de Zoologie et de Botanique Africaines, 46, 301 - 310." type="journal article" year="1951">1951</bibRefCitation>
, but with characters of antenna, epipharynx and mouthparts not clarified) by the following points: left mandible without large inner tooth; tergites 7–10 (as in
<taxonomicName id="4C6F4D2FFFF6FFB3865FBA086D8D500A" authorityName="MacLeay" authorityYear="1819" box="[841,927,1627,1653]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB3865FBA086D8D500A" box="[841,927,1627,1653]" italics="true" pageId="12" pageNumber="529">Figulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D2FFFF6FFB386CFBA086A54500A" authorityName="Westwood" authorityYear="1834" box="[985,1094,1627,1653]" class="Insecta" family="Lucanidae" genus="Cardanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB386CFBA086A54500A" box="[985,1094,1627,1653]" italics="true" pageId="12" pageNumber="529">Cardanus</emphasis>
</taxonomicName>
) without spinules; sternite 10 with raster interrupted on median line by a smooth longitudinal strip; plectrum being a single row of the conjoined tubercles; par stridens with most part of main row of tubercles conjoined, not separated, and without granules outside of main row; claw with a subapical sinuation (as in
<taxonomicName id="4C6F4D2FFFF6FFB38624BA946D9A509E" authorityName="MacLeay" authorityYear="1819" box="[818,904,1735,1761]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB38624BA946D9A509E" box="[818,904,1735,1761]" italics="true" pageId="12" pageNumber="529">Figulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D2FFFF6FFB386A8BA946A39509E" authorityName="Westwood" authorityYear="1834" box="[958,1067,1735,1761]" class="Insecta" family="Lucanidae" genus="Cardanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB386A8BA946A39509E" box="[958,1067,1735,1761]" italics="true" pageId="12" pageNumber="529">Cardanus</emphasis>
</taxonomicName>
), not evenly tapering.
</paragraph>
<paragraph id="8BD036ACFFF6FFB385D1BAB86D2751C6" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="529">
<taxonomicName id="4C6F4D2FFFF6FFB385D1BAB86F62517B" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[199,368,1771,1796]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="12" pageNumber="529" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF6FFB385D1BAB86F62517B" box="[199,368,1771,1796]" italics="true" pageId="12" pageNumber="529">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFF6FFB38469BAB86C1B517B" bold="true" box="[383,521,1770,1797]" pageId="12" pageNumber="529">
<taxonomicNameLabel id="A22857C5FFF6FFB38469BAB86C1B517B" box="[383,521,1770,1797]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
larva is different from the one of
<taxonomicName id="4C6F4D2FFFF6FFB3869FBAB86A0D517A" box="[905,1055,1771,1797]" class="Insecta" family="Lucanidae" genus="Cardanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91BEABEFFF6FFB3869FBAB86DE4517A" box="[905,1014,1771,1797]" italics="true" pageId="12" pageNumber="529">Cardanus</emphasis>
sp.
</taxonomicName>
(larva described in
<bibRefCitation id="EFFE4B5DFFF6FFB381E9BAB86B8A517A" author="Emden, F." box="[1279,1432,1770,1797]" pageId="12" pageNumber="529" pagination="178 - 200" refId="ref9485" refString="Emden, F. (1935) Die Gattungsunterschiede der Hirschkaferlarven, ein Beitrag zum naturlichen System der Familie (Col. Lucan.). Stettiner Entomologische Zeitung, 93, 178 - 200. [in Germany]" type="journal article" year="1935">Emden, 1935</bibRefCitation>
,
<bibRefCitation id="EFFE4B5DFFF6FFB38581BB5D6EC35156" author="Emden, F." box="[151,209,1806,1833]" pageId="12" pageNumber="529" pagination="301 - 310" refId="ref9525" refString="Emden, F. (1951) The larvae of Dendezia and Figulus, with notes on some other larvae of Lucanidae. Revue de Zoologie et de Botanique Africaines, 46, 301 - 310." type="journal article" year="1951">1951</bibRefCitation>
, but with characters of antenna, epipharynx and mouthparts not clarified) by the following points: left mandible without large inner tooth; sternite 10 with distinct raster which is interrupted on median line by a smooth longitudinal strip; tibiotarsus about twice as long as wide (as in
<taxonomicName id="4C6F4D2FFFF6FFB387F0BB046D2E510E" authorityName="MacLeay" authorityYear="1819" box="[742,828,1879,1905]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB387F0BB046D2E510E" box="[742,828,1879,1905]" italics="true" pageId="12" pageNumber="529">Figulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D2FFFF6FFB38660BB046DC7510E" authorityName="MacLeay" authorityYear="1819" box="[886,981,1879,1905]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB38660BB046DC7510E" box="[886,981,1879,1905]" italics="true" pageId="12" pageNumber="529">Nigidius</emphasis>
</taxonomicName>
), not extremely short; plectrum being a single row of the conjoined tubercles; par stridens without granules outside of main row; the two setae of the claw inserted side by side (as in
<taxonomicName id="4C6F4D2FFFF6FFB384D0BBCC6C3751C6" authorityName="MacLeay" authorityYear="1819" box="[454,549,1951,1977]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFF6FFB384D0BBCC6C3751C6" box="[454,549,1951,1977]" italics="true" pageId="12" pageNumber="529">Nigidius</emphasis>
</taxonomicName>
), not behind each other.
