248 lines
31 KiB
XML
248 lines
31 KiB
XML
<document id="869227CD331A9D361192D685AF859D5E" ID-CLB-Dataset="53817" ID-DOI="10.11646/zootaxa.5205.5.2" ID-GBIF-Dataset="0e38201e-1c55-425c-9f76-eda1695fef9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318477" ID-ZooBank="2D332120-D5B2-4B50-A7F2-DE0DE91249D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668421138976" checkinUser="plazi" docAuthor="Tuf, Ivan Hadrián, Barjadze, Shalva & Maghradze, Eter" docDate="2022" docId="03C687B72C46FFC3FF32FA50A9BBF849" docLanguage="en" docName="zootaxa.5205.5.2.pdf" docOrigin="Zootaxa 5205 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptops (Cryptops) datviensis Tuf & Barjadze & Maghradze 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="438" masterDocId="FFFFFFCF2C47FFC1FFA5FFD3A966FFF4" masterDocTitle="The first troglobiotic cryptopid centipede (Chilopoda: Scolopendromorpha: Cryptopidae) from the Caucasus" masterLastPageNumber="444" masterPageNumber="436" pageNumber="437" updateTime="1699394133656" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F06A57D69D5809FDA2E8741C934CC569">
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<mods:title id="E7B3ADCA3A29E928649C7B0C74BAE72C">The first troglobiotic cryptopid centipede (Chilopoda: Scolopendromorpha: Cryptopidae) from the Caucasus</mods:title>
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</mods:titleInfo>
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<mods:name id="A83EF18256D6D78D149F2221013D17FC" type="personal">
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<mods:role id="8EC5578FAF74D73F2C84C0B7B3A65391">
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<mods:roleTerm id="CC5BE67387FFF684031FD0AE13D87785">Author</mods:roleTerm>
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<mods:namePart id="467B262C3E2B79E18412E03EA2025C5C">Tuf, Ivan Hadrián</mods:namePart>
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<mods:affiliation id="21FA7954F5690179901538C4740C8833">Department of Ecology and Environmental Sciences, Faculty of Science, Palacký University Olomouc, Olomouc, Czech Republic Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Republic of Georgia</mods:affiliation>
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</mods:name>
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<mods:name id="592367E46C2A404AE866AA3133719273" type="personal">
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<mods:roleTerm id="83F7EA298C60F9A4CF4CCAE2C76F5ED1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E89C5C599F16F9FD8A0413A0414A72A3">Barjadze, Shalva</mods:namePart>
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<mods:nameIdentifier id="64D0CF1587D4455947B508E7F40664A2" type="ORCID">0000-0001-8992-4987</mods:nameIdentifier>
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<mods:affiliation id="75E014D4EDD6B99C4F765CFE31178BFD">shalva. barjadze @ yahoo. com; https: // orcid. org / 0000 - 0001 - 8992 - 4987</mods:affiliation>
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<mods:nameIdentifier id="EF76FBBD30EFFA56F81E55A0BB16D58E" type="email">shalva.barjadze@yahoo.com</mods:nameIdentifier>
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<mods:roleTerm id="50D328883BD3396ECF81E59CB277B80F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3C507B26282AA5C46028809AEC6108D9">Maghradze, Eter</mods:namePart>
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<mods:nameIdentifier id="C9A2EB992FD777C42C30944935CECCC0" type="ORCID">0000-0002-4796-9439</mods:nameIdentifier>
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<mods:affiliation id="B296A764DB42266F426C6DB61C01105D">eter. magradze. 1 @ iliauni. edu. ge; https: // orcid. org / 0000 - 0002 - 4796 - 9439</mods:affiliation>
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<mods:nameIdentifier id="218F9C4A9D3F2CD536D3C480F8EBFA29" type="email">eter.magradze.1@iliauni.edu.ge</mods:nameIdentifier>
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<mods:title id="3107CB9604239C0670D8657F7B08F750">Zootaxa</mods:title>
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<mods:part id="5ABADBE9E4B1BF15672E9906CAEB0BAB">
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<mods:date id="A07551AB110471C0BA9947C31744181F">2022</mods:date>
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<mods:number id="68A54BCF629DB68CBE8B2001FDB92071">2022-11-10</mods:number>
