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<document id="05D92DFDE307AF797206BA6005CDBA90" ID-DOI="10.5281/zenodo.274629" ID-GBIF-Dataset="1568d832-965a-40c3-b154-ec99dea72e89" ID-ISSN="1175-5326" ID-Zenodo-Dep="274629" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459948516208" checkinUser="plazi" docAuthor="Passos, Paulo &amp; Arredondo, Juan C." docDate="2009" docId="03C687ABF713601D70D10B88A0AC92DD" docLanguage="en" docName="zt01969p068.pdf" docOrigin="Zootaxa 1969" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractus wagleri" docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="FFFFFFD3F712601870460F47A265932F" masterDocTitle="Rediscovery and redescription of the Andean earth-snake Atractus wagleri (Reptilia: Serpentes: Colubridae)" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698230711175" updateUser="plazi">
<mods:mods id="6B9E1DD29B8CDB0D1E876AA4160F707E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9DB471FC98FA83F155E263DBD40F721E">
<mods:title id="5389617B6A7F6BF0F2FD61BD9919E1D9">Rediscovery and redescription of the Andean earth-snake Atractus wagleri (Reptilia: Serpentes: Colubridae)</mods:title>
</mods:titleInfo>
<mods:name id="B235F15681ED8481F3824915F25349B5" type="personal">
<mods:role id="310D145B431DD992DF167CE6B1A5D3FD">
<mods:roleTerm id="9062976E0EA18B59B9E6D2FA1C0B8E20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22014B1461D1B717E1B1CF72882571BC">Passos, Paulo</mods:namePart>
</mods:name>
<mods:name id="3C4A9AF481B788EE21D253628F04AE2D" type="personal">
<mods:role id="32024C75A972131EA011B69E6880F30D">
<mods:roleTerm id="92316DB86004F15D0115592F2BA64A7B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C4A8968E6CAF312948D5151A73056818">Arredondo, Juan C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="51127D39DC557222886A9D7D53DE3005">text</mods:typeOfResource>
<mods:relatedItem id="3F958902AD72B3F26001C419AE909F6A" type="host">
<mods:titleInfo id="0B7A7D0003AFD13CB8EFD2D59DBA2E3A">
<mods:title id="91578D9E3730399BE117A2DE30E63845">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3BB033B3AEFB16B05E27DF59AEC4F9C9">
<mods:date id="7218D839393AAAC36451EDD752433219">2009</mods:date>
<mods:detail id="F83303FD7F2A6174D99C4B03D5078D07" type="volume">
<mods:number id="7B2F16D445F5BD8533E0C632211B8A22">1969</mods:number>
</mods:detail>
<mods:extent id="0591AA2968CD08BB4D0AEAD932CBB701" unit="page">
<mods:start id="7273AAFBDC0C6D547E4812FD8B11B826">59</mods:start>
<mods:end id="4287E78D9420EAA5F02612349204CA2C">68</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="66754573DE9668C112CF24C7F0221B59">journal article</mods:classification>
<mods:identifier id="AB8214B2968809579EF3D300347B9BBA" type="DOI">10.5281/zenodo.274629</mods:identifier>
<mods:identifier id="787EFD21A132B61F54FFC4C61504A866" type="GBIF-Dataset">1568d832-965a-40c3-b154-ec99dea72e89</mods:identifier>
<mods:identifier id="726CD3434A8AD773EDB2160235A24496" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="716C3A406D0E100F5439987EA9B4699D" type="Zenodo-Dep">274629</mods:identifier>
</mods:mods>
<treatment id="03C687ABF713601D70D10B88A0AC92DD" ID-DOI="http://doi.org/10.5281/zenodo.6219359" ID-GBIF-Taxon="119376379" ID-Zenodo-Dep="6219359" LSID="urn:lsid:plazi:treatment:03C687ABF713601D70D10B88A0AC92DD" httpUri="http://treatment.plazi.org/id/03C687ABF713601D70D10B88A0AC92DD" lastPageId="5" lastPageNumber="64" pageId="1" pageNumber="60">
<subSubSection id="C3756536F713601970D10B88A3509623" pageId="1" pageNumber="60" type="nomenclature">
<paragraph id="8BD036BDF713601970D10B88A33A97C6" blockId="1.[151,351,1231,1292]" box="[151,351,1231,1257]" pageId="1" pageNumber="60">
<heading id="D09881D1F713601970D10B88A33A97C6" bold="true" box="[151,351,1231,1257]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<taxonomicName id="4C6F4D3EF713601970D10B88A33A97C6" box="[151,351,1231,1257]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF713601970D10B88A33A97C6" bold="true" box="[151,351,1231,1257]" italics="true" pageId="1" pageNumber="60">Atractus wagleri</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD036BDF713601970D10BB5A3509623" blockId="1.[151,351,1231,1292]" box="[151,309,1266,1292]" pageId="1" pageNumber="60">
