148 lines
17 KiB
XML
148 lines
17 KiB
XML
<document id="58C0B0B1C63E4EA28758486526BF003E" ID-DOI="10.11646/zootaxa.3994.3.5" ID-GBIF-Dataset="cec5741f-6463-4588-b543-45a7fe1e91d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="246106" ID-ZooBank="33D65E23-E4C9-4132-88A6-A082640C88C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461275584842" checkinUser="plazi" docAuthor="Fend, Steven V., Rodriguez, Pilar & Lenat, David R." docDate="2015" docId="03C6879CFFF3FFD2FF00F981B2B89A1D" docLanguage="en" docName="zt03994p424.pdf" docOrigin="Zootaxa 3994 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uktena Fend, Rodriguez & Lenat, 2015, n. gen." docType="treatment" docVersion="8" lastPageNumber="413" masterDocId="FFFFFFE4FFF2FFD0FF97FFA6B7249805" masterDocTitle="Uktena riparia n. gen., n. sp. (Annelida, Clitellata, Lumbriculidae), a new spermatophore-producing oligochaete" masterLastPageNumber="424" masterPageNumber="411" pageNumber="412" updateTime="1698612618969" updateUser="plazi">
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<mods:mods id="8D9200BA1A9FB513CA45AB679EFAADE2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B0CA7F83F3D65546210D388D8A3CCC46">
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<mods:title id="AE5461F260F8CB61390B36C4070C2241">Uktena riparia n. gen., n. sp. (Annelida, Clitellata, Lumbriculidae), a new spermatophore-producing oligochaete</mods:title>
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<mods:name id="4CADAAA79BE3F84D0F260C8A24BE5152" type="personal">
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<mods:role id="A3ED5D2659E54FC2ABBF19B75E7C45FE">
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<mods:namePart id="75C850C27A768F2478B7A98AC2F77273">Fend, Steven V.</mods:namePart>
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<mods:name id="A341A7F093F825645A2E9FB748DABFAD" type="personal">
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<mods:roleTerm id="2C3D246CE9C204537DD4B72504F296AB">Author</mods:roleTerm>
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<mods:namePart id="418277055438A850791A5ACEDFBFFCA3">Rodriguez, Pilar</mods:namePart>
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</mods:name>
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<mods:name id="514341A355B8053511CD9129A82F09D1" type="personal">
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<mods:role id="47AF285BE8D6499533AE2203D7A6CE89">
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<mods:roleTerm id="39FC936E9B52D94CD0C15F12639B6815">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50E409A6AC0C089B1CCBD8D2F1BEFF1E">Lenat, David R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="07441760BE7FBA3470D94A064FD87387">text</mods:typeOfResource>
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<mods:titleInfo id="357DDF89EC35FDB0CE7F2EC77927F88B">
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<mods:title id="C2C02BB28D8F51FBA5450945CB8254C3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8C662F9BEF9E7C7D87D20E172D34E452">
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<mods:date id="2D5E497697A65A63DDD69F2C64413D83">2015</mods:date>
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<mods:detail id="E1586D7180CC952D9DD0A2C2A7C7339E" type="volume">
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<mods:number id="44B9DA8468A20498AEEA758541E85015">3994</mods:number>
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</mods:detail>
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<mods:detail id="31008601FA11D400F02FCE4E7B3D52C6" type="issue">
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<mods:number id="493FD95A0B4DFA6EC95F432C562FC995">3</mods:number>
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<mods:start id="BFB192DDECE88DC2FA17DFDF0539874B">411</mods:start>
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<mods:end id="DE4C72DCEF3BD3BAECA96447E4E740D9">424</mods:end>
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<mods:classification id="3F57B2A0AFD3884CE07933E3EE88E337">journal article</mods:classification>
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<mods:identifier id="CDB902530AD6F27A646A766F46BB5C36" type="DOI">10.11646/zootaxa.3994.3.5</mods:identifier>
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<mods:identifier id="81342F0E2A3CB7DF8125A1D50F3E1165" type="GBIF-Dataset">cec5741f-6463-4588-b543-45a7fe1e91d9</mods:identifier>
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<treatment id="03C6879CFFF3FFD2FF00F981B2B89A1D" ID-DOI="http://doi.org/10.5281/zenodo.6103150" ID-GBIF-Taxon="119639283" ID-Zenodo-Dep="6103150" LSID="urn:lsid:plazi:treatment:03C6879CFFF3FFD2FF00F981B2B89A1D" httpUri="http://treatment.plazi.org/id/03C6879CFFF3FFD2FF00F981B2B89A1D" lastPageId="2" lastPageNumber="413" pageId="1" pageNumber="412">
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<subSubSection id="C3756501FFF3FFD1FF00F981B4AE9E82" pageId="1" pageNumber="412" type="nomenclature">
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<paragraph id="8BD0368AFFF3FFD1FF00F981B66C9E47" blockId="1.[151,328,1575,1602]" box="[151,328,1575,1602]" pageId="1" pageNumber="412">
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<heading id="D09881E6FFF3FFD1FF00F981B66C9E47" bold="true" box="[151,328,1575,1602]" fontSize="11" level="1" pageId="1" pageNumber="412" reason="1">
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<emphasis id="B91BEA98FFF3FFD1FF00F981B66C9E47" bold="true" box="[151,328,1575,1602]" pageId="1" pageNumber="412">
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<taxonomicName id="4C6F4D09FFF3FFD1FF00F981B7CB9E44" ID-CoL="84VK8" box="[151,239,1575,1601]" class="Clitellata" family="Lumbriculidae" genus="Uktena" kingdom="Animalia" order="Lumbriculida" pageId="1" pageNumber="412" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B91BEA98FFF3FFD1FF00F981B7CB9E44" bold="true" box="[151,239,1575,1601]" italics="true" pageId="1" pageNumber="412">Uktena</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22857E3FFF3FFD1FF61F98EB66C9E47" box="[246,328,1576,1602]" pageId="1" pageNumber="412" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD0368AFFF3FFD1FF00F9C8B4AE9E82" blockId="1.[151,906,1646,1671]" box="[151,906,1646,1671]" pageId="1" pageNumber="412">
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Included species:
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<taxonomicName id="4C6F4D09FFF3FFD1FEC8F9C8B5229E83" box="[351,518,1646,1671]" class="Clitellata" family="Lumbriculidae" genus="Uktena" kingdom="Animalia" order="Lumbriculida" pageId="1" pageNumber="412" phylum="Annelida" rank="species" species="riparia" status="sp. nov.">
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<emphasis id="B91BEA98FFF3FFD1FEC8F9C8B5229E83" box="[351,518,1646,1671]" italics="true" pageId="1" pageNumber="412">Uktena riparia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22857E3FFF3FFD1FD9AF9C8B5689E82" box="[525,588,1646,1671]" pageId="1" pageNumber="412" rank="species">
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<emphasis id="B91BEA98FFF3FFD1FD9AF9C8B5689E82" bold="true" box="[525,588,1646,1671]" pageId="1" pageNumber="412">n. sp.</emphasis>
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</taxonomicNameLabel>
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(
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<typeStatus id="54D48828FFF3FFD1FDCCF9C9B5A99E82" box="[603,653,1647,1671]" pageId="1" pageNumber="412">type</typeStatus>
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species by monotypy)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756501FFF3FFD1FF00F910B27B9FEB" pageId="1" pageNumber="412" type="diagnosis">
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<paragraph id="8BD0368AFFF3FFD1FF00F910B27B9FEB" blockId="1.[151,1436,1718,2031]" pageId="1" pageNumber="412">
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<emphasis id="B91BEA98FFF3FFD1FF00F910B62E9ECA" bold="true" box="[151,266,1718,1743]" pageId="1" pageNumber="412">Diagnosis</emphasis>
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(autapomorphies are underlined). Medium-sized to large worms with long, superficially-ringed proboscis. Chaetae simple-pointed. Reproductively-mature worms have several modified chaetae in ventral chaetal bundles of two postatrial segments. Two pairs of lateral blood vessels in posterior segments, with many blind branches. Nephridia begin on 10/11. One pair of testes in VIII, one pair of ovaries in IX. Male pores paired on VIII, in deep copulatory bursae with associated glands. Prosoporous atria elongate-cylindrical, ampulla covered with clusters of prostatic cells; ectal ducts produce tubular spermatophores. One pair of sperm funnels, extending back one or more segments in sperm sac. Vasa deferentia do not penetrate posterior septum (8/9). Female pores on posterior septum of first post-atrial segment. Spermathecae paired in atrial segment, anterior to atria; spermathecal pores within deep bursae containing a glandular, protrusible, copulatory organ for spermatophore attachment.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756501FFF0FFD2FF50FF31B2B89A1D" pageId="2" pageNumber="413" type="discussion">
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<paragraph id="8BD0368AFFF0FFD2FF50FF31B2B89A1D" blockId="2.[151,1436,151,536]" pageId="2" pageNumber="413">
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<emphasis id="B91BEA98FFF0FFD2FF50FF31B61F98B5" bold="true" box="[199,315,151,176]" pageId="2" pageNumber="413">Remarks.</emphasis>
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<taxonomicName id="4C6F4D09FFF0FFD2FED4FF31B6B098B5" box="[323,404,151,176]" class="Clitellata" family="Lumbriculidae" genus="Uktena" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B91BEA98FFF0FFD2FED4FF31B6B098B5" box="[323,404,151,176]" italics="true" pageId="2" pageNumber="413">Uktena</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22857E3FFF0FFD2FE0BFF3EB6CD98B5" box="[412,489,152,176]" pageId="2" pageNumber="413" rank="genus">n. gen.</taxonomicNameLabel>
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has three distinctive autapomorphies, all of which were previously unknown in the family
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<taxonomicName id="4C6F4D09FFF0FFD2FF7CFF1AB6B198D0" box="[235,405,188,213]" class="Clitellata" family="Lumbriculidae" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="family">Lumbriculidae</taxonomicName>
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: a unique arrangement of genital chaetae, a “copulatory organ” within a large, muscular spermathecal bursa, and tubular spermatophores (see Discussion). The presence of spermathecae in the atrial segment is unusual within the family, but this character is shared with at least three other lumbriculid genera, as well as two species of
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<taxonomicName id="4C6F4D09FFF0FFD2FE36FE81B5919945" authority="Piguet, 1913" authorityName="Piguet" authorityYear="1913" box="[417,693,295,320]" class="Clitellata" family="Dorydrilidae" genus="Dorydrilus" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FE36FE81B5389945" box="[417,540,295,320]" italics="true" pageId="2" pageNumber="413">Dorydrilus</emphasis>
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<bibRefCitation id="EFFE4B7BFFF0FFD2FDB2FE81B5919945" author="Piguet" box="[549,693,295,320]" pageId="2" pageNumber="413" refString="Piguet, E. (1913) Notes sur les Oligochetes. Revue Suisse de Zoologie, 21 (4), 111 - 146." type="journal article" year="1913">Piguet, 1913</bibRefCitation>
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</taxonomicName>
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(see Discussion). The combination of a prosoporous atrium and spermathecae in the male segment suggests that
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<taxonomicName id="4C6F4D09FFF0FFD2FD26FEEAB4269960" box="[689,770,332,357]" class="Clitellata" family="Lumbriculidae" genus="Uktena" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B91BEA98FFF0FFD2FD26FEEAB4269960" box="[689,770,332,357]" italics="true" pageId="2" pageNumber="413">Uktena</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="4C6F4D09FFF0FFD2FC71FEEAB2379960" authority="Altman, 1936" authorityName="Altman" authorityYear="1936" box="[998,1299,332,357]" class="Clitellata" family="Lumbriculidae" genus="Kincaidiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FC71FEEAB3559960" box="[998,1137,332,357]" italics="true" pageId="2" pageNumber="413">Kincaidiana</emphasis>
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<bibRefCitation id="EFFE4B7BFFF0FFD2FBE0FEEAB2379960" author="Altman" box="[1143,1299,332,357]" pageId="2" pageNumber="413" refString="Altman, L. C. (1936) Oligochaeta of Washington. University of Washington Publications in Biology, 4, 1 - 137." type="journal article" year="1936">Altman, 1936</bibRefCitation>
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</taxonomicName>
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, also a large worm with a filiform proboscis. However,
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<taxonomicName id="4C6F4D09FFF0FFD2FD0CFEC9B401998D" box="[667,805,367,392]" class="Clitellata" family="Lumbriculidae" genus="Kincaidiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FD0CFEC9B401998D" box="[667,805,367,392]" italics="true" pageId="2" pageNumber="413">Kincaidiana</emphasis>
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</taxonomicName>
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has the male pores in IX (instead of VIII) and has spermathecae also in postatrial segments.
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<taxonomicName id="4C6F4D09FFF0FFD2FDEDFE32B4CB99A9" authority="Michaelsen, 1933" authorityName="Michaelsen" authorityYear="1933" box="[634,1007,404,429]" class="Clitellata" family="Lumbriculidae" genus="Guestphalinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FDEDFE32B43899A8" box="[634,796,404,429]" italics="true" pageId="2" pageNumber="413">Guestphalinus</emphasis>
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<bibRefCitation id="EFFE4B7BFFF0FFD2FCB2FE32B4CB99A9" author="Michaelsen" box="[805,1007,404,429]" pageId="2" pageNumber="413" refString="Michaelsen, W. (1933) Uber Hohlen Oligochaten. Mitteilungen uber Hohlen- und Karstforschung, Berlin, 1, 1 - 19." type="journal article" year="1933">Michaelsen, 1933</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C6F4D09FFF0FFD2FBBCFE32B7EB99D5" authority="Rodriguez & Giani, 1987" authorityName="Rodriguez & Giani" authorityYear="1987" class="Clitellata" family="Lumbriculidae" genus="Cookidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FBBCFE32B39499A8" box="[1067,1200,404,429]" italics="true" pageId="2" pageNumber="413">Cookidrilus</emphasis>
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<bibRefCitation id="EFFE4B7BFFF0FFD2FB2EFE32B7EB99D5" author="Rodriguez" pageId="2" pageNumber="413" refString="Rodriguez, P. & Giani, N. (1987) Sur deux especes de Lumbriculidae (Oligochaeta) cavernicoles: Cookidrilus speluncaeus n. gen., sp. nov., et Trichodrilus diversisetosus Rodriguez, 1986. Bulletin de la Societe d'Histoire naturelle de Toulouse, 123, 45 - 49." type="journal article" year="1987">Rodriguez & Giani, 1987</bibRefCitation>
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</taxonomicName>
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are differentiated from
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<taxonomicName id="4C6F4D09FFF0FFD2FE7CFE11B51899D5" box="[491,572,439,464]" class="Clitellata" family="Lumbriculidae" genus="Uktena" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="B91BEA98FFF0FFD2FE7CFE11B51899D5" box="[491,572,439,464]" italics="true" pageId="2" pageNumber="413">Uktena</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6F4D09FFF0FFD2FDECFE11B42299D5" box="[635,774,439,464]" class="Clitellata" family="Lumbriculidae" genus="Kincaidiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FDECFE11B42299D5" box="[635,774,439,464]" italics="true" pageId="2" pageNumber="413">Kincaidiana</emphasis>
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</taxonomicName>
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by their semiprosoporous male ducts, as well as by the position of atria in IX and X, respectively. The three
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<taxonomicName id="4C6F4D09FFF0FFD2FD74FE7AB44399F0" box="[739,871,476,501]" class="Clitellata" family="Lumbriculidae" genus="Cookidrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="413" phylum="Annelida" rank="genus">
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<emphasis id="B91BEA98FFF0FFD2FD74FE7AB44399F0" box="[739,871,476,501]" italics="true" pageId="2" pageNumber="413">Cookidrilus</emphasis>
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</taxonomicName>
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species are small worms, without a proboscis, and with spermathecae also in post-atrial segments (
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<bibRefCitation id="EFFE4B7BFFF0FFD2FD3CFE59B4E09A1D" author="Rodriguez" box="[683,964,511,536]" pageId="2" pageNumber="413" refString="Rodriguez, P. & Giani, N. (1987) Sur deux especes de Lumbriculidae (Oligochaeta) cavernicoles: Cookidrilus speluncaeus n. gen., sp. nov., et Trichodrilus diversisetosus Rodriguez, 1986. Bulletin de la Societe d'Histoire naturelle de Toulouse, 123, 45 - 49." type="journal article" year="1987">Rodriguez & Giani 1987</bibRefCitation>
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,
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<bibRefCitation id="EFFE4B7BFFF0FFD2FC58FDA6B3B49A1D" author="Route" box="[975,1168,511,536]" pageId="2" pageNumber="413" refString="Route, N., Martinez-Ansemil, E., Sambugar, B. & Giani, N. (2004) On some interesting freshwater Annelida, mainly Oligochaeta, of the underground waters of southwestern France with description of Cookidrilus ruffoi n. sp. Subterranean Biology, 2, 1 - 5." type="journal article" year="2004">
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Route
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<emphasis id="B91BEA98FFF0FFD2FB8FFDA7B36B9A1D" box="[1048,1103,511,536]" italics="true" pageId="2" pageNumber="413">et al.</emphasis>
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2004
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</bibRefCitation>
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,
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<bibRefCitation id="EFFE4B7BFFF0FFD2FB0CFE59B2B59A1D" author="Rodriguez" box="[1179,1425,511,536]" pageId="2" pageNumber="413" refString="Rodriguez, P., Fend, S. V. & Lenat, D. R. (2014) Sylphella puccoon gen. n., sp. n. and two additional new species of aquatic oligochaetes (Lumbriculidae, Clitellata) from poorly-known lotic habitats in North Carolina (USA). ZooKeys, 451, 1 - 32. http: // dx. doi. org / 10.3897 / zookeys. 451.7304" type="journal article" year="2014">
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Rodriguez
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<emphasis id="B91BEA98FFF0FFD2FA81FDA7B2699A1D" box="[1302,1357,511,536]" italics="true" pageId="2" pageNumber="413">et al.</emphasis>
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2014
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |