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<document id="8069C4C99C205737BA4D21DDDA8BF95F" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F073FFE4FF42F96DFDC9FDED" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Coprophanaeoides) pauliani Martinez &amp; Pereira 1960" docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="18" updateTime="1698234910999" updateUser="plazi">
<mods:mods id="75F41CE88EF39AAF210E7BFE902AA02E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2E281D68EBE2D0800C2E5ADFF542DF23">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
</mods:titleInfo>
<mods:name id="6E24A03D8C088F229851A2D2B331A8B5" type="personal">
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<mods:roleTerm id="C036A8EA89A1592BC9AC608B24B6375B">Author</mods:roleTerm>
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<mods:namePart id="DD25D484512065D7964F1881432E6AC3">François Génier</mods:namePart>
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<mods:namePart id="16F8AF0B16A363B9507F94E2C482EAAC">Patrick Arnaud</mods:namePart>
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<mods:typeOfResource id="CAC7C2B30140ABDBDAFA64D899D11270">text</mods:typeOfResource>
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<mods:title id="3B8ADD1BAD4CE6D795181C1576391C80">Zootaxa</mods:title>
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<mods:part id="66659ABB13AFE27F372115FF660DA850">
<mods:date id="52F89E7D0B60EF2802EA89CD5FDF3527">2016</mods:date>
<mods:detail id="F57614CD0E35F2162A66224DBB96B38F" type="volume">
<mods:number id="E0617F7620FA9CE5D8C8237AD327480E">4099</mods:number>
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<mods:detail id="42731AC351CAF9638BBB1B4561DC6766" type="issue">
<mods:number id="A03C08229C33C371E4E31A7BCDECDEF1">1</mods:number>
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<mods:classification id="E5E0AC8DD80304B0BB94373E25F094E8">journal volume</mods:classification>
<mods:identifier id="DE7B69534AF977C07C0516B382FFC290" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
<mods:identifier id="781D561690AD53478E737FED0F7AE889" type="GBIF-Dataset">5d7eab55-66e2-43ed-b764-bac0c6e5b6d6</mods:identifier>
<mods:identifier id="7C60EB08CF743F1295E6863929D4C3CF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8F049ABF9C106000D3A8AD39F9DF1826" type="Zenodo-Dep">268274</mods:identifier>
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<treatment id="03C68782F073FFE4FF42F96DFDC9FDED" ID-DOI="http://doi.org/10.5281/zenodo.5780266" ID-GBIF-Taxon="127631981" ID-Zenodo-Dep="5780266" LSID="urn:lsid:plazi:treatment:03C68782F073FFE4FF42F96DFDC9FDED" httpUri="http://treatment.plazi.org/id/03C68782F073FFE4FF42F96DFDC9FDED" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C375651FF073FFE6FF42F96DFBC7F94A" box="[151,1057,1664,1690]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BD03694F073FFE6FF42F96DFBC7F94A" blockId="17.[151,1057,1664,1723]" box="[151,1057,1664,1690]" pageId="17" pageNumber="18">
<heading id="D09881F8F073FFE6FF42F96DFBC7F94A" bold="true" box="[151,1057,1664,1690]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B91BEA86F073FFE6FF42F96DFBC7F94A" bold="true" box="[151,1057,1664,1690]" pageId="17" pageNumber="18">
8.