(
<figureCitation id="13542A38F713601970D90BB5A3489623" box="[159,301,1266,1292]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,269,1153,1177]" captionStartId-1="3.[151,255,1861,1885]" captionStartId-2="4.[151,255,1651,1675]" captionTargetBox-0="[220,1327,616,1152]" captionTargetBox-1="[151,1435,877,1838]" captionTargetBox-2="[151,1435,834,1627]" captionTargetId-0="figure@2.[220,1367,597,1165]" captionTargetId-1="figure@3.[151,1436,877,1838]" captionTargetId-2="figure@4.[151,1436,834,1628]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 2. Dorsal (1) and ventral (2) views of Atractus wagleri (MHUA 14504) from San Vicenti de Chucurí. SVL 445 mm, CL 96 mm." captionText-1="FIGURE 3. General view of Atractus wagleri (MHUA 14504) in life." captionText-2="FIGURE 4. Sulcate (left) and asulcate (right) views of the hemipenis of Atractus wagleri (MHUA 14504). Scale = 10 mm." httpUri-0="https://zenodo.org/record/274630/files/figure.png" httpUri-1="https://zenodo.org/record/274631/files/figure.png" httpUri-2="https://zenodo.org/record/274632/files/figure.png" pageId="1" pageNumber="60">Figures 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3756536F713601970D10A05A0BB9676" box="[151,734,1346,1369]" pageId="1" pageNumber="60" type="reference_group">
<paragraph id="8BD036BDF713601970D10A05A0BB9676" blockId="1.[151,734,1346,1369]" box="[151,734,1346,1369]" pageId="1" pageNumber="60">
<taxonomicName id="4C6F4D3EF713601970D10A05A3B69676" authority="Prado, 1945" authorityName="Prado" authorityYear="1945" box="[151,467,1346,1369]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF713601970D10A05A32F9676" box="[151,330,1346,1369]" italics="true" pageId="1" pageNumber="60">Atractus wagleri</emphasis>
<bibRefCitation id="EFFE4B4CF713601971160A05A3B69676" author="Prado" box="[336,467,1346,1369]" pageId="1" pageNumber="65" refString="Prado, A. (1945). Um novo Atractus de Colombia. Ciencia (Mexico), 6, 61." type="journal article" year="1945">Prado, 1945</bibRefCitation>
</taxonomicName>
; Ciencia (
<collectingCountry id="F378762DF713601972780A05A0F39676" box="[574,662,1346,1369]" name="Mexico" pageId="1" pageNumber="60">Mexico</collectingCountry>
): 6:61.
</paragraph>
</subSubSection>
<subSubSection id="C3756536F713601970D10ACDA6E396E3" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="8BD036BDF713601970D10ACDA6E396E3" blockId="1.[151,1437,1418,2004]" pageId="1" pageNumber="60">
<emphasis id="B91BEAAFF713601970D10ACDA377968B" bold="true" box="[151,274,1418,1444]" pageId="1" pageNumber="60">
<typeStatus id="54D4881FF713601970D10ACDA36B968B" box="[151,270,1418,1444]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
:
</emphasis>
Female,
<collectionCode id="ED7EAE78F713601971C60ACDA3D9968B" box="[384,444,1418,1444]" httpUri="http://grbio.org/cool/mztg-j2c7" name="Museo del Instituto de La Salle" pageId="1" pageNumber="60">MLS</collectionCode>
228, from Humbo (
<geoCoordinate id="EE5B507AF713601972EC0ACDA174968B" box="[682,785,1418,1444]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="925" value="5.6">05º36’N</geoCoordinate>
,
<geoCoordinate id="EE5B507AF7136019735B0ACDA1E2968B" box="[797,903,1418,1444]" direction="west" orientation="longitude" pageId="1" pageNumber="60" precision="925" value="-74.26667">74º16’W</geoCoordinate>
, ca.
<quantity id="4C979B58F713601973FB0ACDA67C968B" box="[957,1049,1418,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="60" unit="m" value="1000.0">1000 m</quantity>
), Boyacá Department of
<collectingCountry id="F378762DF713601975040ACDA2DB96E3" name="Colombia" pageId="1" pageNumber="60">Colombia</collectingCountry>
, collected on 1948 by Hermanos of La Salle order. Presumed lost (see discussion).
