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<document id="9F12A66691E06F13FE45F616A7BC2DE9" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F02CFFA7FF42F8A6FC0CFC75" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Sulcopaemon) similis Blut 1939" docType="treatment" docVersion="10" lastPageNumber="81" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="79" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="32A09EF6114579728F5F16C2E93F4043">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="9745991CE81DC53BC5CB4BB8D0D8AD01">François Génier</mods:namePart>
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<mods:namePart id="F595B7D59550F240153EB424C2148A9E">Patrick Arnaud</mods:namePart>
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<mods:title id="A6D3FBBB8993FDFD39365E4B0D22D5C2">Zootaxa</mods:title>
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<mods:date id="4501826E761E9994CE2E8ACC3E7E53F2">2016</mods:date>
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<mods:number id="190EA3B3F7A205493FC34EA321E50191">4099</mods:number>
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<mods:number id="5AF4D45DC3AD2FECD367613BBA180D11">1</mods:number>
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<mods:classification id="90F7E12D6F7E9A7FE7303769569DFB9C">journal volume</mods:classification>
<mods:identifier id="2FB1AA857768793B19C30488FBC258AA" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
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<treatment id="03C68782F02CFFA7FF42F8A6FC0CFC75" ID-DOI="http://doi.org/10.5281/zenodo.5780372" ID-GBIF-Taxon="127631993" ID-Zenodo-Dep="5780372" LSID="urn:lsid:plazi:treatment:03C68782F02CFFA7FF42F8A6FC0CFC75" httpUri="http://treatment.plazi.org/id/03C68782F02CFFA7FF42F8A6FC0CFC75" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
<subSubSection id="C375651FF02CFFB9FF42F8A6FCFFF8B5" box="[151,793,1867,1893]" pageId="78" pageNumber="79" type="nomenclature">
<paragraph id="8BD03694F02CFFB9FF42F8A6FCFFF8B5" blockId="78.[151,793,1867,1926]" box="[151,793,1867,1893]" pageId="78" pageNumber="79">
<heading id="D09881F8F02CFFB9FF42F8A6FCFFF8B5" bold="true" box="[151,793,1867,1893]" fontSize="11" level="1" pageId="78" pageNumber="79" reason="1">
<emphasis id="B91BEA86F02CFFB9FF42F8A6FCFFF8B5" bold="true" box="[151,793,1867,1893]" pageId="78" pageNumber="79">
40.
<taxonomicName id="4C6F4D17F02CFFB9FF16F8A6FCFFF8B5" ID-CoL="34R37" authority="Blut, 1939" authorityName="Blut" authorityYear="1939" box="[195,793,1867,1893]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="similis" subGenus="Sulcopaemon">
<emphasis id="B91BEA86F02CFFB9FF16F8A6FD74F8B5" bold="true" box="[195,658,1867,1893]" italics="true" pageId="78" pageNumber="79">Dendropaemon (Sulcopaemon) similis</emphasis>
<bibRefCitation id="EFFE4B65F02CFFB9FD4FF8A6FCFFF8B5" author="Blut" box="[666,793,1867,1893]" pageId="78" pageNumber="79" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut, 1939</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF02CFFB9FF42F880FE4DF856" box="[151,427,1901,1926]" pageId="78" pageNumber="79" type="description">
<paragraph id="8BD03694F02CFFB9FF42F880FE4DF856" blockId="78.[151,793,1867,1926]" box="[151,427,1901,1926]" pageId="78" pageNumber="79">
(
<figureCitation id="13542A11F02CFFB9FF4AF880FF1AF856" box="[159,252,1901,1926]" captionStart="FIGURES 37 – 41" captionStartId="105.[151,264,1956,1979]" captionTargetBox="[258,1329,206,1918]" captionTargetId="figure@105.[242,1345,193,1935]" captionTargetPageId="105" captionText="FIGURES 37 – 41. Dorsal habitus. 37. D. latistriatus, male holotype; 38. D. nitidicollis, male lectotype; 39. D. quadratus, male neotype; 40. D. similis, male holotype; 41. D. denticollis, female holotype." httpUri="https://zenodo.org/record/268281/files/figure.png" pageId="78" pageNumber="79">Figs. 40</figureCitation>