</paragraph>
<caption id="DF106624FFF7FFB28581BBCB6D9D51CF" ID-DOI="http://doi.org/10.5281/zenodo.6375339" ID-Zenodo-Dep="6375339" box="[151,911,1944,1968]" httpUri="https://zenodo.org/record/6375339/files/figure.png" pageId="13" pageNumber="530" startId="13.[151,250,1944,1968]" targetBox="[151,1436,181,1948]" targetPageId="13">
<paragraph id="8BD036ACFFF7FFB28581BBCB6D9D51CF" blockId="13.[151,911,1944,1968]" box="[151,911,1944,1968]" pageId="13" pageNumber="530">
<emphasis id="B91BEABEFFF7FFB28581BBCB6F3251CF" bold="true" box="[151,288,1944,1968]" pageId="13" pageNumber="530">FIGURE 40.</emphasis>
<taxonomicName id="4C6F4D2FFFF7FFB28430BBCA6C7351CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[294,609,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="13" pageNumber="530" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF7FFB28430BBCA6FAC51CF" box="[294,446,1945,1968]" italics="true" pageId="13" pageNumber="530">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF7FFB2877DBBCB6CFA51CF" box="[619,744,1944,1968]" pageId="13" pageNumber="530" rank="species">new species</taxonomicNameLabel>
, tarsi and claws.
</paragraph>
</caption>
<caption id="DF106624FFF7FFB28581BBEF6DDD51AB" ID-DOI="http://doi.org/10.5281/zenodo.6377198" ID-Zenodo-Dep="6377198" box="[151,975,1980,2004]" httpUri="https://zenodo.org/record/6377198/files/figure.png" pageId="13" pageNumber="530" startId="13.[151,264,1980,2004]" targetBox="[151,1436,181,1948]" targetPageId="13">
<paragraph id="8BD036ACFFF7FFB28581BBEF6DDD51AB" blockId="13.[151,975,1980,2004]" box="[151,975,1980,2004]" pageId="13" pageNumber="530">
<emphasis id="B91BEABEFFF7FFB28581BBEF6F4651AB" bold="true" box="[151,340,1980,2004]" pageId="13" pageNumber="530">FIGURES 41–42.</emphasis>
<taxonomicName id="4C6F4D2FFFF7FFB2844CBBEE6C8751AB" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[346,661,1980,2004]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="13" pageNumber="530" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF7FFB2844CBBEE6FE151AB" box="[346,499,1981,2004]" italics="true" pageId="13" pageNumber="530">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF7FFB28789BBEF6D0E51AB" box="[671,796,1980,2004]" pageId="13" pageNumber="530" rank="species">new species</taxonomicNameLabel>
, female genitalia.
</paragraph>
</caption>
<caption id="DF106624FFF4FFB18581BB9B6A26519F" ID-DOI="http://doi.org/10.5281/zenodo.6375343" ID-Zenodo-Dep="6375343" box="[151,1076,1992,2017]" httpUri="https://zenodo.org/record/6375343/files/figure.png" pageId="14" pageNumber="531" startId="14.[151,250,1992,2016]" targetBox="[151,1435,181,1969]" targetPageId="14">
<paragraph id="8BD036ACFFF4FFB18581BB9B6A26519F" blockId="14.[151,1076,1992,2017]" box="[151,1076,1992,2017]" pageId="14" pageNumber="531">
<emphasis id="B91BEABEFFF4FFB18581BB9B6F32519F" bold="true" box="[151,288,1992,2016]" pageId="14" pageNumber="531">FIGURE 43.</emphasis>
<taxonomicName id="4C6F4D2FFFF4FFB18430BB9A6C73519F" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[294,609,1992,2017]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="14" pageNumber="531" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF4FFB18430BB9A6FAC519F" box="[294,446,1993,2016]" italics="true" pageId="14" pageNumber="531">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF4FFB1877DBB9A6CFA519F" box="[619,744,1992,2016]" pageId="14" pageNumber="531" rank="species">new species</taxonomicNameLabel>
, male genitalia (M1—holotype).
