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<document id="44D1EF388A3E60C72D62B5E983440CF9" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F075FFEFFF42FF75FC7AFE25" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Dendropaemon) aenigmaticus Genier &amp; Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="24" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="A2CB982AC7FFD80119FF9913E387A9B7">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:name id="C456A6A6E50772521E775E15DCAF4E0E" type="personal">
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<mods:namePart id="69878038992DAB2779FA0B424B76AA21">François Génier</mods:namePart>
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<mods:namePart id="E66DBD41FE6CC255C3180301A3E4A43B">Patrick Arnaud</mods:namePart>
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<mods:typeOfResource id="2839882E7A34741EB0E27101C0CCAFB9">text</mods:typeOfResource>
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<mods:title id="45878921C6FC79F919BD49DE7290A206">Zootaxa</mods:title>
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<mods:part id="82C4AEDFF0A63A8710E0919AE8B689E3">
<mods:date id="25EB691B861958B30C763906BC99A6B1">2016</mods:date>
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<mods:number id="EFD11850ABC5A04694B771DE7BC4D5D4">4099</mods:number>
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<mods:identifier id="5D536AF88002E995D70065A41398B2CB" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
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<treatment id="03C68782F075FFEFFF42FF75FC7AFE25" ID-DOI="http://doi.org/10.5281/zenodo.5780274" ID-GBIF-Taxon="127631940" ID-Zenodo-Dep="5780274" LSID="urn:lsid:plazi:treatment:03C68782F075FFEFFF42FF75FC7AFE25" httpUri="http://treatment.plazi.org/id/03C68782F075FFEFFF42FF75FC7AFE25" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C375651FF075FFE0FF42FF75FC07FF62" box="[151,993,152,178]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BD03694F075FFE0FF42FF75FC07FF62" blockId="23.[151,993,152,211]" box="[151,993,152,178]" pageId="23" pageNumber="24">
<heading id="D09881F8F075FFE0FF42FF75FC07FF62" bold="true" box="[151,993,152,178]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B91BEA86F075FFE0FF42FF75FC07FF62" bold="true" box="[151,993,152,178]" pageId="23" pageNumber="24">
11.
<taxonomicName id="4C6F4D17F075FFE0FF14FF75FCA2FF62" ID-CoL="7ZMZX" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[193,836,152,178]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="aenigmaticus" status="sp. nov." subGenus="Dendropaemon">
<emphasis id="B91BEA86F075FFE0FF14FF75FDBAFF62" bold="true" box="[193,604,152,178]" italics="true" pageId="23" pageNumber="24">Dendropaemon (D.) aenigmaticus</emphasis>
Génier &amp; Arnaud
</taxonomicName>
,
<taxonomicNameLabel id="A22857FDF075FFE0FC84FF75FC07FF62" box="[849,993,152,178]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF075FFE0FF42FF57FE23FF03" box="[151,453,186,211]" pageId="23" pageNumber="24" type="description">
<paragraph id="8BD03694F075FFE0FF42FF57FE23FF03" blockId="23.[151,993,152,211]" box="[151,453,186,211]" pageId="23" pageNumber="24">
(
<figureCitation id="13542A11F075FFE0FF4AFF57FF1DFF03" box="[159,251,186,211]" captionStart="FIGURES 7 12" captionStartId="100.[151,264,1966,1989]" captionTargetBox="[259,1332,208,1940]" captionTargetId="figure@100.[239,1347,193,1945]" captionTargetPageId="100" captionText="FIGURES 7 12. Dorsal habitus. 7. D. inflatus, female holotype; 8. D. pauliani, female holotype; 9. D. pilosissimus, female holotype; 10. D. renatii, female holotype; 11. D. aenigmaticus, male holotype; 12. D. amazonicus, male holotype." httpUri="https://zenodo.org/record/268276/files/figure.png" pageId="23" pageNumber="24">Figs. 11</figureCitation>
, 5455, 112, 157)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF075FFE0FF42FEEFFB54FECB" box="[151,1202,258,283]" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BD03694F075FFE0FF42FEEFFB54FECB" blockId="23.[151,1437,258,2011]" box="[151,1202,258,283]" pageId="23" pageNumber="24">