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<mods:number id="EDF7DE98B55CA2BCF0184F02E77610C3">5205</mods:number>
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<treatment id="03C687B72C46FFC3FF32FA50A9BBF849" ID-DOI="http://doi.org/10.5281/zenodo.7318475" ID-GBIF-Taxon="210361647" ID-Zenodo-Dep="7318475" LSID="urn:lsid:plazi:treatment:03C687B72C46FFC3FF32FA50A9BBF849" httpUri="http://treatment.plazi.org/id/03C687B72C46FFC3FF32FA50A9BBF849" lastPageId="2" lastPageNumber="438" pageId="1" pageNumber="437">
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<subSubSection id="C375652A2C46FFC0FF32FA50AB0AFA69" box="[151,620,1411,1438]" pageId="1" pageNumber="437" type="nomenclature">
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<paragraph id="8BD036A12C46FFC0FF32FA50AB0AFA69" blockId="1.[151,620,1411,1473]" box="[151,620,1411,1438]" pageId="1" pageNumber="437">
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<heading id="D09881CD2C46FFC0FF32FA50AB0AFA69" bold="true" box="[151,620,1411,1438]" fontSize="11" level="1" pageId="1" pageNumber="437" reason="1">
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<emphasis id="B91BEAB32C46FFC0FF32FA50AB0AFA69" bold="true" box="[151,620,1411,1438]" pageId="1" pageNumber="437">
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<taxonomicName id="4C6F4D222C46FFC0FF32FA50AB6EFA6A" authority="Tuf & Barjadze & Maghradze, 2022" authorityName="Tuf & Barjadze & Maghradze" authorityYear="2022" box="[151,520,1411,1438]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="437" phylum="Arthropoda" rank="species" species="datviensis" status="sp. nov." subGenus="Cryptops">
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<emphasis id="B91BEAB32C46FFC0FF32FA50A865FA69" bold="true" box="[151,259,1411,1437]" italics="true" pageId="1" pageNumber="437">Cryptops</emphasis>
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(
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||
<emphasis id="B91BEAB32C46FFC0FEB1FA50A8E6FA69" bold="true" box="[276,384,1411,1437]" italics="true" pageId="1" pageNumber="437">Cryptops</emphasis>
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)
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<emphasis id="B91BEAB32C46FFC0FE34FA50AB6EFA6A" bold="true" box="[401,520,1411,1438]" italics="true" pageId="1" pageNumber="437">datviensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22857C82C46FFC0FDAAFA50AB0AFA69" box="[527,620,1411,1437]" pageId="1" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C375652A2C46FFC0FF32FA74A856FA35" box="[151,304,1447,1473]" pageId="1" pageNumber="437" type="description">
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<paragraph id="8BD036A12C46FFC0FF32FA74A856FA35" blockId="1.[151,620,1411,1473]" box="[151,304,1447,1473]" pageId="1" pageNumber="437">
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<figureCitation id="13542A242C46FFC0FF32FA74A99DFA35" box="[151,251,1447,1473]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1618,1642]" captionTargetBox="[405,1184,193,1554]" captionTargetId="figure-19@3.[363,1224,181,1594]" captionTargetPageId="3" captionText="FIGURES 2–7. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 2–4. Cryptops datviensis sp. nov.: 2. Anterior part of body (holotype, IZISU TM-T-00001), dorsal view. 3. Head and forcipular segment (holotype, IZISU TM-T-00001), ventral view. 4. Clypeus and labrum (paratype, IZISU TM-T-00002), ventral view. 5–7. Cryptops caucasius Verhoeff, 1934: 5. Anterior part of body, dorsal view. 6. Head and forcipular segment ventral view. 7. Clypeus and labrum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318483" httpUri="https://zenodo.org/record/7318483/files/figure.png" pageId="1" pageNumber="437">Figs 2–4</figureCitation>
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,
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<figureCitation id="13542A242C46FFC0FEA3FA74A856FA35" box="[262,304,1447,1473]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[310,1271,422,1817]" captionTargetId="figure-380@4.[270,1318,374,1878]" captionTargetPageId="4" captionText="FIGURES 8–11. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 8, 9. Cryptops datviensis sp. nov.: 8. Posterior part of body (holotype, IZISU TM-T-00001), lateral view. 9. End of right ultimate leg (paratype, IZISU TM-T-00002), lateral view. 10, 11. Cryptops caucasius Verhoeff, 1934: 10. Posterior part of body, lateral view. 11. End of right ultimate leg, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7318485" httpUri="https://zenodo.org/record/7318485/files/figure.png" pageId="1" pageNumber="437">8, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375652A2C46FFC0FF32FA3CAAC4F96D" pageId="1" pageNumber="437" type="materials_examined">