</paragraph>
</subSubSection>
<subSubSection id="C3756536F713601A70830A9DA39E92DD" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60" type="diagnosis">
<paragraph id="8BD036BDF713601970830A9DA0E39473" blockId="1.[151,1437,1418,2004]" pageId="1" pageNumber="60">
<emphasis id="B91BEAAFF713601970830A9DA32296DB" bold="true" box="[197,327,1498,1524]" pageId="1" pageNumber="60">Diagnosis:</emphasis>
<taxonomicName id="4C6F4D3EF713601971080A9CA07796DB" box="[334,530,1499,1524]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF713601971080A9CA07796DB" box="[334,530,1499,1524]" italics="true" pageId="1" pageNumber="60">Atractus wagleri</emphasis>
</taxonomicName>
is distinguished from all congeners by the combination of the following characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) loreal long; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) seven infralabials, first three contacting chinshields; (7) six or seven maxillary teeth; (8) generally four gular scale rows; (9) four or five preventrals; (10) 174–180 ventrals in females,
<quantity id="4C979B58F713601971C2093DA06C95BB" box="[388,521,1658,1684]" metricMagnitude="0" metricUnit="m" metricValue="4.2036999999999995" metricValueMax="4.4196" metricValueMin="3.9878" pageId="1" pageNumber="60" unit="in" value="165.5" valueMax="174.0" valueMin="157.0">157–174 in</quantity>
males; (11) 43–44 subcaudals in females,
<quantity id="4C979B58F713601973B8093DA60395BB" box="[1022,1126,1658,1684]" metricMagnitude="0" metricUnit="m" metricValue="1.2953999999999999" metricValueMax="1.4224" metricValueMin="1.1684" pageId="1" pageNumber="60" unit="in" value="51.0" valueMax="56.0" valueMin="46.0">46–56 in</quantity>
males; (12) dorsal ground colour in preservative cream-red with an irregular vertebral black stripe connected to lateral black blotches, constituting complete bands anteriorly and decreasing in size posteriorly; (13) venter black with paraventral region cream; (14) moderate body size, females reaching
<quantity id="4C979B58F7136019737C09B5A1F99423" box="[826,924,1778,1804]" metricMagnitude="-1" metricUnit="m" metricValue="4.37" pageId="1" pageNumber="60" unit="mm" value="437.0">437 mm</quantity>
SVL and males
<quantity id="4C979B58F7136019742609B5A6A49423" box="[1120,1217,1778,1804]" metricMagnitude="-1" metricUnit="m" metricValue="4.45" pageId="1" pageNumber="60" unit="mm" value="445.0">445 mm</quantity>
SVL; (15) moderate tail length in females (13.6–15.3% SVL) and long in male (21.61% SVL); (16) hemipenis moderately bilobed, semicapitate, and semicalyculate.
</paragraph>
<paragraph id="8BD036BDF713601A7083082DA39E92DD" blockId="1.[151,1437,1418,2004]" lastBlockId="2.[151,1437,152,498]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
<emphasis id="B91BEAAFF71360197083082DA31094AB" bold="true" box="[197,373,1898,1924]" pageId="1" pageNumber="60">Comparisons:</emphasis>
Among all congeners,
<taxonomicName id="4C6F4D3EF713601972CA082CA13794AB" box="[652,850,1899,1924]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF713601972CA082CA13794AB" box="[652,850,1899,1924]" italics="true" pageId="1" pageNumber="60">Atractus wagleri</emphasis>
</taxonomicName>
occur parapatrically and shared a suite combination of characters with
<taxonomicName id="4C6F4D3EF713601971EF08D4A02A9483" box="[425,591,1939,1964]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B91BEAAFF713601971EF08D4A02A9483" box="[425,591,1939,1964]" italics="true" pageId="1" pageNumber="60">A. sanguineus</emphasis>
</taxonomicName>
: 17 dorsal scale rows, more than 170 ventrals in both sexes, more than 40 subcaudals in males, seven upper and lower labials, first three infralabials contacting chinshields, seven or eight maxillary teeth, long tail in males, dorsum with blotches decreasing in size posteriorly, and venter generally most black.