, 106–107, 154)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF02CFFB8FF42F858FC92FF1C" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="reference_group">
<paragraph id="8BD03694F02CFFB8FF42F858FC92FF1C" blockId="78.[151,996,1973,2026]" lastBlockId="79.[151,1095,152,204]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">
<taxonomicName id="4C6F4D17F02CFFB9FF42F858FE00F81B" ID-CoL="34R37" authority="Blut 1939" authorityName="Blut" authorityYear="1939" box="[151,486,1973,1995]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91BEA86F02CFFB9FF42F858FE9AF81B" box="[151,380,1973,1995]" italics="true" pageId="78" pageNumber="79">Dendropaemon similis</emphasis>
<bibRefCitation id="EFFE4B65F02CFFB9FE57F858FE00F81B" author="Blut" box="[386,486,1973,1995]" pageId="78" pageNumber="79" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">
Blut
<emphasis id="B91BEA86F02CFFB9FE61F85BFE00F81B" bold="true" box="[436,486,1974,1995]" pageId="78" pageNumber="79">1939</emphasis>
</bibRefCitation>
</taxonomicName>
, Arch. Naturg. (N.F.) 8: 282 (original description)
<taxonomicName id="4C6F4D17F02CFFB9FF42F839FDC9F839" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,559,2004,2026]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="simile">
<emphasis id="B91BEA86F02CFFB9FF42F839FE8FF83A" box="[151,361,2004,2026]" italics="true" pageId="78" pageNumber="79">Dendropemon simile</emphasis>
:
<bibRefCitation id="EFFE4B65F02CFFB9FEA3F839FDC9F839" author="Blackwelder" box="[374,559,2004,2026]" pageId="78" pageNumber="79" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F02CFFB9FE28F839FDC9F839" bold="true" box="[509,559,2004,2025]" pageId="78" pageNumber="79">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="F3787604F02CFFB9FDE9F839FD8CF83A" box="[572,618,2004,2026]" name="United States of America" pageId="78" pageNumber="79">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F02CFFB9FDA5F838FD7EF83A" box="[624,664,2005,2026]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="81" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull. 185: 211 (checklist)
<taxonomicName id="4C6F4D17F02DFFB8FF42FF75FDC4FF7D" ID-CoL="34R37" authority="Edmonds 1972" authorityName="Edmonds" authorityYear="1972" box="[151,546,152,174]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91BEA86F02DFFB8FF42FF75FE9AFF7E" box="[151,380,152,174]" italics="true" pageId="79" pageNumber="80">Dendropaemon similis</emphasis>
:
<bibRefCitation id="EFFE4B65F02DFFB8FE5CFF75FDC4FF7D" author="Edmonds" box="[393,546,152,174]" pageId="79" pageNumber="80" refString="Edmonds, W. D. (1972) Comparative Skeletal Morphology, Systematics and Evolution of the Phanaeine Dung Beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin, 49 (11), 731 - 874." type="journal article" year="1972">
Edmonds
<emphasis id="B91BEA86F02DFFB8FE3AFF75FDC4FF7D" bold="true" box="[495,546,152,173]" pageId="79" pageNumber="80">1972</emphasis>
</bibRefCitation>
</taxonomicName>
, Univ. Kansas Sc. Bull. 49: 851 (comment taxonomy)
<taxonomicName id="4C6F4D17F02DFFB8FF42FF5BFDDBFF1C" ID-CoL="34R37" authority="Arnaud 2002" authorityName="Arnaud" authorityYear="2002" box="[151,573,182,204]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="similis" subGenus="Dendropaemon">
<emphasis id="B91BEA86F02DFFB8FF42FF5BFE4DFF1C" box="[151,427,182,204]" italics="true" pageId="79" pageNumber="80">Dendropaemon (D.) similis</emphasis>
:
<bibRefCitation id="EFFE4B65F02DFFB8FE6CFF5BFDDBFF1C" author="Arnaud" box="[441,573,182,204]" pageId="79" pageNumber="80" refString="Arnaud, P. (2002) Phanaeini, Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Les Coleopteres du Monde, 28. Hillside Books, Canterbury, 151 pp." type="book" year="2002">
Arnaud
<emphasis id="B91BEA86F02DFFB8FDDEFF5AFDDBFF1C" bold="true" box="[523,573,183,204]" pageId="79" pageNumber="80">2002</emphasis>
</bibRefCitation>
</taxonomicName>
, Col. Monde 28: 15 (mention)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF02DFFB8FF42FF14FE4CFEC2" box="[151,426,249,274]" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="8BD03694F02DFFB8FF42FF14FE4CFEC2" blockId="79.[151,1437,249,2038]" box="[151,426,249,274]" pageId="79" pageNumber="80">
<emphasis id="B91BEA86F02DFFB8FF42FF14FECBFEC2" bold="true" box="[151,301,249,274]" pageId="79" pageNumber="80">
<typeStatus id="54D48836F02DFFB8FF42FF14FF34FEC2" box="[151,210,249,274]" pageId="79" pageNumber="80">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F3787604F02DFFB8FEE9FF17FE40FEC2" box="[316,422,250,274]" name="Paraguay" pageId="79" pageNumber="80">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF02DFFB8FF12FEF0FCF8FE39" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8BD03694F02DFFB8FF12FEF0FCF8FE39" blockId="79.[151,1437,249,2038]" pageId="79" pageNumber="80">
<emphasis id="B91BEA86F02DFFB8FF12FEF0FEA7FEE5" bold="true" box="[199,321,285,310]" pageId="79" pageNumber="80">Diagnosis.</emphasis>
The small size combined with the three-segmented meso- and metatarsi with the second segment being subequal in length to the first segment will place the species in the
<taxonomicName id="4C6F4D17F02DFFB8FC0AFEACFBB7FE8A" box="[991,1105,321,346]" class="Insecta" family="Scarabaeidae" genus="Enicotarsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B91BEA86F02DFFB8FC0AFEACFBB7FE8A" box="[991,1105,321,346]" italics="true" pageId="79" pageNumber="80">quadratus</emphasis>
</taxonomicName>
species complex. The lateral pronotal fossae with the sharply carinate anterior edge will separate it from most other species in the group. The three-segmented meso- and metatarsi will separate it from
<taxonomicName id="4C6F4D17F02DFFB8FCF3FE67FC5CFE72" box="[806,954,393,418]" class="Insecta" family="Scarabaeidae" genus="Enicotarsus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B91BEA86F02DFFB8FCF3FE67FC5CFE72" box="[806,954,393,418]" italics="true" pageId="79" pageNumber="80">D. quadratus</emphasis>
</taxonomicName>
, the rounded pronotal lateral edges and the obtuse anterior angles in dorsal view will separate it from
<taxonomicName id="4C6F4D17F02DFFB8FCCBFE43FC51FE16" box="[798,951,429,454]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="B91BEA86F02DFFB8FCCBFE43FC51FE16" box="[798,951,429,454]" italics="true" pageId="79" pageNumber="80">D. nitidicollis</emphasis>
</taxonomicName>
and the elytral striae 1 bent laterally before reaching the apical margin will separate it from
<taxonomicName id="4C6F4D17F02DFFB8FD65FE3FFCF1FE39" box="[688,791,465,489]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="fasces">
<emphasis id="B91BEA86F02DFFB8FD65FE3FFCF1FE39" box="[688,791,465,489]" italics="true" pageId="79" pageNumber="80">D. fasces</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF02DFFA7FF12FE18FCCFFE7C" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="description">
<paragraph id="8BD03694F02DFFA7FF12FE18FE1AFF28" blockId="79.[151,1437,249,2038]" lastBlockId="80.[151,1436,151,933]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="B91BEA86F02DFFB8FF12FE18FEB7FDDE" bold="true" box="[199,337,501,526]" pageId="79" pageNumber="80">Description</emphasis>
.
<collectingCountry id="F3787604F02DFFB8FEB6FE18FE7BFDDE" box="[355,413,501,526]" name="Malta" pageId="79" pageNumber="80">Male</collectingCountry>
<typeStatus id="54D48836F02DFFB8FE7DFE18FDECFDDE" box="[424,522,501,526]" pageId="79" pageNumber="80" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F02DFFB8FDCBFE18FD90FDDE" box="[542,630,501,526]" captionStart="FIGURES 37 – 41" captionStartId="105.[151,264,1956,1979]" captionTargetBox="[258,1329,206,1918]" captionTargetId="figure@105.[242,1345,193,1935]" captionTargetPageId="105" captionText="FIGURES 37 – 41. Dorsal habitus. 37. D. latistriatus, male holotype; 38. D. nitidicollis, male lectotype; 39. D. quadratus, male neotype; 40. D. similis, male holotype; 41. D. denticollis, female holotype." httpUri="https://zenodo.org/record/268281/files/figure.png" pageId="79" pageNumber="80">Fig. 40</figureCitation>
).
<emphasis id="B91BEA86F02DFFB8FD58FE18FD2FFDDE" bold="true" box="[653,713,501,526]" pageId="79" pageNumber="80">Body</emphasis>
. Body small, length 9.0 mm, maximum width 4.0 mm; body subrectangular in dorsal view; dorsum narrowly flat.
<emphasis id="B91BEA86F02DFFB8FD3FFDF4FCCDFDE2" bold="true" box="[746,811,537,562]" pageId="79" pageNumber="80">Color</emphasis>
. Dorsal surface dark brown to black, glossy, with green and coppery metallic sheen; head brown along anterior edge of clypeus, with green and coppery metallic sheen on remaining surface; pronotum with green metallic sheen except for anteromedian carina, on anterior portion of disc and surface adjacent to lateral fossae; elytra with uniform green metallic sheen; ventrum dark brown to black; pygidium with green and coppery metallic sheen; legs reddish brown to dark brown.
<emphasis id="B91BEA86F02DFFB8FB59FD44FB2AFD12" bold="true" box="[1164,1228,681,706]" pageId="79" pageNumber="80">Head</emphasis>
. Clypeus broadly arcuate, anterior portion slightly upturned; clypeal teeth ogival; clypeal median emargination broadly u-shaped, clypeal edge emarginate on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with fine blunt transverse rugulae laterally, minutely punctate internally, lacking distinct transverse carina, simply convex; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes small in dorsal view, interocular ratio 5.5.
<emphasis id="B91BEA86F02DFFB8FBD9FC48FB63FC6E" bold="true" box="[1036,1157,933,958]" pageId="79" pageNumber="80">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.4; disc of pronotum minutely punctate on disc, punctures becoming fine on lateral declivities, with a fine longitudinal sulcus on posterior two-third; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a tri-sinuous carina, carina produced into a tubercle medially; anterior angles surface with fine blunt longitudinal rugulae, slightly but distinctly sulcate along posterior edge of anterior margin; lateral fossae rounded and bordered anteriorly by a sharp carina; lateral portions unmodified; pronotal basal fossae small, more or less rounded; posterior margin ill-defined on a short distance on each side of pronotal basal fossae, lacking setae.
<emphasis id="B91BEA86F02DFFB8FEE3FB4CFE64FB6A" bold="true" box="[310,386,1185,1210]" pageId="79" pageNumber="80">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.0; elytral base lacking distinct margin, simply convex; elytral striae 1–4 very wide basally and tapering toward apex, very deeply impressed basally, elytral striae 5 similar to 4 on disc, strial punctures minute, stria 1 well-defined apically, connecting to marginal stria; interstriae slightly convex, minutely punctate with fine irregular depressions throughout, with some microsculpture along striae.