</paragraph>
</caption>
<caption id="DF106624FFF5FFB08581BBC46EDC5187" ID-DOI="http://doi.org/10.5281/zenodo.6375345" ID-Zenodo-Dep="6375345" httpUri="https://zenodo.org/record/6375345/files/figure.png" pageId="15" pageNumber="532" startId="15.[151,264,1943,1967]" targetBox="[194,1393,181,1920]" targetPageId="15">
<paragraph id="8BD036ACFFF5FFB08581BBC46EDC5187" blockId="15.[151,1437,1943,2040]" pageId="15" pageNumber="532">
<emphasis id="B91BEABEFFF5FFB08581BBC46F4751D0" bold="true" box="[151,341,1943,1967]" pageId="15" pageNumber="532">FIGURES 44–48.</emphasis>
<taxonomicName id="4C6F4D2FFFF5FFB0844ABBCB6C8851CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[348,666,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="15" pageNumber="532" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFF5FFB0844ABBCB6FE751D0" box="[348,501,1944,1967]" italics="true" pageId="15" pageNumber="532">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFF5FFB087B3BBCB6D3151CF" box="[677,803,1944,1968]" pageId="15" pageNumber="532" rank="species">new species</taxonomicNameLabel>
, larva and its characters. 44, larva in lateral view; 45, larva in dorsal view; 46, tenth abdominal segment in ventral view to show raster; 47, head in frontal view; 48, left antenna in lateral view.
</paragraph>
</caption>
<caption id="DF106624FFEAFFAF8581BBC46B0651AB" ID-DOI="http://doi.org/10.5281/zenodo.6375349" ID-Zenodo-Dep="6375349" httpUri="https://zenodo.org/record/6375349/files/figure.png" pageId="16" pageNumber="533" startId="16.[151,264,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="16">
<paragraph id="8BD036ACFFEAFFAF8581BBC46B0651AB" blockId="16.[151,1436,1943,2004]" pageId="16" pageNumber="533">
<emphasis id="B91BEABEFFEAFFAF8581BBC46F4151D0" bold="true" box="[151,339,1943,1967]" pageId="16" pageNumber="533">FIGURES 49–52.</emphasis>
<taxonomicName id="4C6F4D2FFFEAFFAF844EBBCB6C8251CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[344,656,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="16" pageNumber="533" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEAFFAF844EBBCB6FE251D0" box="[344,496,1944,1967]" italics="true" pageId="16" pageNumber="533">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFEAFFAF878FBBCB6D0651CF" box="[665,788,1944,1968]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
, larva legs. 49, foreleg in lateral view; 50, midleg in lateral view, with pars stridens enlarged; 51, right hindleg in lateral view, with plectrum enlarged; 52, plectrum of left hindleg.
</paragraph>
</caption>
<caption id="DF106624FFEBFFAE8581BBC46C585187" ID-DOI="http://doi.org/10.5281/zenodo.6375351" ID-Zenodo-Dep="6375351" httpUri="https://zenodo.org/record/6375351/files/figure.png" pageId="17" pageNumber="534" startId="17.[151,264,1943,1967]" targetBox="[194,1393,181,1920]" targetPageId="17">
<paragraph id="8BD036ACFFEBFFAE8581BBC46C585187" blockId="17.[151,1436,1943,2040]" pageId="17" pageNumber="534">
<emphasis id="B91BEABEFFEBFFAE8581BBC46F4551D0" bold="true" box="[151,343,1943,1967]" pageId="17" pageNumber="534">FIGURES 53–57.</emphasis>
<taxonomicName id="4C6F4D2FFFEBFFAE8476BBCB6CB451CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[352,678,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="17" pageNumber="534" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEBFFAE8476BBCB6FE951D0" box="[352,507,1944,1967]" italics="true" pageId="17" pageNumber="534">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFEBFFAE87A5BBCB6D2151CF" box="[691,819,1944,1968]" pageId="17" pageNumber="534" rank="species">new species</taxonomicNameLabel>
, larva characters. 53, left antenna with apical antennomere enlarged; 54, right antenna with apical antennomere enlarged; 55, mandibles in dorsal, inner lateral and ventral views; 56, epipharynx in ventral view; 57, mouthpart.
</paragraph>
</caption>
<caption id="DF106624FFE8FFAD8581BB346A9F5100" ID-DOI="http://doi.org/10.5281/zenodo.6375353" ID-Zenodo-Dep="6375353" box="[151,1165,1895,1920]" httpUri="https://zenodo.org/record/6375353/files/figure.png" pageId="18" pageNumber="535" startId="18.[151,250,1895,1919]" targetBox="[151,1435,181,1871]" targetPageId="18">
<paragraph id="8BD036ACFFE8FFAD8581BB346A9F5100" blockId="18.[151,1165,1895,1920]" box="[151,1165,1895,1920]" pageId="18" pageNumber="535">
<emphasis id="B91BEABEFFE8FFAD8581BB346F325100" bold="true" box="[151,288,1895,1919]" pageId="18" pageNumber="535">FIGURE 58.</emphasis>
<taxonomicName id="4C6F4D2FFFE8FFAD8430BB3B6C735100" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[294,609,1895,1920]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="18" pageNumber="535" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFE8FFAD8430BB3B6FAC5100" box="[294,446,1896,1919]" italics="true" pageId="18" pageNumber="535">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFE8FFAD877DBB3B6CFA5100" box="[619,744,1895,1919]" pageId="18" pageNumber="535" rank="species">new species</taxonomicNameLabel>
, pupa in dorsal, lateral and ventral views.
</paragraph>
</caption>
<caption id="DF106624FFE9FFAC8581BBCA6D6E5186" ID-DOI="http://doi.org/10.5281/zenodo.6375356" ID-Zenodo-Dep="6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="19" pageNumber="536" startId="19.[151,264,1945,1969]" targetBox="[177,1411,181,1922]" targetPageId="19">
<paragraph id="8BD036ACFFE9FFAC8581BBCA6D6E5186" blockId="19.[151,1436,1945,2042]" pageId="19" pageNumber="536">
<emphasis id="B91BEABEFFE9FFAC8581BBCA6F4151CD" bold="true" box="[151,339,1945,1970]" pageId="19" pageNumber="536">FIGURES 59–66.</emphasis>
<taxonomicName id="4C6F4D2FFFE9FFAC844EBBC96C9C51CD" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[344,654,1946,1970]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="19" pageNumber="536" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFE9FFAC844EBBC96FFD51CE" box="[344,495,1946,1969]" italics="true" pageId="19" pageNumber="536">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFE9FFAC8781BBC96D0151CD" box="[663,787,1946,1970]" pageId="19" pageNumber="536" rank="species">new species</taxonomicNameLabel>
, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of
<taxonomicName id="4C6F4D2FFFE9FFAC8054BBED6EDD5186" authorityName="Huang &amp; Chen" authorityYear="2022" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="19" pageNumber="536" phylum="Foraminifera" rank="species" species="qiuae">
<emphasis id="B91BEABEFFE9FFAC8054BBED6EDD5186" italics="true" pageId="19" pageNumber="536">Xizangia qiuae</emphasis>
</taxonomicName>
in cocoon; 65, living adult of
<taxonomicName id="4C6F4D2FFFE9FFAC8714BBB16C895186" authorityName="Huang &amp; Chen" authorityYear="2022" box="[514,667,2018,2041]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="19" pageNumber="536" phylum="Foraminifera" rank="species" species="qiuae">
<emphasis id="B91BEABEFFE9FFAC8714BBB16C895186" box="[514,667,2018,2041]" italics="true" pageId="19" pageNumber="536">Xizangia qiuae</emphasis>
</taxonomicName>
found below ant nest.
</paragraph>
</caption>
<caption id="DF106624FFEEFFAB8581BBC46B8551CF" ID-DOI="http://doi.org/10.5281/zenodo.6375358" ID-Zenodo-Dep="6375358" box="[151,1431,1943,1968]" httpUri="https://zenodo.org/record/6375358/files/figure.png" pageId="20" pageNumber="537" startId="20.[151,250,1943,1967]" targetBox="[237,1356,203,1897]" targetPageId="20">
<paragraph id="8BD036ACFFEEFFAB8581BBC46B8551CF" blockId="20.[151,1431,1943,1968]" box="[151,1431,1943,1968]" pageId="20" pageNumber="537">
<emphasis id="B91BEABEFFEEFFAB8581BBC46F3251D0" bold="true" box="[151,288,1943,1967]" pageId="20" pageNumber="537">FIGURE 67.</emphasis>
Habitus and characters of
<taxonomicName id="4C6F4D2FFFEEFFAB8739BBCB6D1A51CF" box="[559,776,1944,1968]" pageId="20" pageNumber="537">
<emphasis id="B91BEABEFFEEFFAB8739BBCB6CAB51CF" box="[559,697,1944,1968]" italics="true" pageId="20" pageNumber="537">Dolichoderus</emphasis>
species
</taxonomicName>
worker living with
<taxonomicName id="4C6F4D2FFFEEFFAB86C7BBCB6B1E51CF" authority="Huang &amp; Chen" authorityName="Huang &amp; Chen" authorityYear="2022" box="[977,1292,1944,1968]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="20" pageNumber="537" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEEFFAB86C7BBCB6A7851D0" box="[977,1130,1944,1967]" italics="true" pageId="20" pageNumber="537">Xizangia qiuae</emphasis>
Huang &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A22857C5FFEEFFAB8000BBCB6B8151CF" box="[1302,1427,1944,1968]" pageId="20" pageNumber="537" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BD036ACFFEFFFAA85D1BCC46FFF573E" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="538">
As a conclusion,
<taxonomicName id="4C6F4D2FFFEFFFAA849FBCC46FFF56CF" authorityName="Zhang" authorityYear="1988" box="[393,493,151,176]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="genus">
<emphasis id="B91BEABEFFEFFFAA849FBCC46FFF56CF" box="[393,493,151,176]" italics="true" pageId="21" pageNumber="538">Xizangia</emphasis>
</taxonomicName>
differs from
<emphasis id="B91BEABEFFEFFFAA8797BCC46D4356CE" box="[641,849,151,177]" italics="true" pageId="21" pageNumber="538">