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<paragraph id="8BD036A12C46FFC0FF32FA3CA85FF9A5" blockId="1.[151,1437,1519,2014]" pageId="1" pageNumber="437">
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<emphasis id="B91BEAB32C46FFC0FF32FA3CA8E4F9FD" bold="true" box="[151,386,1519,1545]" pageId="1" pageNumber="437">Material examined.</emphasis>
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<materialsCitation id="3B073CFC2C46FFC0FE2FFA3CA85FF9A5" ID-GBIF-Occurrence="4134607303" collectingDate="2020-08-08" collectionCode="IZISU" collectorName="Eter Maghradze" country="Georgia" county="Sataplia-Tskaltubo" latitude="42.374443" location="Datvi Cave" longLatPrecision="1" longitude="42.595833" municipality="Tskaltubo Municipality" pageId="1" pageNumber="437" specimenCode="TM-T-00001" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
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||
<emphasis id="B91BEAB32C46FFC0FE2FFA3CAB66F9FD" bold="true" box="[394,512,1519,1545]" pageId="1" pageNumber="437">
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<typeStatus id="54D488032C46FFC0FE2FFA3CA89DF9FD" box="[394,507,1519,1545]" pageId="1" pageNumber="437" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
<collectingCountry id="F37876312C46FFC0FDADFA3CAB03F9FD" box="[520,613,1519,1545]" name="Georgia" pageId="1" pageNumber="437">Georgia</collectingCountry>
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,
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<collectingRegion id="49ABF8432C46FFC0FDD4FA3CAA72F9FD" box="[625,788,1519,1545]" country="Georgia" name="Imereti" pageId="1" pageNumber="437">Imereti region</collectingRegion>
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,
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||
<collectingCounty id="62B14E2D2C46FFC0FC85FA3CAA94F9FD" box="[800,1010,1519,1545]" pageId="1" pageNumber="437">Sataplia-Tskaltubo</collectingCounty>
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||
karst massif,
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||
<collectingMunicipality id="6BB4ACDB2C46FFC0FB35FA3CACFEF9FD" box="[1168,1432,1519,1545]" pageId="1" pageNumber="437">Tskaltubo Municipality</collectingMunicipality>
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||
,
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<location id="8EB0607A2C46FFC0FF32F9C0A87EF9D9" LSID="urn:lsid:plazi:treatment:03C687B72C46FFC3FF32FA50A9BBF849:8EB0607A2C46FFC0FF32F9C0A87EF9D9" box="[151,280,1555,1581]" country="Georgia" county="Sataplia-Tskaltubo" latitude="42.374443" longLatPrecision="1" longitude="42.595833" municipality="Tskaltubo Municipality" name="Datvi Cave" pageId="1" pageNumber="437" stateProvince="Imereti">Datvi Cave</location>
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(
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<geoCoordinate id="EE5B50662C46FFC0FE8DF9C0A8A4F9D9" box="[296,450,1555,1581]" degrees="42" direction="north" minutes="22" orientation="latitude" pageId="1" pageNumber="437" precision="1" seconds="28.0" value="42.374443">42°22'28.0"N</geoCoordinate>
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<geoCoordinate id="EE5B50662C46FFC0FE6FF9C0AB02F9DA" box="[458,612,1555,1582]" degrees="42" direction="east" minutes="35" orientation="longitude" pageId="1" pageNumber="437" precision="1" seconds="45.0" value="42.595833">42°35'45.0"E</geoCoordinate>
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), dark zone of cave,
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||
<date id="FFD110612C46FFC0FCF7F9C0AA9AF9D9" box="[850,1020,1555,1581]" pageId="1" pageNumber="437" value="2020-08-08">
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<collectingDate id="EF95E9892C46FFC0FCF7F9C0AA9AF9D9" box="[850,1020,1555,1581]" pageId="1" pageNumber="437" value="2020-08-08">8 August 2020</collectingDate>
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</date>