<emphasis id="B91BEA86F02DFFB8FD31FADCFC44FA9A" bold="true" box="[740,930,1329,1354]" pageId="79" pageNumber="80">Thoracic sterna</emphasis>
. Proepisternal carina reduced, present along coxal insertion only; metasternal median lobe bluntly angularly produced anteromedially, ventral ridge ill-defined, keel shaped.
<emphasis id="B91BEA86F02DFFB8FEFCFA94FE87FA42" bold="true" box="[297,353,1401,1426]" pageId="79" pageNumber="80">Legs</emphasis>
. Profemur posterior surface slightly but distinctly convex, rather coarsely punctate and glabrous internally, posterointernal margin rather thick, evenly developed, internal edge rather wide, with a contiguous row of setae along anterointernal edge, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface lacking aligned row of setae internally, with few aligned and isolated setiferous punctures only, surface coarsely microsculptured between punctures; posterior surface with coarse irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur obtusely angular on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur internal edge nearly straight and lateral edge arcuate, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined irregular sulcus on apical half. Metatibia moderately slender, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy basally and slightly irregular apically, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with transverse microsculpture. Metatarsus 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F02DFFB8FAC3F854FF1AF826" bold="true" pageId="79" pageNumber="80">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally slightly convex; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 slightly longitudinally concave medially, shorter than segment 6 along midline; pygidium finely punctate on disc.
<emphasis id="B91BEA86F032FFA7FDF8FF51FD3AFF05" bold="true" box="[557,732,188,213]" pageId="80" pageNumber="81">
<collectingCountry id="F3787604F032FFA7FDF8FF51FD8CFF05" box="[557,618,188,213]" name="Malta" pageId="80" pageNumber="81">Male</collectingCountry>
genitalia
</emphasis>
(Figs. 106–107). Parameres produced onto a lobe laterally; surface smooth, glossy apically.
</paragraph>
<paragraph id="8BD03694F032FFA7FF12FEE9FD43FECC" blockId="80.[151,1436,151,933]" box="[199,677,260,285]" pageId="80" pageNumber="81">
<emphasis id="B91BEA86F032FFA7FF12FEE9FE90FECD" bold="true" box="[199,374,260,285]" pageId="80" pageNumber="81">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF032FFA7FE5DFEE9FE35FECD" box="[392,467,260,285]" pageId="80" pageNumber="81" type="male">1 male</specimenCount>
). Length: 9.0 mm.
</paragraph>
<paragraph id="8BD03694F032FFA7FF12FECAFCCFFE7C" blockId="80.[151,1436,151,933]" pageId="80" pageNumber="81">
<emphasis id="B91BEA86F032FFA7FF12FECAFE45FE90" bold="true" box="[199,419,295,320]" pageId="80" pageNumber="81">
Primary
<typeStatus id="54D48836F032FFA7FEE6FECAFE80FE90" box="[307,358,295,320]" pageId="80" pageNumber="81">type</typeStatus>
data
</emphasis>
(
<figureCitation id="13542A11F032FFA7FE67FECAFDF3FE90" box="[434,533,295,320]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="80" pageNumber="81">Fig. 154</figureCitation>
).