<taxonomicName id="4C6F4D2FFFEFFFAA8797BCC46CCB56CE" authorityName="MacLeay" authorityYear="1819" box="[641,729,151,177]" class="Insecta" family="Lucanidae" genus="Figulus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="genus">Figulus</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFEFFFAA87F2BCC46D4356CE" authorityName="Westwood" authorityYear="1834" box="[740,849,151,177]" class="Insecta" family="Lucanidae" genus="Cardanus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="genus">Cardanus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6F4D2FFFEFFFAA869EBCC46DF556CE" authorityName="MacLeay" authorityYear="1819" box="[904,999,151,177]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFEFFFAA869EBCC46DF556CE" box="[904,999,151,177]" italics="true" pageId="21" pageNumber="538">Nigidius</emphasis>
</taxonomicName>
by having left mandible without large inner tooth, sternite 10 with raster interrupted on median line by a smooth longitudinal strip, plectrum being a single row of the conjoined tubercles, and par stridens without granules outside of main row. This may indicate that
<taxonomicName id="4C6F4D2FFFEFFFAA8581BD506F475762" box="[151,341,259,285]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="519" phylum="Arthropoda" rank="tribe" tribe="Penichrolucanini">Penichrolucanini</taxonomicName>
is separable from Figulini in systematics also by larval characters, but more species of both tribes need to be examined in future.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFEFFFAA85D1BD186FFE571A" box="[199,492,330,357]" pageId="21" pageNumber="538" type="description">
<paragraph id="8BD036ACFFEFFFAA85D1BD186FFE571A" blockId="21.[151,1437,151,897]" box="[199,492,330,357]" pageId="21" pageNumber="538">
<emphasis id="B91BEABEFFEFFFAA85D1BD186F17571A" bold="true" box="[199,261,331,357]" pageId="21" pageNumber="538">Pupa</emphasis>
(
<figureCitation id="13542A29FFEFFFAA8403BD186F74571A" box="[277,358,330,357]" captionStart="FIGURE 58" captionStartId="18.[151,250,1895,1919]" captionTargetBox="[151,1435,181,1871]" captionTargetId="figure-21@18.[151,1435,181,1871]" captionTargetPageId="18" captionText="FIGURE 58. Xizangia qiuae Huang &amp; Chen, new species, pupa in dorsal, lateral and ventral views." figureDoi="http://doi.org/10.5281/zenodo.6375353" httpUri="https://zenodo.org/record/6375353/files/figure.png" pageId="21" pageNumber="538">Fig. 58</figureCitation>
) as figured.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFEFFFAA85D1BD3D6C4B548E" pageId="21" pageNumber="538" type="biology_ecology">
<paragraph id="8BD036ACFFEFFFAA85D1BD3D6C4B548E" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="538">
<emphasis id="B91BEABEFFEFFFAA85D1BD3D6F9157F6" bold="true" box="[199,387,366,393]" pageId="21" pageNumber="538">Biological notes</emphasis>
(
<figureCitation id="13542A29FFEFFFAA8484BD3C6C0E57F6" box="[402,540,366,393]" captionStart-0="FIGURES 59–66" captionStart-1="FIGURE 67" captionStartId-0="19.[151,264,1945,1969]" captionStartId-1="20.[151,250,1943,1967]" captionTargetBox-0="[177,1411,181,1922]" captionTargetBox-1="[237,1356,203,1897]" captionTargetId-0="figure-19@19.[177,1411,181,1922]" captionTargetId-1="figure-21@20.[225,1362,181,1920]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." captionText-1="FIGURE 67. Habitus and characters of Dolichoderus species worker living with Xizangia qiuae Huang &amp; Chen, new species." figureDoi-0="http://doi.org/10.5281/zenodo.6375356" figureDoi-1="http://doi.org/10.5281/zenodo.6375358" httpUri-0="https://zenodo.org/record/6375356/files/figure.png" httpUri-1="https://zenodo.org/record/6375358/files/figure.png" pageId="21" pageNumber="538">Figs. 59–67</figureCitation>
). All adults and third instar larvae (
<figureCitation id="13542A29FFEFFFAA86B1BD3C6A3C57F5" box="[935,1070,367,394]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Figs. 64, 66</figureCitation>
) of
<taxonomicName id="4C6F4D2FFFEFFFAA814DBD3C6B1657F7" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[1115,1284,367,392]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEFFFAA814DBD3C6B1657F7" box="[1115,1284,367,392]" italics="true" pageId="21" pageNumber="538">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFEFFFAA8004BD3C6B8E57F7" bold="true" box="[1298,1436,366,393]" pageId="21" pageNumber="538">
<taxonomicNameLabel id="A22857C5FFEFFFAA8004BD3C6B8E57F7" box="[1298,1436,366,393]" pageId="21" pageNumber="538" rank="species">new species</taxonomicNameLabel>