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,
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<collectorName id="269A53772C46FFC0FBADF9C7ADDBF9D9" box="[1032,1213,1555,1582]" pageId="1" pageNumber="437">Eter Maghradze</collectorName>
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leg., 1 spm (
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||
<collectionCode id="ED7EAE642C46FFC0FAF6F9C0ACFAF9D9" box="[1363,1436,1555,1581]" pageId="1" pageNumber="437">IZISU</collectionCode>
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||
<specimenCode id="DBC99EDA2C46FFC0FF32F9E4A84BF9A5" box="[151,301,1591,1617]" collectionCode="TM-T-" pageId="1" pageNumber="437">TM-T-00001</specimenCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BD036A12C46FFC0FF62F988AAC4F96D" blockId="1.[151,1437,1519,2014]" pageId="1" pageNumber="437">
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<materialsCitation id="3B073CFC2C46FFC0FF62F988ACF1F981" ID-GBIF-Occurrence="4134607301" box="[199,1431,1627,1654]" collectingDate="2020-11-03" collectionCode="IZISU" collectorName="Eter Maghradze" pageId="1" pageNumber="437" specimenCode="TM-T-00002" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B91BEAB32C46FFC0FF62F988A859F981" bold="true" box="[199,319,1627,1653]" pageId="1" pageNumber="437">
|
||
<typeStatus id="54D488032C46FFC0FF62F988A859F981" box="[199,319,1627,1653]" pageId="1" pageNumber="437" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
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||
: same locality and collector: 1 spm (
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||
<collectionCode id="ED7EAE642C46FFC0FD7BF988AA41F981" box="[734,807,1627,1653]" pageId="1" pageNumber="437">IZISU</collectionCode>
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||
<specimenCode id="DBC99EDA2C46FFC0FC8BF988AAA2F981" box="[814,964,1627,1653]" collectionCode="TM-T-" pageId="1" pageNumber="437">TM-T-00002</specimenCode>
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||
),
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||
<date id="FFD110612C46FFC0FC72F988ADC3F981" box="[983,1189,1627,1653]" pageId="1" pageNumber="437" value="2020-11-03">
|
||
<collectingDate id="EF95E9892C46FFC0FC72F988ADC3F981" box="[983,1189,1627,1653]" pageId="1" pageNumber="437" value="2020-11-03">3 November 2020</collectingDate>
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||
</date>
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||
,
|
||
<collectorName id="269A53772C46FFC0FB15F98FAC02F981" box="[1200,1380,1627,1654]" pageId="1" pageNumber="437">Eter Maghradze</collectorName>
|
||
leg.
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B073CFC2C46FFC0FF32F9ACAAC4F96D" ID-GBIF-Occurrence="4134607302" box="[151,930,1663,1690]" collectingDate="2020-11-30" collectionCode="NHMW" collectorName="Eter Maghradze" pageId="1" pageNumber="437" specimenCode="MY10321" specimenCount="1" typeStatus="paratype">
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||
1 spm (
|
||
<collectionCode id="ED7EAE642C46FFC0FF4CF9ACA820F96D" box="[233,326,1663,1689]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="437" type="Museum">NHMW</collectionCode>
|
||
<specimenCode id="DBC99EDA2C46FFC0FEE8F9ACA8A3F96D" box="[333,453,1663,1689]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="437" type="Museum">MY10321</specimenCode>
|
||
),
|
||
<date id="FFD110612C46FFC0FE7DF9ACABD5F96D" box="[472,691,1663,1689]" pageId="1" pageNumber="437" value="2020-11-30">
|
||
<collectingDate id="EF95E9892C46FFC0FE7DF9ACABD5F96D" box="[472,691,1663,1689]" pageId="1" pageNumber="437" value="2020-11-30">30 November 2020</collectingDate>
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||
</date>
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||
,
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<collectorName id="269A53772C46FFC0FD1BF953AA14F96D" box="[702,882,1663,1690]" pageId="1" pageNumber="437">Eter Maghradze</collectorName>
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leg.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C375652A2C46FFC0FF62F970A8D5F8DD" pageId="1" pageNumber="437" type="diagnosis">
|
||
<paragraph id="8BD036A12C46FFC0FF62F970A8D5F8DD" blockId="1.[151,1437,1519,2014]" pageId="1" pageNumber="437">
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||