<taxonomicName id="4C6F4D3EF710601A711E0F87A07993F6" box="[344,540,192,217]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF710601A711E0F87A07993F6" box="[344,540,192,217]" italics="true" pageId="2" pageNumber="61">Atractus wagleri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6F4D3EF710601A72F00F87A13E93F6" box="[694,859,192,217]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B91BEAAFF710601A72F00F87A13E93F6" box="[694,859,192,217]" italics="true" pageId="2" pageNumber="61">A. sanguineus</emphasis>
</taxonomicName>
in having 43–44 subcaudals in females and paravertebral blotches constituting bands anteriorly (vs. 29–38 subcaudals in females and paravertebral blotches never forming bands, except for occasionally a black collar on neck, anteriorly). The only other species of
<taxonomicName id="4C6F4D3EF710601A70D10E7FA29F927E" box="[151,250,312,337]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="B91BEAAFF710601A70D10E7FA29F927E" box="[151,250,312,337]" italics="true" pageId="2" pageNumber="61">Atractus</emphasis>
</taxonomicName>
occurring parapatrycally with
<taxonomicName id="4C6F4D3EF710601A72360E7FA088927E" box="[624,749,312,337]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF710601A72360E7FA088927E" box="[624,749,312,337]" italics="true" pageId="2" pageNumber="61">A. wagleri</emphasis>
</taxonomicName>
at Cordillera Oriental of
<collectingCountry id="F378762DF710601A74630E7FA6FC927D" box="[1061,1177,312,338]" name="Colombia" pageId="2" pageNumber="61">Colombia</collectingCountry>
is
<taxonomicName id="4C6F4D3EF710601A74870E7FA7F1927E" box="[1217,1428,312,337]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="crassicaudatus">
<emphasis id="B91BEAAFF710601A74870E7FA7F1927E" box="[1217,1428,312,337]" italics="true" pageId="2" pageNumber="61">A. crassicaudatus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6F4D3EF710601A70D10E27A3389256" box="[151,349,352,377]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF710601A70D10E27A3389256" box="[151,349,352,377]" italics="true" pageId="2" pageNumber="61">Atractus wagleri</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C6F4D3EF710601A72AB0E27A1DA9256" box="[749,959,352,377]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="crassicaudatus">
<emphasis id="B91BEAAFF710601A72AB0E27A1DA9256" box="[749,959,352,377]" italics="true" pageId="2" pageNumber="61">A. crassicaudatus</emphasis>
</taxonomicName>
by having 174–180 ventrals in females, 43–44 subcaudals in females and
<quantity id="4C979B58F710601A72660ECFA0E2928D" box="[544,647,392,418]" metricMagnitude="0" metricUnit="m" metricValue="1.2953999999999999" metricValueMax="1.4224" metricValueMin="1.1684" pageId="2" pageNumber="61" unit="in" value="51.0" valueMax="56.0" valueMin="46.0">46–56 in</quantity>
males, and tail 21.61% SVL in males, 13.6–15.3% SVL in females (vs. 147–170 ventrals in females; 21–33 subcaudals in males and
<quantity id="4C979B58F710601A73E30EF7A66B92E5" box="[933,1038,432,458]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="6.604" metricValueMin="3.556" pageId="2" pageNumber="61" unit="in" value="20.0" valueMax="26.0" valueMin="14.0">14–26 in</quantity>
females; tail 11.0–17.1% SVL in males, 8.0–13.7% in females).
</paragraph>
</subSubSection>
<caption id="DF106635F710601A70D10BC6A3129794" httpUri="https://zenodo.org/record/274630/files/figure.png" pageId="2" pageNumber="61" targetBox="[220,1327,616,1152]" targetPageId="2">
<paragraph id="8BD036BDF710601A70D10BC6A3129794" blockId="2.[151,1436,1153,1211]" pageId="2" pageNumber="61">
<emphasis id="B91BEAAFF710601A70D10BC6A32797B6" bold="true" box="[151,322,1153,1177]" pageId="2" pageNumber="61">FIGURES 1–2.</emphasis>
Dorsal (1) and ventral (2) views of
<taxonomicName id="4C6F4D3EF710601A72860BC5A11797B6" box="[704,882,1154,1177]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF710601A72860BC5A11797B6" box="[704,882,1154,1177]" italics="true" pageId="2" pageNumber="61">Atractus wagleri</emphasis>
</taxonomicName>
(MHUA 14504) from San Vicenti de Chucurí. SVL 445 mm, CL 96 mm.
</paragraph>
</caption>
<subSubSection id="C3756536F710601A70830BAEA38794FB" pageId="2" pageNumber="61" type="materials_examined">
<paragraph id="8BD036BDF710601A70830BAEA0AD941B" blockId="2.[151,1437,1257,2004]" pageId="2" pageNumber="61">
<emphasis id="B91BEAAFF710601A70830BAEA1D1962C" bold="true" box="[197,948,1257,1283]" pageId="2" pageNumber="61">
Description (based on 3 new specimens;
<specimenCount id="9D69FD34F710601A72800BAEA142962C" box="[710,807,1257,1283]" pageId="2" pageNumber="61" type="male">2 males</specimenCount>
,
<specimenCount id="9D69FD34F710601A73700BAEA1C3962C" box="[822,934,1257,1283]" pageId="2" pageNumber="61" type="female">1 female</specimenCount>
):
</emphasis>
Head twice as long as wide, flattened in lateral view, round in dorsal view; snout truncate in lateral view, round in dorsal view; rostral subtriangular in frontal view, broader than high, little visible in dorsal view; internasal as long as wide; internasal suture sinistral with respect to prefrontal suture; prefrontal longer than wide; supraocular subtrapezoidal, slightly longer than wide; frontal subpentagonal, broader than long; parietal about twice as long as wide; nasal divided; nostril located between prenasal and postnasal; prenasal twice as high as long; postnasal about twice as high as long, higher than prenasal; loreal long, contacting second and third supralabials; pupil round; two postoculars of similar length; upper postocular about twice as high as lower postocular; temporals 1+2; anterior temporal twice as long as high; upper posterior temporal elongate, three times as long as wide; seven supralabials, third and fourth contacting orbit; second supralabial higher than first and smaller than third; third higher and seventh longer than remaining supralabials; symphisial subtriangular, four times broader than long; seven infralabials, first three contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshields contact; chinshields three times as long as wide; three or four gular scale rows; four or five preventrals; 17/17/17 smooth dorsal scale rows; dorsals lacking apical pits, supra-anal tubercles, and keels; caudal spine short, robust, and rhomboid.
</paragraph>
<paragraph id="8BD036BDF710601A70830806A38794FB" blockId="2.[151,1437,1257,2004]" pageId="2" pageNumber="61">
<emphasis id="B91BEAAFF710601A70830806A3EB9474" bold="true" box="[197,398,1857,1883]" pageId="2" pageNumber="61">Maxillary arch:</emphasis>
Arched in dorsal view, with four or five prediastemal and two or three postdiastemal teeth; prediastemal teeth large, curved downward, decreasing gradually in size posteriorly, angular in cross section, robust at base, and narrower at apices; maxillary diastema short; postdiastemal teeth slightly smaller than last prediastemal tooth.
</paragraph>
</subSubSection>
<subSubSection id="C3756536F711601C70830FD0A7149025" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="description">
<paragraph id="8BD036BDF711601B70830FD0A3AE901B" blockId="3.[151,1437,151,820]" pageId="3" pageNumber="62">
<emphasis id="B91BEAAFF711601B70830FD0A029939E" bold="true" box="[197,588,151,177]" pageId="3" pageNumber="62">Colour pattern in preservative:</emphasis>
Dorsum of head black, covered with small beige dots above internasals, nasals, loreal, and prefrontals; background of head black to dorsal margins of supralabials; black descending postocular stripe reaching dorsal edge of sixth and anterior ventral margin of seventh supralabial; temporal region pigmented with irregular beige spots; median region of supralabials cream-white, except for sixth and seventh scales; mental region cream-white, occasionally with disperse black dots; preventrals uniformly cream; venter predominantly cream-white with irregular black dots for first 15 ventrals; venter becomes uniformly black before 15th ventral, except for lateral margin of scales uniformly cream-white; tail black with lateral portion of subcaudals cream-white; dorsal ground colour of body black anteriorly, with cream-white complete transverse bands (one scale long), alternated on flanks; black interspaces (three to four scales long), reaching paraventral region; pale transverse bands of opposite sides connected middorsally only anteriorly, up to a point level with 20th ventral; pale transverse bands increase in size at midbody (two scales long), with black interspaces reducing gradually in size, restricted to paravertebral region (two scales long); posterior to midbody region, dorsal ground colour becomes cream-red with black vertebral line (two scales wide), breaking up to for black paravertebral blotches; flanks with round black blotches (two scales long) at the level of third and fourth scale rows; paraventral region with irregular small black blotches, constituting a barely defined lateral stripe above first two dorsal scale rows; paraventral blotches occasionally connected to paravertebral ones (
<figureCitation id="13542A38F711601B710A0C5DA3D8901B" box="[332,445,794,820]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,269,1153,1177]" captionTargetBox="[220,1327,616,1152]" captionTargetId="figure@2.[220,1367,597,1165]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Dorsal (1) and ventral (2) views of Atractus wagleri (MHUA 14504) from San Vicenti de Chucurí. SVL 445 mm, CL 96 mm." httpUri="https://zenodo.org/record/274630/files/figure.png" pageId="3" pageNumber="62">Figs. 1–2</figureCitation>
).
</paragraph>
<caption id="DF106635F711601B70D10802A1E09472" box="[151,901,1861,1885]" httpUri="https://zenodo.org/record/274631/files/figure.png" pageId="3" pageNumber="62" targetBox="[151,1435,877,1838]" targetPageId="3">
<paragraph id="8BD036BDF711601B70D10802A1E09472" blockId="3.[151,901,1861,1885]" box="[151,901,1861,1885]" pageId="3" pageNumber="62">
<emphasis id="B91BEAAFF711601B70D10802A37C9472" bold="true" box="[151,281,1861,1885]" pageId="3" pageNumber="62">FIGURE 3.</emphasis>
General view of
<taxonomicName id="4C6F4D3EF711601B71940801A0E19472" box="[466,644,1862,1885]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF711601B71940801A0E19472" box="[466,644,1862,1885]" italics="true" pageId="3" pageNumber="62">Atractus wagleri</emphasis>
</taxonomicName>
(MHUA 14504) in life.
</paragraph>
</caption>
<paragraph id="8BD036BDF711601C708308F4A6FB9205" blockId="3.[151,1437,1971,2037]" lastBlockId="4.[151,1437,152,778]" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<emphasis id="B91BEAAFF711601B708308F4A3BF94E2" bold="true" box="[197,474,1971,1997]" pageId="3" pageNumber="62">Colour pattern in life:</emphasis>
Dorsum of head black with yellow dots above internasals, nasals, loreal, prefrontal, and temporal region; ventral margin of supralabials yellow; mental region and venter anteriorly creamyellow; lateral margin of ventrals cream-yellow and remaining belly uniformly black; dorsum of body black anteriorly, with yellow transverse bands; bands gradually change to reddish orange in the median third and uniformly red at posterior region of body; vertebral line and paravertebral blotches black; flanks and paraventral region with cream-red ground colour, covered with irregular black blotches (
<figureCitation id="13542A38F716601C74010E57A6EB9205" box="[1095,1166,272,298]" captionStart="FIGURE 3" captionStartId="3.[151,255,1861,1885]" captionTargetBox="[151,1435,877,1838]" captionTargetId="figure@3.[151,1436,877,1838]" captionTargetPageId="3" captionText="FIGURE 3. General view of Atractus wagleri (MHUA 14504) in life." httpUri="https://zenodo.org/record/274631/files/figure.png" pageId="4" pageNumber="63">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BD036BDF716601C70830E70A7149025" blockId="4.[151,1437,152,778]" pageId="4" pageNumber="63">
<emphasis id="B91BEAAFF716601C70830E70A32F927E" bold="true" box="[197,330,311,337]" pageId="4" pageNumber="63">Hemipenis</emphasis>
(
<figureCitation id="13542A38F716601C711B0E7FA3C2927D" box="[349,423,312,338]" captionStart="FIGURE 4" captionStartId="4.[151,255,1651,1675]" captionTargetBox="[151,1435,834,1627]" captionTargetId="figure@4.[151,1436,834,1628]" captionTargetPageId="4" captionText="FIGURE 4. Sulcate (left) and asulcate (right) views of the hemipenis of Atractus wagleri (MHUA 14504). Scale = 10 mm." httpUri="https://zenodo.org/record/274632/files/figure.png" pageId="4" pageNumber="63">Fig. 4</figureCitation>
; everted organs
<emphasis id="B91BEAAFF716601C722A0E7FA01E927E" box="[620,635,312,337]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2): Organ moderately bilobed, semicapitate, and semicalyculate; lobes distinct and restricted to distal portion of capitulum; lobes cylindrical with round, laterally projecting apices; lateral projections oriented centrifugally, clearly distinct from proximal region of lobes; lobes and capitulum uniformly covered with spinulate calyces; calyces progressively replaced by papillae toward tips of lobes; lobes with calyces losing vertical walls and constituting spinulate transverse flounces; transverse flounces well defined on lateral portion of asulcate side of hemipenis; asulcate side of capitulum with barely conspicuous lobular crests; capitular crotch well marked on both sides of hemipenis; capitulum similar in size to hemipenial body, located at the level of sulcus spermaticus bifurcation; sulcus spermaticus bifurcates in the mid portion of the organ; branches of sulcus spermaticus centrifugal, reaching lobe apices; margins of sulcus spermaticus stout and narrow along hemipenial body, and moderately expanded in the capitular region; hemipenial body subcylindrical, covered with moderate hooked spines; basal naked pocket restricted to basal portion of hemipenial body; proximal portion of hemipenis with longitudinal plicae and disperse spinules.
</paragraph>
</subSubSection>
<caption id="DF106635F716601C70D10934A2879582" httpUri="https://zenodo.org/record/274632/files/figure.png" pageId="4" pageNumber="63" targetBox="[151,1435,834,1627]" targetPageId="4">
<paragraph id="8BD036BDF716601C70D10934A2879582" blockId="4.[151,1436,1651,1709]" pageId="4" pageNumber="63">
<emphasis id="B91BEAAFF716601C70D10934A37995A4" bold="true" box="[151,284,1651,1675]" pageId="4" pageNumber="63">FIGURE 4.</emphasis>
Sulcate (left) and asulcate (right) views of the hemipenis of
<taxonomicName id="4C6F4D3EF716601C738E0933A61A95A4" box="[968,1151,1652,1675]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF716601C738E0933A61A95A4" box="[968,1151,1652,1675]" italics="true" pageId="4" pageNumber="63">Atractus wagleri</emphasis>
</taxonomicName>
(MHUA 14504). Scale = 10mm.
</paragraph>
</caption>
<subSubSection id="C3756536F716601C7083099BA0A29491" pageId="4" pageNumber="63" type="materials_examined">
<paragraph id="8BD036BDF716601C7083099BA0A29491" blockId="4.[151,1436,1756,1982]" pageId="4" pageNumber="63">
<emphasis id="B91BEAAFF716601C7083099BA66595D9" bold="true" box="[197,1024,1756,1782]" pageId="4" pageNumber="63">
Variation (3 new specimens plus Prado's data on female
<typeStatus id="54D4881FF716601C73C7099BA19695D9" box="[897,1011,1756,1782]" pageId="4" pageNumber="63" type="holotype">holotype</typeStatus>
):
</emphasis>
Largest male SVL
<quantity id="4C979B58F716601C74A5099BA72295D9" box="[1251,1351,1756,1782]" metricMagnitude="-1" metricUnit="m" metricValue="4.45" pageId="4" pageNumber="63" unit="mm" value="445.0">445 mm</quantity>
, CL
<quantity id="4C979B58F716601C7539099BA2AD9431" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="63" unit="mm" value="96.0">96 mm</quantity>
; largest female SVL
<quantity id="4C979B58F716601C71870843A0439431" box="[449,550,1796,1822]" metricMagnitude="-1" metricUnit="m" metricValue="4.37" pageId="4" pageNumber="63" unit="mm" value="437.0">437 mm</quantity>
, CL
<quantity id="4C979B58F716601C72260843A0D39431" box="[608,694,1796,1822]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="63" unit="mm" value="67.0">67 mm</quantity>
; tail 21.6% SVL in male, 13.6–15.3% (
<emphasis id="B91BEAAFF716601C74CD0843A6FF9432" box="[1163,1178,1796,1821]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) SVL in females; 157–174 (
<emphasis id="B91BEAAFF716601C7148086BA378946A" box="[270,285,1836,1861]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) ventrals in males, 174–180 (
<emphasis id="B91BEAAFF716601C72E1086BA0D3946A" box="[679,694,1836,1861]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) in females; 45–56 (
<emphasis id="B91BEAAFF716601C7390086BA180946A" box="[982,997,1836,1861]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) subcaudals in males, 43–44 (
<emphasis id="B91BEAAFF716601C7533086BA7E1946A" box="[1397,1412,1836,1861]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) in females; 3 (
<emphasis id="B91BEAAFF716601C71200813A3109442" box="[358,373,1876,1901]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 1 side) or 4 (
<emphasis id="B91BEAAFF716601C72720813A0269442" box="[564,579,1876,1901]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 5 sides) gular scale rows; 4 (
<emphasis id="B91BEAAFF716601C73F90813A1AB9442" box="[959,974,1876,1901]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2) or 5 (
<emphasis id="B91BEAAFF716601C74120813A6079442" box="[1108,1122,1876,1901]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 1) preventrals; 8 (
<emphasis id="B91BEAAFF716601C751A0813A70E9442" box="[1372,1387,1876,1901]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 4 sides) or 9 (
<emphasis id="B91BEAAFF716601C7160083BA35094BA" box="[294,309,1916,1941]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2 side) dorsal scale rows in the level of second subcaudal; 6 (
<emphasis id="B91BEAAFF716601C7471083BA62394BA" box="[1079,1094,1916,1941]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 2 sides) or 7 (
<emphasis id="B91BEAAFF716601C7549083BA77B94BA" box="[1295,1310,1916,1941]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 4 sides) maxillary teeth; 4.5–9.0 mm midbody diameter.
</paragraph>
</subSubSection>
<subSubSection id="C3756536F717601D70830FD0A6B7922D" pageId="5" pageNumber="64" type="distribution">
<paragraph id="8BD036BDF717601D70830FD0A6B7922D" blockId="5.[151,1437,151,498]" pageId="5" pageNumber="64">
<emphasis id="B91BEAAFF717601D70830FD0A30D939E" bold="true" box="[197,360,151,177]" pageId="5" pageNumber="64">Distribution:</emphasis>
Western versant of Cordillera Oriental of
<collectingCountry id="F378762DF717601D73380FDFA190939D" box="[894,1013,152,178]" name="Colombia" pageId="5" pageNumber="64">Colombia</collectingCountry>
, from Floridablanca (
<geoCoordinate id="EE5B507AF717601D75450FDFA7F3939D" box="[1283,1430,152,178]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="15" value="7.0647225">07º03’53’’N</geoCoordinate>
,
<geoCoordinate id="EE5B507AF717601D70D10F87A34A93F5" box="[151,303,192,218]" direction="west" orientation="longitude" pageId="5" pageNumber="64" precision="15" value="-73.08972">73º05’23’’W</geoCoordinate>
) in Santander Department to Humbo (
<geoCoordinate id="EE5B507AF717601D73510F87A11893F5" box="[791,893,192,218]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="925" value="5.6">05º36’N</geoCoordinate>
,
<geoCoordinate id="EE5B507AF717601D73C90F87A19F93F5" box="[911,1018,192,218]" direction="west" orientation="longitude" pageId="5" pageNumber="64" precision="925" value="-74.26667">74º16’W</geoCoordinate>
) in Boyacá Department.