</emphasis>
were found in a large fallen wood (
<figureCitation id="13542A29FFEFFFAA8738BDC06C9657D1" box="[558,644,403,430]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Fig. 62</figureCitation>
), immediately below the bottom of an ant nest. The fallen wood was found in a tea field (
<figureCitation id="13542A29FFEFFFAA849FBDE46FCF57AD" box="[393,477,439,466]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Fig. 60</figureCitation>
) in a tropical montane evergreen broad-leaved forest area (
<figureCitation id="13542A29FFEFFFAA8198BDE46AF057AE" box="[1166,1250,438,465]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Fig. 59</figureCitation>
) at
<quantity id="4C979B49FFEFFFAA8004BDE46B7657AE" box="[1298,1380,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="538" unit="m" value="1600.0">1600m</quantity>
. The ants were identified as
<taxonomicName id="4C6F4D2FFFEFFFAA84BCBD886C7F578A" box="[426,621,475,501]" class="Insecta" family="Formicidae" genus="Dolichoderus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91BEABEFFEFFFAA84BCBD886C50578A" box="[426,578,475,501]" italics="true" pageId="21" pageNumber="538">Dolichoderus</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13542A29FFEFFFAA8797BD886CC45789" box="[641,726,475,502]" captionStart="FIGURE 67" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[237,1356,203,1897]" captionTargetId="figure-21@20.[225,1362,181,1920]" captionTargetPageId="20" captionText="FIGURE 67. Habitus and characters of Dolichoderus species worker living with Xizangia qiuae Huang &amp; Chen, new species." figureDoi="http://doi.org/10.5281/zenodo.6375358" httpUri="https://zenodo.org/record/6375358/files/figure.png" pageId="21" pageNumber="538">Fig. 67</figureCitation>
) (
<taxonomicName id="4C6F4D2FFFEFFFAA87E7BD886D82578A" box="[753,912,475,501]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="4C6F4D2FFFEFFFAA8688BD886A30578A" box="[926,1058,475,501]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
:
<taxonomicName id="4C6F4D2FFFEFFFAA8127BD886AF05789" authorityName="Forel" authorityYear="1878" box="[1073,1250,475,502]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="549" phylum="Arthropoda" rank="subFamily" subFamily="Dolichoderinae">Dolichoderinae</taxonomicName>
), being close to
<taxonomicName id="4C6F4D2FFFEFFFAA8581BDAC6FEE5466" authority="Xu, 1995" authorityName="Xu" authorityYear="1995" box="[151,508,510,537]" class="Insecta" family="Formicidae" genus="Dolichoderus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B91BEABEFFEFFFAA8581BDAC6F955467" box="[151,391,511,537]" italics="true" pageId="21" pageNumber="538">Dolichoderus incisus</emphasis>
<bibRefCitation id="EFFE4B5DFFEFFFAA8487BDAC6FEE5466" author="Xu, Z. - H." box="[401,508,510,537]" pageId="21" pageNumber="538" pagination="33 - 39" refId="ref10435" refString="Xu, Z. - H. (1995) A taxonomic study of the ant genus Dolichoderus Lund in China. Journal of Southwest Forestry College, 15 (1), 33 - 39." type="journal article" year="1995">Xu, 1995</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54D4880EFFEFFFAA871BBDAC6C545466" box="[525,582,511,537]" pageId="21" pageNumber="538">Type</typeStatus>
locality: Lincang,
<collectingRegion id="49ABF84EFFEFFFAA8636BDAC6D695466" box="[800,891,511,537]" country="China" name="Yunnan" pageId="21" pageNumber="538">Yunnan</collectingRegion>
). The nest of the
<taxonomicName id="4C6F4D2FFFEFFFAA815EBDAC6AF25466" authorityName="Lund" authorityYear="1831" box="[1096,1248,511,537]" class="Insecta" family="Formicidae" genus="Dolichoderus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFEFFFAA815EBDAC6AF25466" box="[1096,1248,511,537]" italics="true" pageId="21" pageNumber="538">Dolichoderus</emphasis>
</taxonomicName>
ant was located at the center of the log, consisting of layers of shells; most parts of the log were not decayed. Adults (
<figureCitation id="13542A29FFEFFFAA8004BE706B765442" box="[1298,1380,546,573]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Fig. 65</figureCitation>
) and larvae (mostly in cocoon, see
<figureCitation id="13542A29FFEFFFAA84F6BE146C22541D" box="[480,560,583,610]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Fig. 66</figureCitation>
) of
<taxonomicName id="4C6F4D2FFFEFFFAA874DBE146D11541F" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[603,771,583,608]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEFFFAA874DBE146D11541F" box="[603,771,583,608]" italics="true" pageId="21" pageNumber="538">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFEFFFAA8606BE146D8A541F" bold="true" box="[784,920,582,609]" pageId="21" pageNumber="538">
<taxonomicNameLabel id="A22857C5FFEFFFAA8606BE146D8A541F" box="[784,920,582,609]" pageId="21" pageNumber="538" rank="species">new species</taxonomicNameLabel>
</emphasis>
were gathered in the decayed part of the wood just below the ant nest, not inside the ant nest. The ants (
<figureCitation id="13542A29FFEFFFAA860CBE386DB054F9" box="[794,930,619,646]" captionStart="FIGURES 59–66" captionStartId="19.[151,264,1945,1969]" captionTargetBox="[177,1411,181,1922]" captionTargetId="figure-19@19.[177,1411,181,1922]" captionTargetPageId="19" captionText="FIGURES 59–66. Xizangia qiuae Huang &amp; Chen, new species, biotope and field observations (photo by H. Xu). 59, biotope; 60 and 62, fallen wood in which the ant nest was found; 61 and 63, broken ant nest and ant workers; 64 and 66, larva of Xizangia qiuae in cocoon; 65, living adult of Xizangia qiuae found below ant nest." figureDoi="http://doi.org/10.5281/zenodo.6375356" httpUri="https://zenodo.org/record/6375356/files/figure.png" pageId="21" pageNumber="538">Figs. 61, 63</figureCitation>
) did not visit the part where
<taxonomicName id="4C6F4D2FFFEFFFAA81FABE386B8754FB" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[1260,1429,619,644]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEFFFAA81FABE386B8754FB" box="[1260,1429,619,644]" italics="true" pageId="21" pageNumber="538">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFEFFFAA8581BEDC6F3354D7" bold="true" box="[151,289,654,681]" pageId="21" pageNumber="538">
<taxonomicNameLabel id="A22857C5FFEFFFAA8581BEDC6F3354D7" box="[151,289,654,681]" pageId="21" pageNumber="538" rank="species">new species</taxonomicNameLabel>
</emphasis>
lived but would attack
<taxonomicName id="4C6F4D2FFFEFFFAA873ABEDC6CC454D7" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[556,726,655,680]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEFFFAA873ABEDC6CC454D7" box="[556,726,655,680]" italics="true" pageId="21" pageNumber="538">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFEFFFAA87F3BEDC6D7D54D7" bold="true" box="[741,879,654,681]" pageId="21" pageNumber="538">
<taxonomicNameLabel id="A22857C5FFEFFFAA87F3BEDC6D7D54D7" box="[741,879,654,681]" pageId="21" pageNumber="538" rank="species">new species</taxonomicNameLabel>
</emphasis>
if the wall between the ant nest and the
<taxonomicName id="4C6F4D2FFFEFFFAA802FBEDC6B8F54D7" authorityName="Zhang" authorityYear="1988" box="[1337,1437,655,680]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="genus">
<emphasis id="B91BEABEFFEFFFAA802FBEDC6B8F54D7" box="[1337,1437,655,680]" italics="true" pageId="21" pageNumber="538">Xizangia</emphasis>
</taxonomicName>
nest was broken. Also
<taxonomicName id="4C6F4D2FFFEFFFAA8487BEE06D4754B1" authority="Boileau, 1899" authorityName="Boileau" authorityYear="1899" box="[401,853,691,718]" class="Insecta" family="Lucanidae" genus="Aegus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="taurus">
<emphasis id="B91BEABEFFEFFFAA8487BEE06C3654B3" box="[401,548,691,717]" italics="true" pageId="21" pageNumber="538">Aegus taurus</emphasis>
Boileau, 1899 (Lucanidae)
</taxonomicName>
was found living together with
<taxonomicName id="4C6F4D2FFFEFFFAA81A0BEE06B4C54B3" authority="Huang &amp; Chen, 2022" authorityName="Huang &amp; Chen" authorityYear="2022" box="[1206,1374,691,716]" class="Globothalamea" family="Orbitolinidae" genus="Xizangia" kingdom="Chromista" order="Loftusiida" pageId="21" pageNumber="538" phylum="Foraminifera" rank="species" species="qiuae" status="sp. nov.">
<emphasis id="B91BEABEFFEFFFAA81A0BEE06B4C54B3" box="[1206,1374,691,716]" italics="true" pageId="21" pageNumber="538">Xizangia qiuae</emphasis>
</taxonomicName>
,
<emphasis id="B91BEABEFFEFFFAA807ABEE06EFB548F" bold="true" pageId="21" pageNumber="538">
<taxonomicNameLabel id="A22857C5FFEFFFAA807ABEE06EFB548F" pageId="21" pageNumber="538" rank="species">new species</taxonomicNameLabel>
</emphasis>
in the decayed part of the wood.