<emphasis id="B91BEA86F075FFE0FF42FEEFFECBFECB" bold="true" box="[151,301,258,283]" pageId="23" pageNumber="24">
<typeStatus id="54D48836F075FFE0FF42FEEFFF34FECB" box="[151,210,258,283]" pageId="23" pageNumber="24">Type</typeStatus>
locality
</emphasis>
. PK 125, Route N2 (Cayenne-Saint-Georges-de-LOyapock),
<collectingCountry id="F3787604F075FFE0FC32FEEEFBD8FECB" box="[999,1086,259,283]" name="French Guiana" pageId="23" pageNumber="24">Guyane</collectingCountry>
française.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF075FFE0FF12FECBFA8DFDC7" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BD03694F075FFE0FF12FECBFA8DFDC7" blockId="23.[151,1437,258,2011]" pageId="23" pageNumber="24">
<emphasis id="B91BEA86F075FFE0FF12FECBFEA7FEEE" bold="true" box="[199,321,294,319]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
The two-segmented meso- and metatarsi of which the first segment is approximately 3 times as long as wide at apex, the completely glossy black body and the clypeal teeth lacking emargination laterally will place this species in the
<taxonomicName id="4C6F4D17F075FFE0FE7BFE82FE13FE56" box="[430,501,367,390]" class="Insecta" family="Scarabaeidae" genus="Eurysternus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B91BEA86F075FFE0FE7BFE82FE13FE56" box="[430,501,367,390]" italics="true" pageId="23" pageNumber="24">piceus</emphasis>
</taxonomicName>
complex. From closely related species it can be separated from
<taxonomicName id="4C6F4D17F075FFE0FB01FE82FA97FE57" box="[1236,1393,366,391]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="angustulus">
<emphasis id="B91BEA86F075FFE0FB01FE82FA97FE57" box="[1236,1393,366,391]" italics="true" pageId="23" pageNumber="24">D. angustulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D17F075FFE0FA54FE82FF03FE7B" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="larseni">
<emphasis id="B91BEA86F075FFE0FA54FE82FF03FE7B" italics="true" pageId="23" pageNumber="24">D. larseni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D17F075FFE0FF20FE79FE90FE7B" box="[245,374,402,427]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="telephus">
<emphasis id="B91BEA86F075FFE0FF20FE79FE90FE7B" box="[245,374,402,427]" italics="true" pageId="23" pageNumber="24">D. telephus</emphasis>
</taxonomicName>
by its lager body size and more robust shape and dorsoventrally compressed body; from
<taxonomicName id="4C6F4D17F075FFE0FA54FE79FEFAFE1F" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="flechtmanni">
<emphasis id="B91BEA86F075FFE0FA54FE79FEFAFE1F" italics="true" pageId="23" pageNumber="24">D. flechtmanni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D17F075FFE0FE8DFE5AFDE4FE1F" box="[344,514,438,463]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B91BEA86F075FFE0FE8DFE5AFDE4FE1F" box="[344,514,438,463]" italics="true" pageId="23" pageNumber="24">D. vazdemelloi</emphasis>
</taxonomicName>
by its much finer elytral striae 14; from
<taxonomicName id="4C6F4D17F075FFE0FC22FE5AFBAFFE1E" box="[1015,1097,439,462]" class="Insecta" family="Scarabaeidae" genus="Enicotarsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91BEA86F075FFE0FC22FE5AFBAFFE1E" box="[1015,1097,439,462]" italics="true" pageId="23" pageNumber="24">D. ater</emphasis>
</taxonomicName>
by its flat posterior protibial surface and nearly semicircular clypeal edge. The more slender and gradually tapering toward apex meso and metatibia will separate it from
<taxonomicName id="4C6F4D17F075FFE0FE3AFE12FDBEFDC6" box="[495,600,511,534]" class="Insecta" family="Scarabaeidae" genus="Eurysternus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="B91BEA86F075FFE0FE3AFE12FDBEFDC6" box="[495,600,511,534]" italics="true" pageId="23" pageNumber="24">D. piceus</emphasis>
</taxonomicName>
, from which it also differs by its only minutely punctate elytral striae.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF075FFEFFF12FDCFFE26FF28" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="8BD03694F075FFE0FF12FDCFFD6BF867" blockId="23.[151,1437,258,2011]" pageId="23" pageNumber="24">
<emphasis id="B91BEA86F075FFE0FF12FDCFFEB7FDEB" bold="true" box="[199,337,546,571]" pageId="23" pageNumber="24">Description</emphasis>
.
<collectingCountry id="F3787604F075FFE0FEB7FDCFFE7AFDEB" box="[354,412,546,571]" name="Malta" pageId="23" pageNumber="24">Male</collectingCountry>
<typeStatus id="54D48836F075FFE0FE73FDCFFDEEFDEB" box="[422,520,546,571]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F075FFE0FDCFFDCFFD89FDEB" box="[538,623,546,571]" captionStart="FIGURES 7 12" captionStartId="100.[151,264,1966,1989]" captionTargetBox="[259,1332,208,1940]" captionTargetId="figure@100.[239,1347,193,1945]" captionTargetPageId="100" captionText="FIGURES 7 12. Dorsal habitus. 7. D. inflatus, female holotype; 8. D. pauliani, female holotype; 9. D. pilosissimus, female holotype; 10. D. renatii, female holotype; 11. D. aenigmaticus, male holotype; 12. D. amazonicus, male holotype." httpUri="https://zenodo.org/record/268276/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
).
<emphasis id="B91BEA86F075FFE0FD52FDCFFD22FDEB" bold="true" box="[647,708,546,571]" pageId="23" pageNumber="24">Body</emphasis>
. Body moderately large, length
<quantity id="4C979B71F075FFE0FBE9FDCFFB43FDEA" box="[1084,1189,546,571]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="23" pageNumber="24" unit="mm" value="14.5">14.5 mm</quantity>
, maximum width 8.0 mm; body subrectangular in dorsal view; dorsum largely flat.
<emphasis id="B91BEA86F075FFE0FC9AFDABFC76FD8F" bold="true" box="[847,912,582,607]" pageId="23" pageNumber="24">Color</emphasis>
. Dorsal surface black, glossy, lacking metallic sheen; ventrum black; pygidium black; legs black.
<emphasis id="B91BEA86F075FFE0FD22FD87FCD0FD53" bold="true" box="[759,822,618,643]" pageId="23" pageNumber="24">Head</emphasis>
. Clypeus narrowly arcuate, anterior portion slightly upturned; clypeal teeth triangular; clypeal median emargination v-shaped, clypeal edge lacking emargination on external side of each clypeal tooth, clypeal teeth ventral surface with a well-defined v-shaped carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface simply punctate, with a long and acute transverse carina; clypeofrontal carina low, more than 6 times wider than high, straight in dorsal view, simply carinate, clypeofrontal carina apical edge slightly sinuous medially in frontal view; eyes small in dorsal view, interocular ratio 4.4.
<emphasis id="B91BEA86F075FFE0FEDBFC8BFE61FCAF" bold="true" box="[270,391,870,895]" pageId="23" pageNumber="24">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.5; disc of pronotum minutely punctate throughout, with a fine ill-defined longitudinal sulcus on posterior half; pronotal anterior margin only slightly wider and flat lateral to eye; anterior portion with a broad and fine tectiform carina tuberculate medially; anterior angles surface simply punctate, similar to lateral margin along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions slightly explanate; pronotal basal fossae ill-defined, moderately concave; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F075FFE0FBE3FBF7FB64FBE3" bold="true" box="[1078,1154,1050,1075]" pageId="23" pageNumber="24">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.1; elytral base lacking distinct margin, simply convex; elytral striae 14 fine on disc, finer and lacking minute carina laterally on apical declivity, evenly impressed throughout, elytral striae 5 atrophied, lacking fine carina on each side on disc, strial punctures minute, stria 1 well-defined apically, connecting to marginal stria; interstriae flat, minutely punctate throughout, surface glossy.
<emphasis id="B91BEA86F075FFE0FF3FFB23FE4EFB37" bold="true" box="[234,424,1230,1255]" pageId="23" pageNumber="24">Thoracic sterna</emphasis>
. Proepisternal carina reduced, present along coxal insertion only; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, keel-shaped.
<emphasis id="B91BEA86F075FFE0FC2DFB1FFBD6FADB" bold="true" box="[1016,1072,1266,1291]" pageId="23" pageNumber="24">Legs</emphasis>
. Profemur posterior surface flat and glabrous internally, posterointernal margin rather thick, evenly developed, internal edge rather narrow, with a brush of long setae along anterointernal edge, remaining surface with irregular ill-defined punctures and glossy. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some ill-defined irregular punctures externally to median carina, surface finely and irregularly microsculptured between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather slender, gradually widening toward apex in anterior view; anteroapical edge straight in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge carinate, sharp. Mesotarsus similar in shape to metatarsus, 2-segmented, first segment elongate, more than three times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on apical half. Metatibia rather slender, gradually widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with transverse microsculpture. Metatarsus 2-segmented (
<figureCitation id="13542A11F075FFE0FF4AF8E3FEE5F8F7" box="[159,259,1806,1831]" captionStart="FIGURES 110 117" captionStartId="112.[151,264,1933,1956]" captionTargetBox="[219,1362,202,1901]" captionTargetId="figure@112.[212,1375,193,1913]" captionTargetPageId="112" captionText="FIGURES 110 117. Metatarsus, ventral view. 110. D. bluti; 111. D. carinifer; 112. D. aenigmaticus; 113. D. amazonicus; 114. D. angustulus; 115. D. viridis; 116. D. viridipennis; 117. D. convexus." httpUri="https://zenodo.org/record/268282/files/figure.png" pageId="23" pageNumber="24">Fig. 112</figureCitation>
), first segment elongate, more than three time as long as wide at apex, with anterointernal carina illdefined.
<emphasis id="B91BEA86F075FFE0FF29F8DFFE16F89B" bold="true" box="[252,496,1842,1867]" pageId="23" pageNumber="24">Abdominal sternites</emphasis>
. Sternites 36 longitudinally flat; sternites 46 with 13 unaligned rows of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium minutely punctate on disc.
<emphasis id="B91BEA86F075FFE0FD48F897FCAEF843" bold="true" box="[669,840,1914,1939]" pageId="23" pageNumber="24">
<collectingCountry id="F3787604F075FFE0FD48F897FD3CF843" box="[669,730,1914,1939]" name="Malta" pageId="23" pageNumber="24">Male</collectingCountry>
genitalia
</emphasis>
(Figs. 5455). Parameres simply rounded apically in dorsal view; surface smooth, glossy apically.
</paragraph>
<paragraph id="8BD03694F075FFE0FF12F82FFD54F80A" blockId="23.[151,1437,258,2011]" box="[199,690,1986,2011]" pageId="23" pageNumber="24">
<emphasis id="B91BEA86F075FFE0FF12F82FFE90F80B" bold="true" box="[199,374,1986,2011]" pageId="23" pageNumber="24">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF075FFE0FE5DF82EFE35F80B" box="[392,467,1986,2011]" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
). Length:
<quantity id="4C979B71F075FFE0FD92F82FFD48F80A" box="[583,686,1986,2011]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="23" pageNumber="24" unit="mm" value="14.5">14.5 mm</quantity>
.
</paragraph>
<paragraph id="8BD03694F07AFFEFFF12FF7AFE26FF28" blockId="24.[151,1436,151,501]" pageId="24" pageNumber="25">
<emphasis id="B91BEA86F07AFFEFFF12FF7AFE55FF60" bold="true" box="[199,435,151,176]" pageId="24" pageNumber="25">
Primary
<typeStatus id="54D48836F07AFFEFFEEEFF7AFE88FF60" box="[315,366,151,176]" pageId="24" pageNumber="25">type</typeStatus>
data
</emphasis>
.
<typeStatus id="54D48836F07AFFEFFE1CFF7AFDD7FF60" box="[457,561,151,176]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
male (CPFA): [
<date id="FFD11054F07AFFEFFD2FFF75FC9BFF60" box="[762,893,152,176]" pageId="24" pageNumber="25" value="2007-12-24">24/12/2007</date>
PI/ rt st georges PK125/ JL GIUGLARIS]; WORLD/ SCARAB./ DATABASE/ WSD00017631]; [
<typeStatus id="54D48836F07AFFEFFCD2FF51FC7BFF04" box="[775,925,188,212]" pageId="24" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6F4D17F07AFFEFFC7AFF51FBBDFF05" box="[943,1115,188,213]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F07AFFEFFBB8FF51FAE7FF05" box="[1133,1281,188,213]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="aenigmaticus" status="sp. nov.">aenigmaticus</taxonomicName>
<taxonomicNameLabel id="A22857FDF07AFFEFFADFFF51FADBFF04" box="[1290,1341,188,212]" pageId="24" pageNumber="25" rank="species">n.sp.</taxonomicNameLabel>
/ Génier &amp; Arnaud, 2014] red card.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF07AFFEFFF12FEE9FD61FECD" box="[199,647,260,285]" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BD03694F07AFFEFFF12FEE9FD61FECD" blockId="24.[151,1436,151,501]" box="[199,647,260,285]" pageId="24" pageNumber="25">
<emphasis id="B91BEA86F07AFFEFFF12FEE9FE4FFECD" bold="true" box="[199,425,260,285]" pageId="24" pageNumber="25">Material examined</emphasis>
. Primary
<typeStatus id="54D48836F07AFFEFFDCDFEE8FDAEFECD" box="[536,584,261,285]" pageId="24" pageNumber="25">type</typeStatus>
only.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF07AFFEFFF12FECAFDE4FE58" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BD03694F07AFFEFFF12FECAFC09FEB4" blockId="24.[151,1436,151,501]" pageId="24" pageNumber="25">
<emphasis id="B91BEA86F07AFFEFFF12FECAFEA1FE90" bold="true" box="[199,327,295,320]" pageId="24" pageNumber="25">Etymology</emphasis>
.
<taxonomicName id="4C6F4D17F07AFFEFFE8CFEC5FE17FEEF" box="[345,497,296,319]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="118" phylum="Arthropoda" rank="species" species="aenigmaticus">
<emphasis id="B91BEA86F07AFFEFFE8CFEC5FE17FEEF" box="[345,497,296,319]" italics="true" pageId="24" pageNumber="25">Aenigmaticus</emphasis>
</taxonomicName>
, an adjective relating to the status of this species known only by the
<typeStatus id="54D48836F07AFFEFFAEFFECAFA7AFE90" box="[1338,1436,295,320]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
despite extensive collecting in
<collectingCountry id="F3787604F07AFFEFFE25FEA1FD70FEB5" box="[496,662,332,357]" name="French Guiana" pageId="24" pageNumber="25">French Guiana</collectingCountry>
using flight interception traps.
</paragraph>
<paragraph id="8BD03694F07AFFEFFF12FE82FDE4FE58" blockId="24.[151,1436,151,501]" box="[199,514,367,392]" pageId="24" pageNumber="25">
<emphasis id="B91BEA86F07AFFEFFF12FE82FE9BFE58" bold="true" box="[199,381,367,392]" pageId="24" pageNumber="25">Natural history</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF07AFFEFFF12FE79FC7AFE25" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BD03694F07AFFEFFF12FE79FD43FE7D" blockId="24.[151,1436,151,501]" box="[199,677,404,429]" pageId="24" pageNumber="25">
<emphasis id="B91BEA86F07AFFEFFF12FE79FED2FE7D" bold="true" box="[199,308,404,429]" pageId="24" pageNumber="25">Remarks</emphasis>
. Female and variation unknown.
</paragraph>
<paragraph id="8BD03694F07AFFEFFF12FE55FC7AFE25" blockId="24.[151,1436,151,501]" pageId="24" pageNumber="25">
At the moment we feel confident that this individual represents a new species, especially as the sclerites of the internal sac also differs from
<taxonomicName id="4C6F4D17F07AFFEFFE0AFE30FDCBFE24" box="[479,557,477,500]" class="Insecta" family="Scarabaeidae" genus="Enicotarsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91BEA86F07AFFEFFE0AFE30FDCBFE24" box="[479,557,477,500]" italics="true" pageId="24" pageNumber="25">D. ater</emphasis>
</taxonomicName>
, its most closely related species.
</paragraph>
</subSubSection>
</treatment>
</document>