<typeStatus id="54D48836F032FFA7FDFDFECAFD76FE90" box="[552,656,295,320]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
male (MTD): [
<collectingCountry id="F3787604F032FFA7FCE9FEC5FC40FE90" box="[828,934,296,320]" name="Paraguay" pageId="80" pageNumber="81">Paraguay</collectingCountry>
/ Dr Bohls]/ [♂]; [Coll. C. Felsche/ Kauf 20, 1918] green card; [
<typeStatus id="54D48836F032FFA7FEA1FEA0FE58FEB5" box="[372,446,333,357]" pageId="80" pageNumber="81">Typus</typeStatus>
] red card; [
<taxonomicName id="4C6F4D17F032FFA7FD93FEA1FD13FEB5" box="[582,757,332,357]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F032FFA7FCDDFEA1FC6AFEB5" authority="Blut" authorityName="Blut" box="[776,908,332,357]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="similis">similis Blut</taxonomicName>
/
<collectingCountry id="F3787604F032FFA7FC75FEA0FBEFFEB5" box="[928,1033,333,357]" name="Paraguay" pageId="80" pageNumber="81">Paraguay</collectingCountry>
, Dr. Bohls] hanwritten; [WORLD/ SCARAB./ DATABASE/ WSD00016771]; [
<typeStatus id="54D48836F032FFA7FD78FE9DFCA5FE58" box="[685,835,368,392]" pageId="80" pageNumber="81" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6F4D17F032FFA7FCB5FE82FBEBFE58" box="[864,1037,367,392]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F032FFA7FCB5FE82FBEBFE58" box="[864,1037,367,392]" italics="true" pageId="80" pageNumber="81">Dendropaemon</emphasis>
</taxonomicName>
/
<taxonomicName id="4C6F4D17F032FFA7FBFDFE82FB97FE58" box="[1064,1137,367,392]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91BEA86F032FFA7FBFDFE82FB97FE58" box="[1064,1137,367,392]" italics="true" pageId="80" pageNumber="81">similis</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFFE4B65F032FFA7FB59FE82FAF4FE58" author="Blut" box="[1164,1298,367,392]" pageId="80" pageNumber="81" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut, 1939</bibRefCitation>
] red card; [
<taxonomicName id="4C6F4D17F032FFA7FF48FE79FEAAFE7D" box="[157,332,404,429]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F032FFA7FE89FE79FE40FE7D" box="[348,422,404,429]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="similis">similis</taxonomicName>
/
<bibRefCitation id="EFFE4B65F032FFA7FE60FE79FDCCFE7C" author="Blut" box="[437,554,404,429]" pageId="80" pageNumber="81" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut, 1939</bibRefCitation>
/ vid. F. Génier, 2014].
</paragraph>
</subSubSection>
<subSubSection id="C375651FF032FFA7FF12FE5AFDE4FE25" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="8BD03694F032FFA7FF12FE5AFD61FE00" blockId="80.[151,1436,151,933]" box="[199,647,439,464]" pageId="80" pageNumber="81">
<emphasis id="B91BEA86F032FFA7FF12FE5AFE4FFE00" bold="true" box="[199,425,439,464]" pageId="80" pageNumber="81">Material examined</emphasis>
. Primary
<typeStatus id="54D48836F032FFA7FDCDFE55FDAEFE00" box="[536,584,440,464]" pageId="80" pageNumber="81">type</typeStatus>
only.
</paragraph>
<paragraph id="8BD03694F032FFA7FF12FE31FDE4FE25" blockId="80.[151,1436,151,933]" box="[199,514,476,501]" pageId="80" pageNumber="81">
<emphasis id="B91BEA86F032FFA7FF12FE31FE9BFE25" bold="true" box="[199,381,476,501]" pageId="80" pageNumber="81">Natural history</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF032FFA7FF12FE12FC0CFC75" pageId="80" pageNumber="81" type="discussion">
<paragraph id="8BD03694F032FFA7FF12FE12FD43FDC8" blockId="80.[151,1436,151,933]" box="[199,677,511,536]" pageId="80" pageNumber="81">
<emphasis id="B91BEA86F032FFA7FF12FE12FED2FDC8" bold="true" box="[199,308,511,536]" pageId="80" pageNumber="81">Remarks</emphasis>
. Female and variation unknown.
</paragraph>
<paragraph id="8BD03694F032FFA7FF12FDC9FC68FD1D" blockId="80.[151,1436,151,933]" pageId="80" pageNumber="81">
<collectingCountry id="F3787604F032FFA7FF12FDC9FF00FDEC" box="[199,230,548,572]" name="American Samoa" pageId="80" pageNumber="81">As</collectingCountry>
noted by
<bibRefCitation id="EFFE4B65F032FFA7FE8FFDC9FE39FDEC" author="Blut" box="[346,479,548,573]" pageId="80" pageNumber="81" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut (1939)</bibRefCitation>
, this species is externally extremely similar to
<taxonomicName id="4C6F4D17F032FFA7FC22FDC8FBB9FDEC" box="[1015,1119,548,572]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fasces">
<emphasis id="B91BEA86F032FFA7FC22FDC8FBB9FDEC" box="[1015,1119,548,572]" italics="true" pageId="80" pageNumber="81">D. fasces</emphasis>
</taxonomicName>
. In the original description, Blut states that the
<typeStatus id="54D48836F032FFA7FE54FDAAFE05FDB0" box="[385,483,583,608]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
male is from San
<collectingCountry id="F3787604F032FFA7FD12FDAAFCCDFDB0" box="[711,811,583,608]" name="El Salvador" pageId="80" pageNumber="81">Salvador</collectingCountry>
(
<collectingCountry id="F3787604F032FFA7FCEBFDA5FC4BFDB0" box="[830,941,584,608]" name="Paraguay" pageId="80" pageNumber="81">Paraguay</collectingCountry>
). However, the locality data labels on the specimen only state “
<collectingCountry id="F3787604F032FFA7FE47FD81FDE4FD54" box="[402,514,620,644]" name="Paraguay" pageId="80" pageNumber="81">Paraguay</collectingCountry>
”. Two specimens of
<taxonomicName id="4C6F4D17F032FFA7FD2CFD80FC85FD54" box="[761,867,620,644]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fasces">
<emphasis id="B91BEA86F032FFA7FD2CFD80FC85FD54" box="[761,867,620,644]" italics="true" pageId="80" pageNumber="81">D. fasces</emphasis>
</taxonomicName>
studied by Blut are clearly labeled as from San
<collectingCountry id="F3787604F032FFA7FF42FD62FF1DFD78" box="[151,251,655,680]" name="El Salvador" pageId="80" pageNumber="81">Salvador</collectingCountry>
and both of these specimens are from the
<typeStatus id="54D48836F032FFA7FD0CFD7DFCEFFD78" box="[729,777,656,680]" pageId="80" pageNumber="81">type</typeStatus>
series of
<taxonomicName id="4C6F4D17F032FFA7FCA2FD7CFC39FD78" box="[887,991,656,680]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fasces">
<emphasis id="B91BEA86F032FFA7FCA2FD7CFC39FD78" box="[887,991,656,680]" italics="true" pageId="80" pageNumber="81">D. fasces</emphasis>
</taxonomicName>
studied by Blut and are identical to the
<typeStatus id="54D48836F032FFA7FF42FD59FEE4FD1D" box="[151,258,692,717]" pageId="80" pageNumber="81" type="lectotype">lectotype</typeStatus>
. This is certainly casting doubts on the exact
<typeStatus id="54D48836F032FFA7FD28FD58FCCBFD1D" box="[765,813,693,717]" pageId="80" pageNumber="81">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BD03694F032FFA7FF12FD35FC0CFC75" blockId="80.[151,1436,151,933]" pageId="80" pageNumber="81">
In addition to characters used in the description the following differences can also separate
<taxonomicName id="4C6F4D17F032FFA7FB1AFD34FADDFD20" box="[1231,1339,727,752]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B91BEA86F032FFA7FB1AFD34FADDFD20" box="[1231,1339,727,752]" italics="true" pageId="80" pageNumber="81">D. similis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C6F4D17F032FFA7FA54FD34FF3DFCC4" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fasces">
<emphasis id="B91BEA86F032FFA7FA54FD34FF3DFCC4" italics="true" pageId="80" pageNumber="81">D. fasces</emphasis>
</taxonomicName>
: the punctures of the posterior portion of the profemur are much larger and confluent, the surface exhibit coarse microsculpture, the same is true for the meso- and metafemora. Although much more difficult to quantify and describe, the posterior portion of the pronotum is slightly more convex laterally, the posterior border is, as a result, well defined on a longer distance. Finally, the pygidial surface show distinct microsculpture and the parameres are less explanate apically in dorsal view and are slightly shorter.
</paragraph>
</subSubSection>
</treatment>
</document>
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