</paragraph>
</subSubSection>
<subSubSection id="C3756527FFEFFFAA85D1BEA86D0555FE" pageId="21" pageNumber="538" type="reference_group">
<paragraph id="8BD036ACFFEFFFAA85D1BEA86D0555FE" blockId="21.[151,1437,151,897]" pageId="21" pageNumber="538">
According to
<bibRefCitation id="EFFE4B5DFFEFFFAA8473BEA86C6C556A" author="Geiselhardt, S. F. &amp; Peschke, K. &amp; Nagel, P." box="[357,638,763,789]" pageId="21" pageNumber="538" pagination="871 - 894" refId="ref9562" refString="Geiselhardt, S. F., Peschke, K. &amp; Nagel, P. (2007) A review of myrmecophily in ant nest beetles (Coleoptera: Carabidae: Paussinae): linking early observations with recent findings. Naturwissenschaften, 94, 871 - 894. https: // doi. org / 10.1007 / s 00114 - 007 - 0271 - x" type="journal article" year="2007">
Geiselhardt
<emphasis id="B91BEABEFFEFFFAA84F8BEA86C3A556A" box="[494,552,763,789]" italics="true" pageId="21" pageNumber="538">et al.</emphasis>
(2007)
</bibRefCitation>
, the following
<taxonomicName id="4C6F4D2FFFEFFFAA863ABEA86D88556A" ID-CoL="8KTQL" box="[812,922,763,789]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="subFamily" subFamily="Paussinae">Paussinae</taxonomicName>
taxa have been recorded as the guests of the Dolichoderini ants (
<taxonomicName id="4C6F4D2FFFEFFFAA846BBF4C6C205545" ID-CoL="JGK" authorityName="Forel" authorityYear="1878" box="[381,562,799,826]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="549" phylum="Arthropoda" rank="subFamily" subFamily="Dolichoderinae">Dolichoderinae</taxonomicName>
):
<taxonomicName id="4C6F4D2FFFEFFFAA8751BF4C6DFE5546" ID-CoL="3MHM2" authority="Kolbe, 1920" authorityName="Kolbe" authorityYear="1920" box="[583,1004,799,825]" class="Insecta" family="Carabidae" genus="Homopterus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="steinbachi">
<emphasis id="B91BEABEFFEFFFAA8751BF4C6D425546" box="[583,848,799,825]" italics="true" pageId="21" pageNumber="538">Homopterus steinbachi</emphasis>
Kolbe, 1920
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFEFFFAA86EDBF4C6AA55546" box="[1019,1207,799,825]" class="Insecta" family="Carabidae" genus="Arthropterus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91BEABEFFEFFFAA86EDBF4C6A995546" box="[1019,1163,799,825]" italics="true" pageId="21" pageNumber="538">Arthropterus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C6F4D2FFFEFFFAA81D1BF4C6F3F5522" ID-CoL="7672Z" authority="Reiche, 1860" authorityName="Reiche" authorityYear="1860" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="thomsonii">
<emphasis id="B91BEABEFFEFFFAA81D1BF4C6B8E5546" box="[1223,1436,799,825]" italics="true" pageId="21" pageNumber="538">Paussus thomsonii</emphasis>
Reiche, 1860
</taxonomicName>
and
<taxonomicName id="4C6F4D2FFFEFFFAA8470BF106D075522" ID-CoL="766WX" authority="Westwood, 1838" authorityName="Westwood" authorityYear="1838" box="[358,789,835,861]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="cochlearius">
<emphasis id="B91BEABEFFEFFFAA8470BF106C5C5522" box="[358,590,835,861]" italics="true" pageId="21" pageNumber="538">Paussus cochlearius</emphasis>
Westwood, 1838
</taxonomicName>
; only
<taxonomicName id="4C6F4D2FFFEFFFAA864BBF106AE35522" ID-CoL="3MHM2" authority="Kolbe 1920" authorityName="Kolbe" authorityYear="1920" box="[861,1265,835,861]" class="Insecta" family="Carabidae" genus="Homopterus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="species" species="steinbachi">
<emphasis id="B91BEABEFFEFFFAA864BBF106A715522" box="[861,1123,835,861]" italics="true" pageId="21" pageNumber="538">Homopterus steinbachi</emphasis>
Kolbe
<quantity id="4C979B49FFEFFFAA81AFBF106AE35522" box="[1209,1265,835,861]" metricMagnitude="1" metricUnit="m" metricValue="4.8768" pageId="21" pageNumber="538" unit="in" value="1920.0">1920</quantity>
</taxonomicName>
in Neotropical Region is recorded as the guest of the
<taxonomicName id="4C6F4D2FFFEFFFAA8755BF346CC955FE" ID-CoL="62R6N" authorityName="Lund" authorityYear="1831" box="[579,731,871,897]" class="Insecta" family="Formicidae" genus="Dolichoderus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEABEFFEFFFAA8755BF346CC955FE" box="[579,731,871,897]" italics="true" pageId="21" pageNumber="538">Dolichoderus</emphasis>
</taxonomicName>
ants.
</paragraph>
</subSubSection>
</treatment>
</document>
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