<emphasis id="B91BEAB32C46FFC0FF62F970A824F949" bold="true" box="[199,322,1699,1725]" pageId="1" pageNumber="437">Diagnosis.</emphasis>
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A
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<taxonomicName id="4C6F4D222C46FFC0FEF9F977AB7AF949" box="[348,540,1699,1726]" pageId="1" pageNumber="437">
|
||
<emphasis id="B91BEAB32C46FFC0FEF9F977A8A5F94A" box="[348,451,1700,1726]" italics="true" pageId="1" pageNumber="437">Cryptops</emphasis>
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species
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</taxonomicName>
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, about
|
||
<quantity id="4C979B442C46FFC0FDC2F970ABFAF94A" box="[615,668,1699,1726]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="437" unit="cm" value="2.0">2 cm</quantity>
|
||
long with antenna extending back to tergite 6; incomplete paramedian sutures on cephalic plate; anterior transverse suture on tergite 1; labrum tridentate; prefemur and femur of ultimate leg at least 4 times longer than wide, femur and tibia of ultimate leg lacking distal spinose processes; tibia with 7, tarsus 1 with 4 saw teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375652A2C46FFC3FF62F8E0A9BDF8A5" lastPageId="2" lastPageNumber="438" pageId="1" pageNumber="437" type="description">
|
||
<paragraph id="8BD036A12C46FFC0FF62F8E0AA08F885" blockId="1.[151,1437,1519,2014]" pageId="1" pageNumber="437">
|
||
<emphasis id="B91BEAB32C46FFC0FF62F8E0A83FF8B9" bold="true" box="[199,345,1843,1869]" pageId="1" pageNumber="437">Description.</emphasis>
|
||
Length
|
||
<quantity id="4C979B442C46FFC0FE16F8E0AB67F8BA" box="[435,513,1843,1870]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="437" unit="mm" value="19.0">19 mm</quantity>
|
||
(
|
||
<typeStatus id="54D488032C46FFC0FDABF8E0AB11F8B9" box="[526,631,1843,1869]" pageId="1" pageNumber="437" type="holotype">holotype</typeStatus>
|
||
). In ethanol stored specimen, head pale yellow and trunk (including legs) whitish with pale yellow posteromedian parts of tergites (
|
||
<figureCitation id="13542A242C46FFC0FCBEF884AA06F885" box="[795,864,1879,1905]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1618,1642]" captionTargetBox="[405,1184,193,1554]" captionTargetId="figure-19@3.[363,1224,181,1594]" captionTargetPageId="3" captionText="FIGURES 2–7. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 2–4. Cryptops datviensis sp. nov.: 2. Anterior part of body (holotype, IZISU TM-T-00001), dorsal view. 3. Head and forcipular segment (holotype, IZISU TM-T-00001), ventral view. 4. Clypeus and labrum (paratype, IZISU TM-T-00002), ventral view. 5–7. Cryptops caucasius Verhoeff, 1934: 5. Anterior part of body, dorsal view. 6. Head and forcipular segment ventral view. 7. Clypeus and labrum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318483" httpUri="https://zenodo.org/record/7318483/files/figure.png" pageId="1" pageNumber="437">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD036A12C46FFC3FF62F8A8AC0FFF0D" blockId="1.[151,1437,1519,2014]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="438" pageId="1" pageNumber="437">
|
||
Length and width of cephalic plate nearly equal (
|
||
<quantity id="4C979B442C46FFC0FCA1F8A8AA04F861" box="[772,866,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="437" unit="mm" value="1.2">1.2 mm</quantity>
|
||
), posterior corners rounded, sides weakly convex outwards for much of their length; two paramedian sutures on cephalic plate are incomplete, expressed anteriorly and posteriorly up to one fourth of cephalic plate length only. Antennae
|
||
<quantity id="4C979B442C46FFC0FC6FF817AD6AF82A" box="[970,1036,1988,2014]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="437" unit="mm" value="7.0">7 mm</quantity>
|
||
long (
|
||
<typeStatus id="54D488032C46FFC0FBF0F810ADDBF829" box="[1109,1213,1987,2013]" pageId="1" pageNumber="437" type="holotype">holotype</typeStatus>
|
||
), with 17 elongated articles; basal article stout, articles then gradually increasing in length to a maximum along articles 6–14 (ca 2 times longer than their width), then gradually shortening along articles 15–17. Articles 1–3 bearing strong setae of various length, these setae round proximal end of articles 4–5; from article 6, short fine setae form a fur-like covering.