<taxonomicName id="4C6F4D3EF717601D757F0F87A294922E" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF717601D757F0F87A294922E" italics="true" pageId="5" pageNumber="64">Atractus wagleri</emphasis>
</taxonomicName>
has been found in sub-Andean Tropical Forest at elevations of
<quantity id="4C979B58F717601D73910FAFA60E922D" box="[983,1131,232,258]" metricMagnitude="2" metricUnit="m" metricValue="9.7" metricValueMax="12.0" metricValueMin="7.4" pageId="5" pageNumber="64" unit="m" value="970.0" valueMax="1200.0" valueMin="740.0">740–1200 m</quantity>
(
<figureCitation id="13542A38F717601D743D0FAFA6A6922D" box="[1147,1219,232,258]" captionStart="FIGURE 5" captionStartId="5.[151,255,1291,1315]" captionTargetBox="[166,1436,561,1255]" captionTargetId="figure@5.[153,1438,551,1285]" captionTargetPageId="5" captionText="FIGURE 5. Geographical distribution of Atractus sanguineus and Atractus wagleri. Open symbols correspond to respective type localities of the species." httpUri="https://zenodo.org/record/274633/files/figure.png" pageId="5" pageNumber="64">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756536F717601D70830E48A0AC92DD" pageId="5" pageNumber="64" type="discussion">
<paragraph id="8BD036BDF717601D70830E48A0AC92DD" blockId="5.[151,1437,151,498]" pageId="5" pageNumber="64">
<emphasis id="B91BEAAFF717601D70830E48A3249206" bold="true" box="[197,321,271,297]" pageId="5" pageNumber="64">Remarks:</emphasis>
<bibRefCitation id="EFFE4B4CF717601D710B0E57A3899205" author="Prado" box="[333,492,272,298]" pageId="5" pageNumber="65" refString="Prado, A. (1945). Um novo Atractus de Colombia. Ciencia (Mexico), 6, 61." type="journal article" year="1945">Prado (1945)</bibRefCitation>
cited the
<typeStatus id="54D4881FF717601D722A0E57A0FA9205" box="[620,671,272,298]" pageId="5" pageNumber="64">type</typeStatus>
locality of
<taxonomicName id="4C6F4D3EF717601D73750E57A1D79206" box="[819,946,272,297]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="B91BEAAFF717601D73750E57A1D79206" box="[819,946,272,297]" italics="true" pageId="5" pageNumber="64">A. wagleri</emphasis>
</taxonomicName>
as Humbo (Boyacá),
<collectingCountry id="F378762DF717601D74830E57A7599205" box="[1221,1340,272,298]" name="Colombia" pageId="5" pageNumber="64">Colombia</collectingCountry>
. Subsequently, some authors cited Muzo as the proper
<typeStatus id="54D4881FF717601D729F0E7FA16E927D" box="[729,779,312,338]" pageId="5" pageNumber="64">type</typeStatus>
locality of the species (Medem, 1965; Pérez-Santos &amp; Moreno, 1998). This contradiction is probably because Humbo (
<geoCoordinate id="EE5B507AF717601D73870E27A64D9255" box="[961,1064,352,378]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="925" value="5.6">05º36’N</geoCoordinate>
,
<geoCoordinate id="EE5B507AF717601D747D0E27A6C09255" box="[1083,1189,352,378]" direction="west" orientation="longitude" pageId="5" pageNumber="64" precision="925" value="-74.26667">74º16’W</geoCoordinate>
) was a small town included originally in the boundaries of Muzo (
<geoCoordinate id="EE5B507AF717601D72960ECFA153928D" box="[720,822,392,418]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="925" value="5.5333333">05º32’N</geoCoordinate>
, 74º06) municipality. However, the municipality of Muzo was redefined and now does not contain Humbo, which is located instead within Quípama (
<geoCoordinate id="EE5B507AF717601D75770EF7A7F292E5" box="[1329,1431,432,458]" direction="north" orientation="latitude" pageId="5" pageNumber="64" precision="925" value="5.5166664">05º31’N</geoCoordinate>
,
<geoCoordinate id="EE5B507AF717601D70D10E9FA29A92DD" box="[151,255,472,498]" direction="west" orientation="longitude" pageId="5" pageNumber="64" precision="925" value="-74.183334">74º11’W</geoCoordinate>
) municipality (J. Lynch, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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