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<document id="4E9926EE6BFF272B8EF3AD6282FE7FCA" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F05CFFC8FF42FE64FD7BFA4D" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Paradendropaemon) ganglbaueri Felsche 1909" docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="63" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="478E71C3732CA3ABD203BD534E34ECE2">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="029C54088BECB797A6F8611BF6F037C4">François Génier</mods:namePart>
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<mods:namePart id="A5B3FD4C88F346BA08447C1355512CF8">Patrick Arnaud</mods:namePart>
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<mods:title id="DD2BF6A670120C73A20391A16B8DD5F5">Zootaxa</mods:title>
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<mods:date id="6E7593ABA31E3065B3754535572767F8">2016</mods:date>
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<mods:number id="9A22C43899E3855B85B44AA513A6F19D">4099</mods:number>
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<treatment id="03C68782F05CFFC8FF42FE64FD7BFA4D" ID-DOI="http://doi.org/10.5281/zenodo.5780342" ID-GBIF-Taxon="127631972" ID-Zenodo-Dep="5780342" LSID="urn:lsid:plazi:treatment:03C68782F05CFFC8FF42FE64FD7BFA4D" httpUri="http://treatment.plazi.org/id/03C68782F05CFFC8FF42FE64FD7BFA4D" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<subSubSection id="C375651FF05CFFC9FF42FE64FC36FE73" box="[151,976,393,419]" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8BD03694F05CFFC9FF42FE64FC36FE73" blockId="62.[151,976,393,453]" box="[151,976,393,419]" pageId="62" pageNumber="63">
<heading id="D09881F8F05CFFC9FF42FE64FC36FE73" bold="true" box="[151,976,393,419]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B91BEA86F05CFFC9FF42FE64FC36FE73" bold="true" box="[151,976,393,419]" pageId="62" pageNumber="63">
31.
<taxonomicName id="4C6F4D17F05CFFC9FF16FE64FC36FE73" ID-CoL="34R2W" authority="Felsche, 1909" authorityName="Felsche" authorityYear="1909" box="[195,976,393,419]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri" subGenus="Paradendropaemon">
<emphasis id="B91BEA86F05CFFC9FF16FE64FCC4FE73" bold="true" box="[195,802,393,419]" italics="true" pageId="62" pageNumber="63">Dendropaemon (Paradendropaemon) ganglbaueri</emphasis>
<bibRefCitation id="EFFE4B65F05CFFC9FCFCFE64FC36FE73" author="Felsche" box="[809,976,393,419]" pageId="62" pageNumber="63" refString="Felsche, C. (1909) Neue und alte coprophage Scarabaeiden. (Col.). Deutsche Entomologische Zeitschrift, 1909 (6), 751 - 765. http: // dx. doi. org / 10.1002 / mmnd. 48019090608" type="journal article" year="1909">Felsche, 1909</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05CFFC9FF42FE41FE4DFE15" box="[151,427,428,453]" pageId="62" pageNumber="63" type="description">
<paragraph id="8BD03694F05CFFC9FF42FE41FE4DFE15" blockId="62.[151,976,393,453]" box="[151,427,428,453]" pageId="62" pageNumber="63">
(
<figureCitation id="13542A11F05CFFC9FF4AFE41FF1AFE14" box="[159,252,428,453]" captionStart="FIGURES 31 – 36" captionStartId="104.[151,264,1957,1980]" captionTargetBox="[266,1316,210,1927]" captionTargetId="figure@104.[251,1336,193,1937]" captionTargetPageId="104" captionText="FIGURES 31 – 36. Dorsal habitus. 31. D. ganglbaueri, male holotype; 32. D. vazdemelloi, male holotype; 33. D. refulgens, male holotype; 34. D. fractipes, male holotype; 35. D. fasces, male lectotype; 36. D. haroldi, male lectotype." httpUri="https://zenodo.org/record/268280/files/figure.png" pageId="62" pageNumber="63">Figs. 31</figureCitation>
,
<figureCitation id="13542A11F05CFFC9FEDDFE41FED2FE15" box="[264,308,428,453]" captionStart="FIGURES 126 – 133" captionStartId="114.[151,264,1932,1955]" captionTargetBox="[221,1363,201,1897]" captionTargetId="figure@114.[211,1376,193,1911]" captionTargetPageId="114" captionText="FIGURES 126 – 133. Metatarsus, ventral view. 126. D. ganglbaueri, holotype; 127. D. vazdemelloi, holotype; 128. D. refulgens, holotype; 129. D. fractipes; 130. D. fasces; 131. D. nitidicollis; 132. D. quadratus; 133. D. denticollis." httpUri="https://zenodo.org/record/268284/files/figure.png" pageId="62" pageNumber="63">126</figureCitation>
,
<figureCitation id="13542A11F05CFFC9FE95FE41FE8AFE14" box="[320,364,428,452]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="62" pageNumber="63">148</figureCitation>
,
<figureCitation id="13542A11F05CFFC9FEADFE41FE45FE15" box="[376,419,428,453]" captionStart="FIGURE 159" captionStartId="119.[151,250,1249,1271]" captionTargetBox="[150,1436,193,1227]" captionTargetId="figure@119.[150,1437,193,1228]" captionTargetPageId="119" captionText="FIGURE 159. Known distribution of D. amyntas, D. attalus, D. lydiae, D. morettoi, D. ganglbaueri, D. vazdemelloi." httpUri="https://zenodo.org/record/268289/files/figure.png" pageId="62" pageNumber="63">159</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05CFFC9FF42FE19FAB7FCAA" pageId="62" pageNumber="63" type="reference_group">
<paragraph id="8BD03694F05CFFC9FF42FE19FBCAFDDA" blockId="62.[151,1361,500,890]" box="[151,1068,500,522]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FE19FDDDFDD9" ID-CoL="8HFDR" authority="Ganglbaueri Felsche 1909" authorityName="Ganglbaueri Felsche" authorityYear="1909" box="[151,571,500,522]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05CFFC9FF42FE19FE48FDDA" box="[151,430,500,522]" italics="true" pageId="62" pageNumber="63">Dendropemon Ganglbaueri</emphasis>
<bibRefCitation id="EFFE4B65F05CFFC9FE60FE19FDDDFDD9" author="Felsche" box="[437,571,500,522]" pageId="62" pageNumber="63" refString="Felsche, C. (1909) Neue und alte coprophage Scarabaeiden. (Col.). Deutsche Entomologische Zeitschrift, 1909 (6), 751 - 765. http: // dx. doi. org / 10.1002 / mmnd. 48019090608" type="journal article" year="1909">
Felsche
<emphasis id="B91BEA86F05CFFC9FDDDFE19FDDDFDD9" bold="true" box="[520,571,500,521]" pageId="62" pageNumber="63">1909</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD9DFE18FD02FDDA" box="[584,740,501,522]" italics="true" pageId="62" pageNumber="63">Deut. Ent. Zeit.</emphasis>
1909: 755 (original description)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FDFFFCB2FDF8" blockId="62.[151,1361,500,890]" box="[151,852,530,552]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FDFFFE48FDF8" ID-CoL="8HFDR" authority="Ganglbaueri" authorityName="Ganglbaueri" box="[151,430,530,552]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05CFFC9FF42FDFFFE48FDF8" box="[151,430,530,552]" italics="true" pageId="62" pageNumber="63">Dendropemon Ganglbaueri</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE69FDFFFDCBFDF7" author="Gillet" box="[444,557,530,552]" pageId="62" pageNumber="63" refString="Gillet, J. J. E. (1911) Coleopterorum Catalogus, Scarabaeidae: Coprinae I. Vol. 38. W Junk, Berlin, 263 pp." type="book" year="1911">
Gillet
<emphasis id="B91BEA86F05CFFC9FE29FDFFFDCBFDF7" bold="true" box="[508,557,530,551]" pageId="62" pageNumber="63">1911</emphasis>
</bibRefCitation>
,
<emphasis id="B91BEA86F05CFFC9FDEFFDFFFD71FDF8" box="[570,663,530,552]" italics="true" pageId="62" pageNumber="63">Col. Cat.</emphasis>
38: 88 (catalogue)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FDDCFC71FD97" blockId="62.[151,1361,500,890]" box="[151,919,561,583]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FDDCFD82FD96" ID-CoL="34R2W" authority="Olsoufieff 1924" authorityName="Olsoufieff" authorityYear="1924" box="[151,612,561,583]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FDDCFE50FD97" box="[151,438,561,583]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE16FDDCFD82FD96" author="Olsoufieff" box="[451,612,561,583]" pageId="62" pageNumber="63" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">
Olsoufieff
<emphasis id="B91BEA86F05CFFC9FDE4FDDCFD82FD96" bold="true" box="[561,612,561,582]" pageId="62" pageNumber="63">1924</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FDA5FDDFFD5FFD97" box="[624,697,562,583]" italics="true" pageId="62" pageNumber="63">Insecta</emphasis>
13: 121 (monograph)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FDBDFBFEFD73" blockId="62.[151,1361,500,890]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FDBDFD75FDB5" ID-CoL="34R2W" authority="Pessoa &amp; Lane 1936" authorityName="Pessoa &amp; Lane" authorityYear="1936" box="[151,659,592,614]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FDBDFE50FDB6" box="[151,438,592,614]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE16FDBDFD75FDB5" author="Pessoa" box="[451,659,592,614]" pageId="62" pageNumber="63" refString="Pessoa, S. B. &amp; Lane, F. (1936) Sobre os coleopteros do genero Dendropaemon Perty, de S. Paulo e regioes visinhas. Revista de Biologia e Hygiene, 7 (2), 89 - 93." type="journal article" year="1936">
Pessôa &amp; Lane
<emphasis id="B91BEA86F05CFFC9FDB4FDBDFD75FDB5" bold="true" box="[609,659,592,613]" pageId="62" pageNumber="63">1936</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD75FDBCFCBDFDB6" box="[672,859,592,614]" italics="true" pageId="62" pageNumber="63">Rev. Biol. Hygiene</emphasis>
7: 89 (identification key, comment)
<taxonomicName id="4C6F4D17F05CFFC9FF42FD83FD75FD54" ID-CoL="34R2W" authority="Pessoa &amp; Lane 1941" authorityName="Pessoa &amp; Lane" authorityYear="1941" box="[151,659,622,644]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FD83FE50FD54" box="[151,438,622,644]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE16FD82FD75FD54" author="Pessoa" box="[451,659,622,644]" pageId="62" pageNumber="63" refString="Pessoa, S. B. &amp; Lane, F. (1941) Coleopteros necrophagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de Sao Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo, 2, 389 - 504." type="journal article" year="1941">
Pessôa &amp; Lane
<emphasis id="B91BEA86F05CFFC9FDB4FD82FD75FD54" bold="true" box="[609,659,623,644]" pageId="62" pageNumber="63">1941</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD75FD82FC85FD54" box="[672,867,622,644]" italics="true" pageId="62" pageNumber="63">Arq. Zool. S. Paulo</emphasis>
2: 490 (identification key, comment)
<taxonomicName id="4C6F4D17F05CFFC9FF42FD60FD88FD72" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,622,653,675]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FD60FE4FFD73" box="[151,425,653,675]" italics="true" pageId="62" pageNumber="63">Dendropemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE63FD60FD88FD72" author="Blackwelder" box="[438,622,653,675]" pageId="62" pageNumber="63" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F05CFFC9FDE9FD60FD88FD72" bold="true" box="[572,622,653,674]" pageId="62" pageNumber="63">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FDA9FD63FCAEFD73" box="[636,840,653,675]" italics="true" pageId="62" pageNumber="63">
<collectingCountry id="F3787604F05CFFC9FDA9FD63FD4EFD73" box="[636,680,654,675]" name="United States of America" pageId="62" pageNumber="63">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F05CFFC9FD78FD63FD33FD73" box="[685,725,654,675]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull.
</emphasis>
185: 210 (checklist)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FD41FC18FD12" blockId="62.[151,1361,500,890]" box="[151,1022,684,706]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FD41FDADFD11" authority="Martinez 1944" authorityName="Martinez" authorityYear="1944" box="[151,587,684,706]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FD41FE4FFD12" box="[151,425,684,706]" italics="true" pageId="62" pageNumber="63">Dendropemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE63FD41FDADFD11" author="Martinez" box="[438,587,684,706]" pageId="62" pageNumber="63" refString="Martinez, A. (1944) Insectos nuevos o poco conocidos II. Revista Argentina de Entomologia, 2, 34 - 40." type="journal article" year="1944">
Martínez
<emphasis id="B91BEA86F05CFFC9FDCDFD41FDADFD11" bold="true" box="[536,587,684,705]" pageId="62" pageNumber="63">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD82FD40FD02FD12" box="[599,740,684,706]" italics="true" pageId="62" pageNumber="63">Rev. Arg. Ent.</emphasis>
2: 35 (comment taxonomy)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FD27FC10FD2F" blockId="62.[151,1361,500,890]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FD27FCC9FD0F" authority="Edmonds 1972" authorityName="Edmonds" authorityYear="1972" box="[151,815,714,736]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri" subGenus="Paradendropaemon">
<emphasis id="B91BEA86F05CFFC9FF42FD27FD6CFD30" box="[151,650,714,736]" italics="true" pageId="62" pageNumber="63">Dendropemon (Paradendropaemon) ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FD42FD27FCC9FD0F" author="Edmonds" box="[663,815,714,736]" pageId="62" pageNumber="63" refString="Edmonds, W. D. (1972) Comparative Skeletal Morphology, Systematics and Evolution of the Phanaeine Dung Beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin, 49 (11), 731 - 874." type="journal article" year="1972">
Edmonds
<emphasis id="B91BEA86F05CFFC9FD28FD27FCC9FD0F" bold="true" box="[765,815,714,735]" pageId="62" pageNumber="63">1972</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FCE9FD26FBF1FD30" box="[828,1047,714,736]" italics="true" pageId="62" pageNumber="63">Univ. Kansas Sc. Bull</emphasis>
. 49: 849 (misidentification)
<taxonomicName id="4C6F4D17F05CFFC9FF42FD04FE50FD2F" ID-CoL="34R2W" box="[151,438,745,767]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FD04FE50FD2F" box="[151,438,745,767]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
</taxonomicName>
: Vaz-de-Mello
<emphasis id="B91BEA86F05CFFC9FD81FD04FD61FD2E" bold="true" box="[596,647,745,766]" pageId="62" pageNumber="63">2000</emphasis>
,
<emphasis id="B91BEA86F05CFFC9FD41FD07FCB3FD2F" box="[660,853,746,767]" italics="true" pageId="62" pageNumber="63">Hac. Proy. CYTED</emphasis>
: 192 (faunistic)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FCE5FC1AFCEC" blockId="62.[151,1361,500,890]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FCE5FCCEFCCD" ID-CoL="34R2W" authority="Arnaud 2002" authorityName="Arnaud" authorityYear="2002" box="[151,808,776,798]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri" subGenus="Paradendropaemon">
<emphasis id="B91BEA86F05CFFC9FF42FCE5FD70FCCE" box="[151,662,776,798]" italics="true" pageId="62" pageNumber="63">Dendropaemon (Paradendropaemon) ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FD71FCE5FCCEFCCD" author="Arnaud" box="[676,808,776,798]" pageId="62" pageNumber="63" refString="Arnaud, P. (2002) Phanaeini, Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Les Coleopteres du Monde, 28. Hillside Books, Canterbury, 151 pp." type="book" year="2002">
Arnaud
<emphasis id="B91BEA86F05CFFC9FD23FCE5FCCEFCCD" bold="true" box="[758,808,776,797]" pageId="62" pageNumber="63">2002</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FCE0FCE5FC4AFCCE" box="[821,940,776,798]" italics="true" pageId="62" pageNumber="63">Col. Monde</emphasis>
28: 14 (misidentification)
<taxonomicName id="4C6F4D17F05CFFC9FF42FCCBFD9FFCEC" ID-CoL="34R2W" authority="Philips et al. 2004" authorityName="Philips et al." authorityYear="2004" box="[151,633,806,828]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FCCBFE50FCEC" box="[151,438,806,828]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE16FCCBFD9FFCEC" author="Philips" box="[451,633,806,828]" pageId="62" pageNumber="63" refString="Philips, T. K., Edmonds, W. D. &amp; Scholtz, C. H. (2004) A phylogenetic analysis of the New World tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae): Hypotheses on relationships and origins. Insect Systematics &amp; Evolution, 35, 43 - 63. http: // dx. doi. org / 10.1163 / 187631204788964664" type="journal article" year="2004">
Philips et al.
<emphasis id="B91BEA86F05CFFC9FD92FCCAFD9FFCEC" bold="true" box="[583,633,807,828]" pageId="62" pageNumber="63">2004</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD53FCC5FCD2FCEC" box="[646,820,806,829]" italics="true" pageId="62" pageNumber="63">Insect Syst. Evol.</emphasis>
35: 51 (phylogeny)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FCA8FC81FC8B" blockId="62.[151,1361,500,890]" box="[151,871,837,859]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FCA8FDA8FC8A" ID-CoL="34R2W" authority="Barbosa 2008" authorityName="Barbosa" authorityYear="2008" box="[151,590,837,859]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FF42FCA8FE50FC8B" box="[151,438,837,859]" italics="true" pageId="62" pageNumber="63">Dendropaemon ganglbaueri</emphasis>
:
<bibRefCitation id="EFFE4B65F05CFFC9FE16FCA8FDA8FC8A" author="Barbosa" box="[451,590,837,859]" pageId="62" pageNumber="63" refString="Barbosa Silva, F. A. (2008) Estudo da Comunidade de Scarabaeinae (Coleoptera: Scarabaeidae) em dois Ambientes do Refugio Ecologico Charles Darwin, Igarassu-PE. M. Sc. Thesis, Universidade Federal de Pernambuco-UFPE, Recife, 79 pp." type="book" year="2008">
Barbosa
<emphasis id="B91BEA86F05CFFC9FDC9FCA8FDA8FC8A" bold="true" box="[540,590,837,858]" pageId="62" pageNumber="63">2008</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05CFFC9FD8EFCABFD3BFC8B" box="[603,733,838,859]" italics="true" pageId="62" pageNumber="63">Comm. Scar.</emphasis>
: 19 (biology)
</paragraph>
<paragraph id="8BD03694F05CFFC9FF42FC89FAB7FCAA" blockId="62.[151,1361,500,890]" box="[151,1361,868,890]" pageId="62" pageNumber="63">
<taxonomicName id="4C6F4D17F05CFFC9FF42FC89FD70FCAA" ID-CoL="34R2W" box="[151,662,868,890]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri" subGenus="Paradendropaemon">
<emphasis id="B91BEA86F05CFFC9FF42FC89FD70FCAA" box="[151,662,868,890]" italics="true" pageId="62" pageNumber="63">Dendropaemon (Paradendropaemon) ganglbaueri</emphasis>
</taxonomicName>
: Vaz-de-Mello &amp; Génier
<emphasis id="B91BEA86F05CFFC9FC4FFC89FC2AFCA9" bold="true" box="[922,972,868,889]" pageId="62" pageNumber="63">2009</emphasis>
,
<emphasis id="B91BEA86F05CFFC9FC0CFC89FBDAFCAA" box="[985,1084,868,890]" italics="true" pageId="62" pageNumber="63">Col. Bull.</emphasis>
63: 364 (misidentification)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05CFFC9FF42FC49FDBBFC6D" box="[151,605,932,957]" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8BD03694F05CFFC9FF42FC49FDBBFC6D" blockId="62.[151,1437,932,2037]" box="[151,605,932,957]" pageId="62" pageNumber="63">
<emphasis id="B91BEA86F05CFFC9FF42FC49FECBFC6D" bold="true" box="[151,301,932,957]" pageId="62" pageNumber="63">
<typeStatus id="54D48836F05CFFC9FF42FC49FF34FC6D" box="[151,210,932,957]" pageId="62" pageNumber="63">Type</typeStatus>
locality
</emphasis>
. Sao Paulo, Süd-Brasilien.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05CFFC9FF12FC2AFE0CFBA0" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8BD03694F05CFFC9FF12FC2AFE0CFBA0" blockId="62.[151,1437,932,2037]" pageId="62" pageNumber="63">
<emphasis id="B91BEA86F05CFFC9FF12FC2AFEA7FC0F" bold="true" box="[199,321,967,992]" pageId="62" pageNumber="63">Diagnosis.</emphasis>
The long, approximately 3.5 times as long as wide at apex, first metatarsomere will place
<taxonomicName id="4C6F4D17F05CFFC9FA54FC24FEC6FBD5" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91BEA86F05CFFC9FA54FC24FEC6FBD5" italics="true" pageId="62" pageNumber="63">D. ganglbaueri</emphasis>
</taxonomicName>
in
<taxonomicName id="4C6F4D17F05CFFC9FE9BFC01FDCBFBD5" box="[334,557,1004,1029]" class="Insecta" family="Scarabaeidae" genus="Paradendropaemon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05CFFC9FE9BFC01FDCBFBD5" box="[334,557,1004,1029]" italics="true" pageId="62" pageNumber="63">Paradendropaemon</emphasis>
</taxonomicName>
. Differs from the only other species in the group,
<taxonomicName id="4C6F4D17F05CFFC9FB44FC00FAD8FBD5" box="[1169,1342,1004,1029]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B91BEA86F05CFFC9FB44FC00FAD8FBD5" box="[1169,1342,1004,1029]" italics="true" pageId="62" pageNumber="63">D. vazdemelloi</emphasis>
</taxonomicName>
, by the longitudinally compressed first metatarsomere, the metatibial anterior surface which is glossy between the fine irregular punctures, the anterior pronotal margin which is distinctly enlarged and flat lateral to the eye and by its seemingly larger average size.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05CFFC8FF12FB91FB94FC18" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
<paragraph id="8BD03694F05CFFC8FF12FB91FD19FD20" blockId="62.[151,1437,932,2037]" lastBlockId="63.[151,1437,151,1437]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<emphasis id="B91BEA86F05CFFC9FF12FB91FEB7FB45" bold="true" box="[199,337,1148,1173]" pageId="62" pageNumber="63">Description</emphasis>
.
<collectingCountry id="F3787604F05CFFC9FEB7FB91FE7DFB45" box="[354,411,1148,1173]" name="Malta" pageId="62" pageNumber="63">Male</collectingCountry>
<typeStatus id="54D48836F05CFFC9FE70FB91FDE1FB45" box="[421,519,1148,1173]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F05CFFC9FDCCFB91FD89FB44" box="[537,623,1148,1173]" captionStart="FIGURES 31 – 36" captionStartId="104.[151,264,1957,1980]" captionTargetBox="[266,1316,210,1927]" captionTargetId="figure@104.[251,1336,193,1937]" captionTargetPageId="104" captionText="FIGURES 31 – 36. Dorsal habitus. 31. D. ganglbaueri, male holotype; 32. D. vazdemelloi, male holotype; 33. D. refulgens, male holotype; 34. D. fractipes, male holotype; 35. D. fasces, male lectotype; 36. D. haroldi, male lectotype." httpUri="https://zenodo.org/record/268280/files/figure.png" pageId="62" pageNumber="63">Fig. 31</figureCitation>
).
<emphasis id="B91BEA86F05CFFC9FD53FB91FD24FB45" bold="true" box="[646,706,1148,1173]" pageId="62" pageNumber="63">Body</emphasis>
. Body large, length 20.0 mm, maximum width 12.0 mm; body elongate-oval in dorsal view; dorsum convex.
<emphasis id="B91BEA86F05CFFC9FD71FB72FD03FB68" bold="true" box="[676,741,1183,1208]" pageId="62" pageNumber="63">Color</emphasis>
. Dorsal surface dark brown to black, glossy, lacking metallic sheen; ventrum dark brown to black; pygidium dark brown; legs dark reddish brown to black.
<emphasis id="B91BEA86F05CFFC9FB3BFB29FACBFB0D" bold="true" box="[1262,1325,1220,1245]" pageId="62" pageNumber="63">Head</emphasis>
. Clypeus broadly arcuate, anterior portion slightly upturned; clypeal teeth obtusely triangular; clypeal median emargination v-shaped, clypeal edge emarginate on external side of each clypeal tooth, clypeal teeth ventral surface with an arcuate irregular carina, clypeal margin well-defined and sharply carinate posteriorly, clypeal surface with transverse rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with fine blunt transverse rugulae, lacking distinct transverse carina, concave laterally and convex internally; clypeofrontal carina rather low, approximately 4 times wider than high, straight in dorsal view, tuberculate, clypeofrontal carina apical edge trilobate, median lobe much higher than laterals in frontal view; eyes small in dorsal view, interocular ratio 5.9.
<emphasis id="B91BEA86F05CFFC9FF18FA09FEA0FA2D" bold="true" box="[205,326,1508,1533]" pageId="62" pageNumber="63">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.6; disc of pronotum finely punctate basally, punctures changing into fine rough rugulae anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin unmodified lateral to eye; anterior portion with a nearly straight and slightly tuberculate medially transverse carina bordering anteriorly a large concavity; anterior angles surface with fine rugulae, similar to lateral margin along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions slightly explanate; pronotal basal fossae ill-defined, slightly concave; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F05CFFC9FD23F951FCA4F905" bold="true" box="[758,834,1724,1749]" pageId="62" pageNumber="63">Elytra</emphasis>
. Elytra slightly transverse, elytral combined width/ length ratio 1.3; elytral base lacking distinct margin, simply convex; elytral striae 1–4 moderately wide, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures minute, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, finely punctate throughout, surface glossy.
<emphasis id="B91BEA86F05CFFC9FDA8F8A1FCDDF8B5" bold="true" box="[637,827,1868,1893]" pageId="62" pageNumber="63">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, keel-shaped.
<emphasis id="B91BEA86F05CFFC9FBCDF882FBB6F858" bold="true" box="[1048,1104,1903,1928]" pageId="62" pageNumber="63">Legs</emphasis>
. Profemur posterior surface convex, glabrous and punctate internally, posterointernal margin rather thin, evenly developed, internal edge wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface finely punctate along setae. Protibia with four teeth on lateral edge; internal basal angle unmodified; anterior surface lacking aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface lacking punctures externally to median carina, surface finely and irregularly microsculptured, with several unaligned and contiguous setal row along lateral teeth. Mesofemur unmodified on anterointernal edge apically. Mesotibia abruptly widening before apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge angular, lacking distinct emargination internally; external edge more or less flat, bordered anteriorly and posteriorly by an almost complete setal row. Mesotarsus differently shaped than metatarsus, 2-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia abruptly widening before apex in anterior view, anterior surface with distinct row of setae, with fine irregular punctures on a glossy surface, metatibial posterior surface concave between internal and lateral edge, glossy between punctures. Metatarsus 2-segmented (
<figureCitation id="13542A11F05DFFC8FD77FDAAFCE1FDB0" box="[674,775,583,608]" captionStart="FIGURES 126 – 133" captionStartId="114.[151,264,1932,1955]" captionTargetBox="[221,1363,201,1897]" captionTargetId="figure@114.[211,1376,193,1911]" captionTargetPageId="114" captionText="FIGURES 126 – 133. Metatarsus, ventral view. 126. D. ganglbaueri, holotype; 127. D. vazdemelloi, holotype; 128. D. refulgens, holotype; 129. D. fractipes; 130. D. fasces; 131. D. nitidicollis; 132. D. quadratus; 133. D. denticollis." httpUri="https://zenodo.org/record/268284/files/figure.png" pageId="63" pageNumber="64">Fig. 126</figureCitation>
), first segment elongate, more than three time as long as wide at apex, with anterointernal carina indistinguishable.
<emphasis id="B91BEA86F05DFFC8FCF8FD81FBC6FD55" bold="true" box="[813,1056,620,645]" pageId="63" pageNumber="64">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with more than three unaligned row of setae laterally, narrowly glabrous medially; sternite 7 approximately longitudinally flat medially, as long as segment 5–6 combined along midline; pygidium finely punctate on disc.
<emphasis id="B91BEA86F05DFFC8FE8EFD3AFDE3FD20" bold="true" box="[347,517,727,752]" pageId="63" pageNumber="64">
<collectingCountry id="F3787604F05DFFC8FE8EFD3AFE71FD20" box="[347,407,727,752]" name="Malta" pageId="63" pageNumber="64">Male</collectingCountry>
genitalia
</emphasis>
. Parameres unknown.
</paragraph>
<paragraph id="8BD03694F05DFFC8FF12FD11FD54FCC4" blockId="63.[151,1437,151,1437]" box="[199,690,764,789]" pageId="63" pageNumber="64">
<emphasis id="B91BEA86F05DFFC8FF12FD11FE90FCC5" bold="true" box="[199,374,764,789]" pageId="63" pageNumber="64">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF05DFFC8FE5DFD11FE35FCC5" box="[392,467,764,789]" pageId="63" pageNumber="64" type="male">1 male</specimenCount>
). Length: 20.0 mm.
</paragraph>
<paragraph id="8BD03694F05DFFC8FF12FCF2FB94FC18" blockId="63.[151,1437,151,1437]" pageId="63" pageNumber="64">
<emphasis id="B91BEA86F05DFFC8FF12FCF2FE55FCE8" bold="true" box="[199,435,799,824]" pageId="63" pageNumber="64">
Primary
<typeStatus id="54D48836F05DFFC8FEEEFCF2FE88FCE8" box="[315,366,799,824]" pageId="63" pageNumber="64">type</typeStatus>
data
</emphasis>
(
<figureCitation id="13542A11F05DFFC8FE1FFCF2FDD3FCE8" box="[458,565,799,824]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="63" pageNumber="64">Fig. 148</figureCitation>
).
<collectingCountry id="F3787604F05DFFC8FD84FCF2FD6CFCE8" box="[593,650,799,824]" name="Malta" pageId="63" pageNumber="64">Male</collectingCountry>
<typeStatus id="54D48836F05DFFC8FD4CFCF2FD1DFCE8" box="[665,763,799,824]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
(MTD): [Sao Paulo]; [
<taxonomicName id="4C6F4D17F05DFFC8FBC9FCF2FB4EFCE8" box="[1052,1192,799,824]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="120" phylum="Arthropoda" rank="species" species="ganglbaueri">Ganglbaueri</taxonomicName>
/ Felsche/
<collectingCountry id="F3787604F05DFFC8FAF8FCF2FA73FCE8" box="[1325,1429,799,824]" name="Brazil" pageId="63" pageNumber="64">Brasilien</collectingCountry>
] handwritten; [♀] disc; [Coll. C. Felsche/ Kauf 20, 1918] green card; [
<typeStatus id="54D48836F05DFFC8FC1FFCA9FBCEFC8C" box="[970,1064,836,860]" pageId="63" pageNumber="64">TYPUS</typeStatus>
] red card; [Staatl. Museum für/ Tierkunde Dresden]; [
<taxonomicName id="4C6F4D17F05DFFC8FE4CFC8AFDAEFC50" box="[409,584,871,896]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F05DFFC8FD8BFC8AFD04FC50" box="[606,738,871,896]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ganglbaueri">ganglbaueri</taxonomicName>
♀/ Felsche/ Sao Paulo. Blut det] handwritten; [WORLD/ SCARAB./ DATABASE/ WSD00002076]; [
<typeStatus id="54D48836F05DFFC8FD42FC61FCCBFC74" box="[663,813,908,932]" pageId="63" pageNumber="64" type="holotype">HOLOTYPE</typeStatus>
/
<emphasis id="B91BEA86F05DFFC8FC97FC61FBE3FC75" box="[834,1029,908,933]" italics="true" pageId="63" pageNumber="64">Dendrop[a]emon</emphasis>
/
<emphasis id="B91BEA86F05DFFC8FBCCFC61FB48FC75" box="[1049,1198,908,933]" italics="true" pageId="63" pageNumber="64">ganglebaueri</emphasis>
/
<bibRefCitation id="EFFE4B65F05DFFC8FB17FC61FA80FC74" author="Felsche" box="[1218,1382,908,933]" pageId="63" pageNumber="64" refString="Felsche, C. (1909) Neue und alte coprophage Scarabaeiden. (Col.). Deutsche Entomologische Zeitschrift, 1909 (6), 751 - 765. http: // dx. doi. org / 10.1002 / mmnd. 48019090608" type="journal article" year="1909">Felsche, 1909</bibRefCitation>
] red card; [
<taxonomicName id="4C6F4D17F05DFFC8FF09FC42FE6DFC18" box="[220,395,943,968]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F05DFFC8FE4EFC42FDF8FC18" box="[411,542,943,968]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ganglbaueri">ganglbaueri</taxonomicName>
/ Felsche ♂/ vid. F. Génier, 2012] partly handwritten.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05DFFC8FF12FC39FDE4FBC0" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="8BD03694F05DFFC8FF12FC39FD61FC3D" blockId="63.[151,1437,151,1437]" box="[199,647,980,1005]" pageId="63" pageNumber="64">
<emphasis id="B91BEA86F05DFFC8FF12FC39FE4FFC3D" bold="true" box="[199,425,980,1005]" pageId="63" pageNumber="64">Material examined</emphasis>
. Primary
<typeStatus id="54D48836F05DFFC8FDCDFC38FDAEFC3D" box="[536,584,981,1005]" pageId="63" pageNumber="64">type</typeStatus>
only.
</paragraph>
<paragraph id="8BD03694F05DFFC8FF12FC1AFDE4FBC0" blockId="63.[151,1437,151,1437]" box="[199,514,1015,1040]" pageId="63" pageNumber="64">
<emphasis id="B91BEA86F05DFFC8FF12FC1AFE9BFBC0" bold="true" box="[199,381,1015,1040]" pageId="63" pageNumber="64">Natural history</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05DFFC8FF12FBF1FD7BFA4D" pageId="63" pageNumber="64" type="discussion">
<paragraph id="8BD03694F05DFFC8FF12FBF1FD43FBE5" blockId="63.[151,1437,151,1437]" box="[199,677,1052,1077]" pageId="63" pageNumber="64">
<emphasis id="B91BEA86F05DFFC8FF12FBF1FED2FBE5" bold="true" box="[199,308,1052,1077]" pageId="63" pageNumber="64">Remarks</emphasis>
. Female and variation unknown.
</paragraph>
<paragraph id="8BD03694F05DFFC8FF12FBADFEC5FB37" blockId="63.[151,1437,151,1437]" pageId="63" pageNumber="64">
<collectingCountry id="F3787604F05DFFC8FF12FBADFF00FB88" box="[199,230,1088,1112]" name="American Samoa" pageId="63" pageNumber="64">As</collectingCountry>
opposed to what Felsche and Blut stated, the
<typeStatus id="54D48836F05DFFC8FD30FBD2FCA1FB88" box="[741,839,1087,1112]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
is a male, but the aedeagus is missing. Removing the viscera and stuffing cotton in the cavity was often done when preparing large specimens in the 19th century. In addition to the characters mentioned above, the metasternal keel differs in being more developed and less arcuate in lateral view and the first metatarsomere has a single distinct fine longitudinal sulcus instead of two in
<taxonomicName id="4C6F4D17F05DFFC8FA54FB40FEFAFB38" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B91BEA86F05DFFC8FA54FB40FEFAFB38" italics="true" pageId="63" pageNumber="64">D. vazdemelloi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD03694F05DFFC8FF12FB19FD7BFA4D" blockId="63.[151,1437,151,1437]" pageId="63" pageNumber="64">
Unfortunately, the single known specimen only possess a generic (contemporary?) printed label stating “Sao Paulo”. Therefore, Felsche stated that the
<typeStatus id="54D48836F05DFFC8FDA3FAF5FD40FAE0" box="[630,678,1304,1328]" pageId="63" pageNumber="64">type</typeStatus>
locality is “Sao Paulo”, it is however not clear if it is taken in the sense of the state or the city. The original manuscript label only state “
<collectingCountry id="F3787604F05DFFC8FC37FAD1FBAAFA85" box="[994,1100,1340,1365]" name="Brazil" pageId="63" pageNumber="64">Brasilien</collectingCountry>
” suggesting that it might be anywhere in
<collectingCountry id="F3787604F05DFFC8FEFDFAB2FE96FAA8" box="[296,368,1375,1400]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
. No additional specimens have ever been collected from the city or the state of São Paulo since it was originally described over a century ago.
</paragraph>
</subSubSection>
</treatment>
</document>
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