<taxonomicName id="4C6F4D17F073FFE6FF61F96DFBC7F94A" ID-CoL="34R2G" authority="Martinez &amp; Pereira, 1960" authorityName="Martinez &amp; Pereira" authorityYear="1960" box="[180,1057,1664,1690]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pauliani" subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F073FFE6FF61F96DFD3FF94A" bold="true" box="[180,729,1664,1690]" italics="true" pageId="17" pageNumber="18">Dendropaemon (Coprophanaeoides) pauliani</emphasis>
<bibRefCitation id="EFFE4B65F073FFE6FD35F96DFBC7F94A" author="Martinez" box="[736,1057,1664,1690]" pageId="17" pageNumber="18" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">Martínez &amp; Pereira, 1960</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF073FFE6FF42F94FFE80F96B" box="[151,358,1698,1723]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BD03694F073FFE6FF42F94FFE80F96B" blockId="17.[151,1057,1664,1723]" box="[151,358,1698,1723]" pageId="17" pageNumber="18">
(
<figureCitation id="13542A11F073FFE6FF4AF94FFF0BF96B" box="[159,237,1698,1723]" captionStart="FIGURES 7 – 12" captionStartId="100.[151,264,1966,1989]" captionTargetBox="[259,1332,208,1940]" captionTargetId="figure@100.[239,1347,193,1945]" captionTargetPageId="100" captionText="FIGURES 7 – 12. Dorsal habitus. 7. D. inflatus, female holotype; 8. D. pauliani, female holotype; 9. D. pilosissimus, female holotype; 10. D. renatii, female holotype; 11. D. aenigmaticus, male holotype; 12. D. amazonicus, male holotype." httpUri="https://zenodo.org/record/268276/files/figure.png" pageId="17" pageNumber="18">Figs. 8</figureCitation>
,
<figureCitation id="13542A11F073FFE6FF2FF94FFEC0F96B" box="[250,294,1698,1723]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="17" pageNumber="18">135</figureCitation>
,
<figureCitation id="13542A11F073FFE6FEE7F94FFEBBF96B" box="[306,349,1698,1723]" captionStart="FIGURE 156" captionStartId="116.[151,250,1237,1259]" captionTargetBox="[150,1436,193,1215]" captionTargetId="figure@116.[150,1436,193,1215]" captionTargetPageId="116" captionText="FIGURE 156. Known distribution of D. bluti, D. carinifer, D. compressipennis, D. cribrosus, D. furtadoi, D. hirticollis, D. inflatus, D. pauliani, D. pilosissimus, D. renatii." httpUri="https://zenodo.org/record/268286/files/figure.png" pageId="17" pageNumber="18">156</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF073FFE6FF42F907FC2FF8CF" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BD03694F073FFE6FF42F907FC2FF8CF" blockId="17.[151,1207,1770,1823]" pageId="17" pageNumber="18">
<taxonomicName id="4C6F4D17F073FFE6FF42F907FD77F92F" ID-CoL="34R2G" authority="Martinez &amp; Pereira 1960" authorityName="Martinez &amp; Pereira" authorityYear="1960" box="[151,657,1770,1792]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B91BEA86F073FFE6FF42F907FE69F8D0" box="[151,399,1770,1792]" italics="true" pageId="17" pageNumber="18">Dendropaemon pauliani</emphasis>
<bibRefCitation id="EFFE4B65F073FFE6FE41F907FD77F92F" author="Martinez" box="[404,657,1770,1792]" pageId="17" pageNumber="18" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">
Martínez &amp; Pereira
<emphasis id="B91BEA86F073FFE6FD8BF907FD77F92F" bold="true" box="[606,657,1770,1791]" pageId="17" pageNumber="18">1960</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F073FFE6FD48F906FC72F8D0" box="[669,916,1771,1792]" italics="true" pageId="17" pageNumber="18">
Rev. Soc. Ent.
<collectingCountry id="F3787604F073FFE6FCFAF906FC72F8D0" box="[815,916,1771,1792]" name="Argentina" pageId="17" pageNumber="18">Argentina</collectingCountry>
</emphasis>
22: 81 (original description)
<taxonomicName id="4C6F4D17F073FFE6FF42F8E4FD78F8CE" ID-CoL="34R2G" authority="Hamel-Leigue et al. 2009" authorityName="Hamel-Leigue et al." authorityYear="2009" box="[151,670,1801,1823]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B91BEA86F073FFE6FF42F8E4FE68F8CF" box="[151,398,1801,1823]" italics="true" pageId="17" pageNumber="18">Dendropaemon pauliani</emphasis>
:
<bibRefCitation id="EFFE4B65F073FFE6FE4EF8E4FD78F8CE" author="Hamel-Leigue" box="[411,670,1801,1823]" pageId="17" pageNumber="18" refString="Hamel-Leigue, A. C., Herzog, S. K., Mann, D. J., Larsen, T. H., Gill, B. D., Edmonds, W. D. &amp; Spector, S. (2009) Distribucion e historia natural de Escarabajos coprofagos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana, 5 (2), 43 - 95." type="journal article" year="2009">
Hamel-Leigue et al.
<emphasis id="B91BEA86F073FFE6FDB9F8E4FD78F8CE" bold="true" box="[620,670,1801,1822]" pageId="17" pageNumber="18">2009</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F073FFE6FD7EF8E4FCC7F8CF" box="[683,801,1801,1823]" italics="true" pageId="17" pageNumber="18">Kempffiana</emphasis>
5: 49 (faunistic)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF073FFE6FF42F8A1FBDDF8B5" box="[151,1083,1868,1893]" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BD03694F073FFE6FF42F8A1FBDDF8B5" blockId="17.[151,1436,1868,2037]" box="[151,1083,1868,1893]" pageId="17" pageNumber="18">
<emphasis id="B91BEA86F073FFE6FF42F8A1FECBF8B5" bold="true" box="[151,301,1868,1893]" pageId="17" pageNumber="18">
<typeStatus id="54D48836F073FFE6FF42F8A1FF34F8B5" box="[151,210,1868,1893]" pageId="17" pageNumber="18">Type</typeStatus>
locality
</emphasis>
. Parapeti, provincia Cordillera, departamento de Santa Cruz,
<collectingCountry id="F3787604F073FFE6FC35F8A1FBD0F8B5" box="[992,1078,1868,1893]" name="Bolivia" pageId="17" pageNumber="18">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF073FFE6FF12F882FE6BF825" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BD03694F073FFE6FF12F882FE6BF825" blockId="17.[151,1436,1868,2037]" pageId="17" pageNumber="18">
<emphasis id="B91BEA86F073FFE6FF12F882FEA7F857" bold="true" box="[199,321,1903,1928]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
The acutely notched external side of each clypeal tooth will place this species in
<taxonomicName id="4C6F4D17F073FFE6FF42F879FE8DF87D" box="[151,363,1940,1965]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F073FFE6FF42F879FE8DF87D" box="[151,363,1940,1965]" italics="true" pageId="17" pageNumber="18">Coprophanaeoides</emphasis>
</taxonomicName>
. Within
<taxonomicName id="4C6F4D17F073FFE6FE05F879FD42F87D" box="[464,676,1940,1965]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F073FFE6FE05F879FD42F87D" box="[464,676,1940,1965]" italics="true" pageId="17" pageNumber="18">Coprophanaeoides</emphasis>
</taxonomicName>
, the glabrous clypeus and pronotum will separate it from all other species except
<taxonomicName id="4C6F4D17F073FFE6FE95F854FE70F801" box="[320,406,1976,2001]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bluti">
<emphasis id="B91BEA86F073FFE6FE95F854FE70F801" box="[320,406,1976,2001]" italics="true" pageId="17" pageNumber="18">D. bluti</emphasis>
</taxonomicName>
, which has smaller eyes,
<taxonomicName id="4C6F4D17F073FFE6FD6DF854FCC9F801" box="[696,815,1976,2001]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B91BEA86F073FFE6FD6DF854FCC9F801" box="[696,815,1976,2001]" italics="true" pageId="17" pageNumber="18">D. inflatus</emphasis>
</taxonomicName>
which has atrophied elytral striae and
<taxonomicName id="4C6F4D17F073FFE6FB31F854FAA9F800" box="[1252,1359,1977,2000]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F073FFE6FB31F854FAA9F800" box="[1252,1359,1977,2000]" italics="true" pageId="17" pageNumber="18">D. renatii</emphasis>
</taxonomicName>
which is known from
<collectingCountry id="F3787604F073FFE6FE94F831FE6FF825" box="[321,393,2012,2037]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF070FFE5FF12FF7AFDE8F845" pageId="18" pageNumber="19" type="description">
<paragraph id="8BD03694F070FFE5FF12FF7AFC6DF96D" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B91BEA86F070FFE5FF12FF7AFEB7FF60" bold="true" box="[199,337,151,176]" pageId="18" pageNumber="19">Description</emphasis>
. Female
<typeStatus id="54D48836F070FFE5FE68FF7AFDF9FF60" box="[445,543,151,176]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F070FFE5FDE4FF7AFD91FF60" box="[561,631,151,176]" captionStart="FIGURES 7 – 12" captionStartId="100.[151,264,1966,1989]" captionTargetBox="[259,1332,208,1940]" captionTargetId="figure@100.[239,1347,193,1945]" captionTargetPageId="100" captionText="FIGURES 7 – 12. Dorsal habitus. 7. D. inflatus, female holotype; 8. D. pauliani, female holotype; 9. D. pilosissimus, female holotype; 10. D. renatii, female holotype; 11. D. aenigmaticus, male holotype; 12. D. amazonicus, male holotype." httpUri="https://zenodo.org/record/268276/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
).
<emphasis id="B91BEA86F070FFE5FD5BFF7AFD2DFF60" bold="true" box="[654,715,151,176]" pageId="18" pageNumber="19">Body</emphasis>
. Body moderately large, length 12.0 mm, maximum width 7.0 mm; body subrectangular in dorsal view; dorsum narrowly flat.
<emphasis id="B91BEA86F070FFE5FCB3FF51FC41FF05" bold="true" box="[870,935,188,213]" pageId="18" pageNumber="19">Color</emphasis>
. Dorsal surface dark brown to black, glossy, with blue and green metallic sheen; head with faint bluish and greenish metallic sheen on posterior portion of clypeus, genae and frons; pronotum with faint bluish and greenish metallic sheen on anterior and lateral declivities; elytra with faint bluish metallic sheen; ventrum with faint greenish and coppery metallic sheen; pygidium with faint bluish metallic sheen; legs light to dark brown, with faint greenish metallic sheen.
<emphasis id="B91BEA86F070FFE5FB44FEA1FB37FEB5" bold="true" box="[1169,1233,332,357]" pageId="18" pageNumber="19">Head</emphasis>
. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth ogival; clypeal median emargination v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, bordered posteriorly by a more or less regular row of punctures, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with fine sharp tubercle and rugulae on disc, lacking distinct transverse carina, simply convex; clypeofrontal carina low, more than 6 times wider than high, straight in dorsal view, simply carinate, clypeofrontal carina apical edge broadly arcuate in frontal view; eyes moderately large in dorsal view, interocular ratio 3.3.
<emphasis id="B91BEA86F070FFE5FCA9FDAAFC12FDB0" bold="true" box="[892,1012,583,608]" pageId="18" pageNumber="19">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.6; disc of pronotum smooth basally, with transverse rugulae anteriorly, with a fine illdefined longitudinal sulcus on posterior half; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a simple anteriorly arcuate carina with lateral extremities connecting to anterior margin; anterior angles surface with more or less rough and irregular fine tubercles, similar to lateral margin along posterior edge of anterior margin; lateral fossae oval, bordered laterally by a sharp carina and anteriorly by a blunt tubercle; lateral portions strongly explanate; pronotal basal fossae absent; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F070FFE5FF08FCA9FECFFC8D" bold="true" box="[221,297,836,861]" pageId="18" pageNumber="19">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.1; elytral base distinctly marginate; elytral striae 1–4 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures minute, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae moderately convex, minutely punctate and with some additional ill-defined irregular punctures along striae, surface glossy.
<emphasis id="B91BEA86F070FFE5FAE4FC39FF04FBC0" bold="true" pageId="18" pageNumber="19">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, y-shaped.
<emphasis id="B91BEA86F070FFE5FE70FBF1FE3BFBE5" bold="true" box="[421,477,1052,1077]" pageId="18" pageNumber="19">Legs</emphasis>
. Profemur posterior surface flat and glabrous internally, posterointernal margin rather thin, uneven, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured; posterior surface with some ill-defined rugose punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge straight in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia rather slender, gradually widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with ill-defined microsculpture. Metatarsus 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and reaching apical edge.
<emphasis id="B91BEA86F070FFE5FE47F9B1FD60F9A5" bold="true" box="[402,646,1628,1653]" pageId="18" pageNumber="19">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with more than three unaligned row of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium minutely punctate on disc.
</paragraph>
<paragraph id="8BD03694F070FFE5FF12F92AFD2EF90F" blockId="18.[151,1437,151,2013]" box="[199,712,1735,1760]" pageId="18" pageNumber="19">
<emphasis id="B91BEA86F070FFE5FF12F92AFE90F930" bold="true" box="[199,374,1735,1760]" pageId="18" pageNumber="19">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF070FFE5FE5DF925FE0FF930" box="[392,489,1735,1760]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
). Length: 12.0 mm.
</paragraph>
<paragraph id="8BD03694F070FFE5FF12F901FDE8F845" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="19">
<emphasis id="B91BEA86F070FFE5FF12F901FE47F8D5" bold="true" box="[199,417,1772,1797]" pageId="18" pageNumber="19">
Primary
<typeStatus id="54D48836F070FFE5FEE7F901FE83F8D5" box="[306,357,1772,1797]" pageId="18" pageNumber="19">type</typeStatus>
data
</emphasis>
(
<figureCitation id="13542A11F070FFE5FE7AF901FDF7F8D5" box="[431,529,1772,1797]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="18" pageNumber="19">Fig. 135</figureCitation>
).
<typeStatus id="54D48836F070FFE5FDF1F901FD6AF8D5" box="[548,652,1772,1797]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
female (MACN): [Ene.-959/
<collectingCountry id="F3787604F070FFE5FC02F901FBAFF8D4" box="[983,1097,1772,1796]" name="Bolivia" pageId="18" pageNumber="19">BOLIVIA</collectingCountry>
/ Dº Sta-Cruz/ Pcia Cordillera/ PARAPETI/ coll. Martínez] handwritten; [
<taxonomicName id="4C6F4D17F070FFE5FDA0F8E2FCC2F8F8" box="[629,804,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/ n.sp./ A. MARTINEZ-DET 1959] partly handwritten]; [
<typeStatus id="54D48836F070FFE5FF48F8D9FEACF89C" box="[157,330,1844,1868]" pageId="18" pageNumber="19" type="holotype">HOLOTYPUS</typeStatus>
] red card; [
<taxonomicName id="4C6F4D17F070FFE5FE00F8D9FD62F89D" box="[469,644,1844,1869]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F070FFE5FD4DF8D9FD14F89D" box="[664,754,1844,1869]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pauliani">pauliani</taxonomicName>
/ sp.n./ A.MARTINEZ DET 1960] partly handwritten, red card; [WORLD/ SCARAB./ DATABASE/ WSD00016734];
<taxonomicName id="4C6F4D17F070FFE5FC99F8BAFC1EF8A0" box="[844,1016,1879,1904]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F070FFE5FBDFF8BAFB82F8A0" box="[1034,1124,1879,1904]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pauliani">pauliani</taxonomicName>
♀/
<bibRefCitation id="EFFE4B65F070FFE5FB41F8BAFAA7F8A0" author="Martinez" box="[1172,1345,1879,1904]" pageId="18" pageNumber="19" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">Martinez, 1960</bibRefCitation>
/ vid. F. Génier, 2014] partly handwritten.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF070FFE4FF12F84DFDC9FDED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BD03694F070FFE5FF12F84DFD61F869" blockId="18.[151,1437,151,2013]" box="[199,647,1952,1977]" pageId="18" pageNumber="19">
<emphasis id="B91BEA86F070FFE5FF12F84DFE4FF869" bold="true" box="[199,425,1952,1977]" pageId="18" pageNumber="19">Material examined</emphasis>
. Primary
<typeStatus id="54D48836F070FFE5FDCDF84CFDAEF869" box="[536,584,1953,1977]" pageId="18" pageNumber="19">type</typeStatus>
only.
</paragraph>
<paragraph id="8BD03694F070FFE4FF12F829FD6AFF28" blockId="18.[151,1437,151,2013]" lastBlockId="19.[151,1436,151,573]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B91BEA86F070FFE5FF12F829FE66F80D" bold="true" box="[199,384,1988,2013]" pageId="18" pageNumber="19">Natural history</emphasis>
. The only known specimen of this species was collected at night in “typical chaco boreal” forest as it was attracted to artificial light.