|
||
</paragraph>
|
||
<paragraph id="8BD036A12C45FFC3FF62FED0A890FE91" blockId="2.[151,1437,151,357]" pageId="2" pageNumber="438">
|
||
Clypeus well defined anteriorly, with a triangular group of about 16 setae (
|
||
<figureCitation id="13542A242C45FFC3FBB3FED0AD3DFEE9" box="[1046,1115,259,285]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1618,1642]" captionTargetBox="[405,1184,193,1554]" captionTargetId="figure-19@3.[363,1224,181,1594]" captionTargetPageId="3" captionText="FIGURES 2–7. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 2–4. Cryptops datviensis sp. nov.: 2. Anterior part of body (holotype, IZISU TM-T-00001), dorsal view. 3. Head and forcipular segment (holotype, IZISU TM-T-00001), ventral view. 4. Clypeus and labrum (paratype, IZISU TM-T-00002), ventral view. 5–7. Cryptops caucasius Verhoeff, 1934: 5. Anterior part of body, dorsal view. 6. Head and forcipular segment ventral view. 7. Clypeus and labrum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318483" httpUri="https://zenodo.org/record/7318483/files/figure.png" pageId="2" pageNumber="438">Fig. 4</figureCitation>
|
||
), about 10 of them arranged in line along the posterior border. Labrum with side-pieces notched at their medial angle, tridentate, although the middle tooth is shorter (
|
||
<figureCitation id="13542A242C45FFC3FE06FE98A88FFE91" box="[419,489,331,357]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1618,1642]" captionTargetBox="[405,1184,193,1554]" captionTargetId="figure-19@3.[363,1224,181,1594]" captionTargetPageId="3" captionText="FIGURES 2–7. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 2–4. Cryptops datviensis sp. nov.: 2. Anterior part of body (holotype, IZISU TM-T-00001), dorsal view. 3. Head and forcipular segment (holotype, IZISU TM-T-00001), ventral view. 4. Clypeus and labrum (paratype, IZISU TM-T-00002), ventral view. 5–7. Cryptops caucasius Verhoeff, 1934: 5. Anterior part of body, dorsal view. 6. Head and forcipular segment ventral view. 7. Clypeus and labrum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318483" httpUri="https://zenodo.org/record/7318483/files/figure.png" pageId="2" pageNumber="438">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1066292C45FFC3FF32FBBBAC24FB74" ID-DOI="http://doi.org/10.5281/zenodo.7318479" ID-Zenodo-Dep="7318479" box="[151,1346,1128,1153]" httpUri="https://zenodo.org/record/7318479/files/figure.png" pageId="2" pageNumber="438" startId="2.[151,250,1128,1152]" targetBox="[338,1251,404,1076]" targetPageId="2">
|
||
<paragraph id="8BD036A12C45FFC3FF32FBBBAC24FB74" blockId="2.[151,1346,1128,1153]" box="[151,1346,1128,1153]" pageId="2" pageNumber="438">
|
||
<emphasis id="B91BEAB32C45FFC3FF32FBBBA875FB74" bold="true" box="[151,275,1128,1152]" pageId="2" pageNumber="438">FIGURE 1.</emphasis>
|
||
Entrance to Datvi Cave, a type locality of
|
||
<taxonomicName id="4C6F4D222C45FFC3FD61FBBAAAEBFB74" authority="Tuf & Barjadze & Maghradze, 2022" authorityName="Tuf & Barjadze & Maghradze" authorityYear="2022" box="[708,909,1128,1152]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="438" phylum="Arthropoda" rank="species" species="datviensis" status="sp. nov.">
|
||
<emphasis id="B91BEAB32C45FFC3FD61FBBAAAEBFB74" box="[708,909,1128,1152]" italics="true" pageId="2" pageNumber="438">Cryptops datviensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91BEAB32C45FFC3FC36FBBBAA8DFB74" bold="true" box="[915,1003,1128,1152]" pageId="2" pageNumber="438">
|
||
<taxonomicNameLabel id="A22857C82C45FFC3FC36FBBBAA80FB74" box="[915,998,1128,1152]" pageId="2" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
with the speleologist descending.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD036A12C45FFC3FF62FB7DAA38FAC1" blockId="2.[151,1437,1198,1981]" pageId="2" pageNumber="438">
|
||
Forcipular coxosternum with anterior edge slightly rounded with weak median notch (
|
||
<figureCitation id="13542A242C45FFC3FB02FB7DAD88FB3D" box="[1191,1262,1198,1225]" captionStart="FIGURES 2–7" captionStartId="3.[151,264,1618,1642]" captionTargetBox="[405,1184,193,1554]" captionTargetId="figure-19@3.[363,1224,181,1594]" captionTargetPageId="3" captionText="FIGURES 2–7. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 2–4. Cryptops datviensis sp. nov.: 2. Anterior part of body (holotype, IZISU TM-T-00001), dorsal view. 3. Head and forcipular segment (holotype, IZISU TM-T-00001), ventral view. 4. Clypeus and labrum (paratype, IZISU TM-T-00002), ventral view. 5–7. Cryptops caucasius Verhoeff, 1934: 5. Anterior part of body, dorsal view. 6. Head and forcipular segment ventral view. 7. Clypeus and labrum, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318483" httpUri="https://zenodo.org/record/7318483/files/figure.png" pageId="2" pageNumber="438">Fig. 3</figureCitation>
|
||
). Coxosternum bearing only a few sparse, moderately long or short setae. Trochanteroprefemur scattered with long, fine setae. Tarsungulum long, slender. Apical claw of second maxilla with slender hook and a flange along its inner edge. Dorsal brush dense, running along distal two-thirds of article 3.
|
||
</paragraph>
|
||
<paragraph id="8BD036A12C45FFC3FF62FAEDA880F9F8" blockId="2.[151,1437,1198,1981]" pageId="2" pageNumber="438">Tergites 2–7 broader than long, tergites 8–19 longer than broad. Overlap of tergite 1 by cephalic plate hides complete anterior transverse suture. Tergites 2–20 with anterior transverse suture hidden by preceding tergite. Tergites 3–20 with longitudinal paramedian sutures; oblique lateral sutures on tergites 3–5. Tergite of ultimate legbearing segment slightly longer than wide, posterior margin rounded; shallow longitudinal median depression on its posterior half. Sternites sparsely scattered with moderately long, thin setae. Posterior margin of sternite of ultimate leg-bearing segment rounded.</paragraph>
|
||
<paragraph id="8BD036A12C45FFC3FF62F9C4AA6BF8FD" blockId="2.[151,1437,1198,1981]" pageId="2" pageNumber="438">
|
||
Legs relatively long and slender. All tarsi strongly bipartite. All legs covered by sparse long setae, ventral face of tarsi with row of slender setae. Pair of accessory spurs half-length of pretarsus on legs 1–20. Ultimate leg of
|
||
<typeStatus id="54D488032C45FFC3FF32F98DA99FF98C" box="[151,249,1630,1656]" pageId="2" pageNumber="438" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13542A242C45FFC3FEADF98DA82AF98D" box="[264,332,1630,1657]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[310,1271,422,1817]" captionTargetId="figure-380@4.[270,1318,374,1878]" captionTargetPageId="4" captionText="FIGURES 8–11. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 8, 9. Cryptops datviensis sp. nov.: 8. Posterior part of body (holotype, IZISU TM-T-00001), lateral view. 9. End of right ultimate leg (paratype, IZISU TM-T-00002), lateral view. 10, 11. Cryptops caucasius Verhoeff, 1934: 10. Posterior part of body, lateral view. 11. End of right ultimate leg, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7318485" httpUri="https://zenodo.org/record/7318485/files/figure.png" pageId="2" pageNumber="438">Fig. 8</figureCitation>
|
||
): prefemur
|
||
<quantity id="4C979B442C45FFC3FE75F98DAB40F98D" box="[464,550,1630,1657]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="438" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long, femur
|
||