<bibRefCitation id="EFFE4B65F071FFE4FDA3FF7AFCC8FF60" author="Martinez" box="[630,814,151,176]" pageId="19" pageNumber="20" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">Martinez (1960)</bibRefCitation>
describes the flight pattern as being similar to those of
<taxonomicName id="4C6F4D17F071FFE4FF42FF51FE02FF05" box="[151,484,188,213]" class="Insecta" family="Geotrupidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Bolboceratinae">Bolboceratinae (Geotrupidae)</taxonomicName>
of the genera
<taxonomicName id="4C6F4D17F071FFE4FD51FF51FCBFFF05" authority="MacLeay" authorityName="MacLeay" box="[644,857,188,213]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F071FFE4FD51FF51FD01FF05" box="[644,743,188,213]" italics="true" pageId="19" pageNumber="20">Athyreus</emphasis>
MacLeay
</taxonomicName>
,
<taxonomicName id="4C6F4D17F071FFE4FCB1FF51FB67FF04" authority="Boucomont" authorityName="Boucomont" box="[868,1153,188,213]" class="Insecta" family="Bolboceratidae" genus="Bolborhinum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F071FFE4FCB1FF51FC11FF05" box="[868,1015,188,213]" italics="true" pageId="19" pageNumber="20">Bolborhinum</emphasis>
Boucomont
</taxonomicName>
and
<taxonomicName id="4C6F4D17F071FFE4FB62FF51FA70FF04" authority="Martinez." authorityName="Martinez." box="[1207,1430,188,212]" class="Insecta" family="Bolboceratidae" genus="Zefevazia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F071FFE4FB62FF51FAC5FF04" box="[1207,1315,188,212]" italics="true" pageId="19" pageNumber="20">Zefevazia</emphasis>
Martínez.
</taxonomicName>
<emphasis id="B91BEA86F071FFE4FF12FF32FED2FF28" bold="true" box="[199,308,223,248]" pageId="19" pageNumber="20">Remarks</emphasis>
.
<collectingCountry id="F3787604F071FFE4FE97FF32FE9AFF28" box="[322,380,223,248]" name="Malta" pageId="19" pageNumber="20">Male</collectingCountry>
and variation unknown.
</paragraph>
<paragraph id="8BD03694F071FFE4FF12FEE9FDC9FDED" blockId="19.[151,1436,151,573]" pageId="19" pageNumber="20">
At first we considered the only specimen known of the species as an odd small female of
<taxonomicName id="4C6F4D17F071FFE4FB60FEE8FAC7FECC" box="[1205,1313,261,284]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F071FFE4FB60FEE8FAC7FECC" box="[1205,1313,261,284]" italics="true" pageId="19" pageNumber="20">D. renatii</emphasis>
</taxonomicName>
. However, after closer examination, we prefer to maintain
<taxonomicName id="4C6F4D17F071FFE4FD15FEC4FCA4FE90" box="[704,834,295,320]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B91BEA86F071FFE4FD15FEC4FCA4FE90" box="[704,834,295,320]" italics="true" pageId="19" pageNumber="20">D. pauliani</emphasis>
</taxonomicName>
as valid. The peculiar configuration of the pronotal anterior carina (see description) was believed to be the minor configuration, but similar sized female of
<taxonomicName id="4C6F4D17F071FFE4FAFAFEA0FA7AFEB4" box="[1327,1436,333,356]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F071FFE4FAFAFEA0FA7AFEB4" box="[1327,1436,333,356]" italics="true" pageId="19" pageNumber="20">D. renatii</emphasis>
</taxonomicName>
are similar to well-developed specimens and have the carina set away from the anterior margin. In addition to this character, the flat anterior femur (slightly but distinctly convex in
<taxonomicName id="4C6F4D17F071FFE4FC4CFE78FBEEFE7C" box="[921,1032,405,428]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F071FFE4FC4CFE78FBEEFE7C" box="[921,1032,405,428]" italics="true" pageId="19" pageNumber="20">D. renatii</emphasis>
</taxonomicName>
) and absence of pronotal posterior fossae seems to further exclude it from
<taxonomicName id="4C6F4D17F071FFE4FDB2FE54FD30FE00" box="[615,726,441,464]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="renatii">
<emphasis id="B91BEA86F071FFE4FDB2FE54FD30FE00" box="[615,726,441,464]" italics="true" pageId="19" pageNumber="20">D. renatii</emphasis>
</taxonomicName>
. In addition, the body seems to be slightly more compressed dorsoventrally; the last tarsal segment appears simply spiniform and lack the internal spiniform projection beyond apical setae in the only known specimen, this however might be due to abrasion. Finally, the species is seemingly restricted to the Bolivian dry Chaco.
</paragraph>
</subSubSection>
</treatment>
</document>
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