<quantity id="4C979B442C45FFC3FD10F98CAA6BF98D" box="[693,781,1631,1657]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="438" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, tibia
|
||
<quantity id="4C979B442C45FFC3FCF4F98CAACFF98D" box="[849,937,1631,1657]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="438" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, tarsus 1
|
||
<quantity id="4C979B442C45FFC3FBB4F98DAD0FF98D" box="[1041,1129,1630,1657]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="438" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, tarsus 2
|
||
<quantity id="4C979B442C45FFC3FB77F98CAC4CF98D" box="[1234,1322,1631,1657]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="438" unit="mm" value="0.9">0.9 mm</quantity>
|
||
; prefemur and femur ca 4 times and 5 times longer than their maximal widths at distal ends, respectively. Robust setae on anterior, posterior and ventral sides of prefemur and femur, mostly slender setae on dorsal side; tibial and tarsal setae slender; 7 saw teeth on tibia, 4 on tarsus 1 (
|
||
<figureCitation id="13542A242C45FFC3FDD3F919ABDCF911" box="[630,698,1738,1765]" captionStart="FIGURES 8–11" captionStartId="4.[151,264,1901,1925]" captionTargetBox="[310,1271,422,1817]" captionTargetId="figure-380@4.[270,1318,374,1878]" captionTargetPageId="4" captionText="FIGURES 8–11. Cryptops datviensis sp. nov. and C. caucasius Verhoeff, 1934. 8, 9. Cryptops datviensis sp. nov.: 8. Posterior part of body (holotype, IZISU TM-T-00001), lateral view. 9. End of right ultimate leg (paratype, IZISU TM-T-00002), lateral view. 10, 11. Cryptops caucasius Verhoeff, 1934: 10. Posterior part of body, lateral view. 11. End of right ultimate leg, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7318485" httpUri="https://zenodo.org/record/7318485/files/figure.png" pageId="2" pageNumber="438">Fig. 9</figureCitation>
|
||
, the same situation on
|
||
<typeStatus id="54D488032C45FFC3FC12F918AD4EF911" box="[951,1064,1739,1765]" pageId="2" pageNumber="438" type="paratype">paratypes</typeStatus>
|
||
); inner margin of tarsus 2 a ridge. Apical claw well developed, without basal sensory spur.
|
||
</paragraph>
|
||
<paragraph id="8BD036A12C45FFC3FF62F8C1A9BDF8A5" blockId="2.[151,1437,1198,1981]" pageId="2" pageNumber="438">Coxopleural pore field elliptical, with ca 35–40 pores and 4–5 slender, short setae interspersed among the pores.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375652A2C45FFC3FF62F888AAF0F880" box="[199,918,1882,1909]" pageId="2" pageNumber="438" type="etymology">
|
||
<paragraph id="8BD036A12C45FFC3FF62F888AAF0F880" blockId="2.[151,1437,1198,1981]" box="[199,918,1882,1909]" pageId="2" pageNumber="438">
|
||
<emphasis id="B91BEAB32C45FFC3FF62F888A828F881" bold="true" box="[199,334,1883,1909]" pageId="2" pageNumber="438">Etymology.</emphasis>
|
||
The specific name originates from the
|
||
<typeStatus id="54D488032C45FFC3FCA0F888AA53F881" box="[773,821,1883,1909]" pageId="2" pageNumber="438">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375652A2C45FFC3FF62F8ADA9BBF849" pageId="2" pageNumber="438" type="distribution">
|
||
<paragraph id="8BD036A12C45FFC3FF62F8ADA9BBF849" blockId="2.[151,1437,1198,1981]" pageId="2" pageNumber="438">
|
||
<emphasis id="B91BEAB32C45FFC3FF62F8ADA807F86C" bold="true" box="[199,353,1918,1944]" pageId="2" pageNumber="438">Distribution.</emphasis>
|
||
The new species is known from the
|
||
<typeStatus id="54D488032C45FFC3FD4EF8ACAA7DF86D" box="[747,795,1919,1945]" pageId="2" pageNumber="438">type</typeStatus>
|
||
locality only. Probably it can be found in the neighbouring caves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |