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<document id="C1149CBDD0DB0EC79B9E97636F616919" ID-CLB-Dataset="28891" ID-DOI="10.5852/ejt.2018.464" ID-GBIF-Dataset="2fc5fa89-0438-4096-b785-78d14b6bfb76" ID-ISSN="2118-9773" ID-Zenodo-Dep="1469475" ID-ZooBank="urn:lsid:zoobank.org:pub:3C501DE6-AC85-4199-8CDC-D28E42753050" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1540291417600" checkinUser="plazi" docAuthor="Zallot, Enrico, Fehér, Zoltán, Bamberger, Sonja &amp; Gittenberger, Edmund" docDate="2018" docId="03C6575CFFFEFFE0C121E456FBDCFA40" docLanguage="en" docName="ejt-464_zallot_feher_bamberger_gittenberger.pdf.imf" docOrigin="European Journal of Taxonomy 464" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Lovcenia) lanatum Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger 2018, sp. nov." docType="treatment" docUuid="E615F65C-B3DC-4978-883C-4D10A06606E6" docUuidSource="ZooBank" docVersion="14" lastPageNumber="22" masterDocId="FFFF2F24FFEAFFF5C052E55DFF9AFFC3" masterDocTitle="Cochlostoma revised: the subgenus Lovcenia Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="21" updateTime="1701997563338" updateUser="plazi">
<mods:mods id="433511E630A230BBD5F0F047C0A6885F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F1792A17F2900EF1762EACB459FE398">
<mods:title id="6160DD1A109006D588022F7048646637">Cochlostoma revised: the subgenus Lovcenia Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="CC4197C5D4A7E50FC7EA53BF263BCC93" type="personal">
<mods:role id="A3D2BA186798952C38E014D929B3E205">
<mods:roleTerm id="21E5B4E24790AC7D923D0441A51DFCC1">Author</mods:roleTerm>
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<mods:namePart id="0F4C99909EF423BAE2B42B72EB90CB31">Zallot, Enrico</mods:namePart>
</mods:name>
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<mods:namePart id="EFC47075A975227F22E96687171C1E3F">Fehér, Zoltán</mods:namePart>
</mods:name>
<mods:name id="D0C43FE3259117CFEF064A9D1487D082" type="personal">
<mods:role id="0D81A866C2B58CDC5A83FDD5CF9F5BE8">
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</mods:role>
<mods:namePart id="AB56E5725649239B74B05823604D1041">Bamberger, Sonja</mods:namePart>
</mods:name>
<mods:name id="60FF9998A1C9977A3BD9145D3556386B" type="personal">
<mods:role id="57FBCE41D47A628B880853DBA8E1A6EE">
<mods:roleTerm id="43B17CBE2CED46288EF2668D49C136A9">Author</mods:roleTerm>
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<mods:namePart id="123C57526BD7DACD26E51F8149BB68E0">Gittenberger, Edmund</mods:namePart>
</mods:name>
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<mods:title id="6A9D6AACAA83461C6313A7752FC0EBB2">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7CBFE228A51D5F5F53FC0C0AC492571E">
<mods:date id="0F6E9B0B3E8EA15ADF617FBC607084DC">2018</mods:date>
<mods:detail id="2A1E97DA70FCB53A252FABC448159E6B" type="pubDate">
<mods:number id="63C1B59FD3760DDADA481B3EB1DA7655">2018-10-02</mods:number>
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<mods:number id="0706375EBC1F9B5CB9C68B894D5806CE">464</mods:number>
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<mods:start id="2E4D2C7ADC680DA4E6EF2C58191DD79B">1</mods:start>
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<mods:classification id="6C0D8554683E37FEF8696A33B238788B">journal article</mods:classification>
<mods:identifier id="60492C198EC01E05E0A13089CB2B0E92" type="CLB-Dataset">28891</mods:identifier>
<mods:identifier id="397FCF76D06A8667A3AFA94E3B99DFA2" type="DOI">10.5852/ejt.2018.464</mods:identifier>
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<treatment id="03C6575CFFFEFFE0C121E456FBDCFA40" ID-DOI="http://doi.org/10.5281/zenodo.5631647" ID-GBIF-Taxon="148967379" ID-Zenodo-Dep="5631647" LSID="urn:lsid:zoobank.org:act:E615F65C-B3DC-4978-883C-4D10A06606E6" httpUri="http://treatment.plazi.org/id/03C6575CFFFEFFE0C121E456FBDCFA40" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="C375B5C1FFFEFFE1C121E456FB59FE8A" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BD0E64AFFFEFFE1C121E456FB59FE8A" blockId="20.[369,1219,267,365]" pageId="20" pageNumber="21">
<taxonomicName id="4C6F9DC9FFFEFFE1C121E456FBC5FEE6" authority="Zallot, Feher &amp; Gittenberger" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[371,1119,267,294]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="lanatum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFEFFE1C121E456FC9DFEE5" bold="true" box="[371,775,267,294]" italics="true" pageId="20" pageNumber="21">Cochlostoma (Lovcenia) lanatum</emphasis>
Zallot, Fehér &amp; Gittenberger
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFEFFE1C434E451FB5BFEE5" box="[1126,1217,268,294]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
urn:lsid:zoobank.org:act:E615F65C-B3DC-4978-883C-4D10A06606E6
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFEFFE1C2EDE40FFCEEFEAF" box="[703,884,338,365]" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BD0E64AFFFEFFE1C2EDE40FFCEEFEAF" blockId="20.[369,1219,267,365]" box="[703,884,338,365]" pageId="20" pageNumber="21">
<figureCitation id="1354FACFFFFEFFE1C2EDE40FFC86FEAE" box="[703,796,338,365]" captionStart="Fig. 5" captionStartId="10.[189,232,1859,1885]" captionTargetBox="[189,1398,625,1829]" captionTargetId="figure@10.[189,1398,625,1829]" captionTargetPageId="10" captionText="Fig. 5. Female specimens of Cochlostoma (Lovcenia) Zallot et al., 2015 used for anatomy (shells were destroyed before dissection). A. C. (Lovcenia) erika (A.J. Wagner, 1906) (HNHM 86387). B. C. (Lovcenia) dalmatinum (L. Pfeiffer, 1863) (NHMW 110430/MN/0162). C. C. (Lovcenia) tropojanum sp. nov. (NHMW 111248). D. C. (Lovcenia) jakschae sp. nov. (NHMW 111249). E. C. (Lovcenia) lanatum sp. nov. (HNHM 99861)." httpUri="https://zenodo.org/record/1469485/files/figure.png" pageId="20" pageNumber="21">Figs 5E</figureCitation>
,
<figureCitation id="1354FACFFFFEFFE1C37AE40FFCD1FEAF" box="[808,843,338,364]" captionStart="Fig. 6" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[191,1395,878,1778]" captionTargetId="figure@11.[191,1474,844,1819]" captionTargetPageId="11" captionText="Fig. 6. Female genital morphology. See Table 1 for localities and Table 3 for measurements. A. Cochlostoma (Lovcenia) erika (A.J. Wagner, 1906) (HNHM 86387). B. Cochlostoma (Lovcenia) dalmatinum (L. Pfeiffer, 1863) (NHMW 110430/MN/0162). C. Cochlostoma (Lovcenia) tropojanum sp. nov. (NHMW 111248). D1. Cochlostoma (Lovcenia) jakschae sp. nov., Bugarska Peak population (NHMW 111249). D2. Cochlostoma (Lovcenia) jakschae sp. nov., Pejë Pass population (HNHM 97221). E. Cochlostoma (Lovcenia) lanatum sp. nov. (HNHM 99861)." httpUri="https://zenodo.org/record/1469487/files/figure.png" pageId="20" pageNumber="21">6E</figureCitation>
,
<figureCitation id="1354FACFFFFEFFE1C305E40FFCEEFEAF" box="[855,884,338,364]" captionStart="Fig. 12" captionStartId="20.[189,232,1964,1990]" captionTargetBox="[189,1398,1182,1935]" captionTargetId="figure@20.[189,1398,1182,1935]" captionTargetPageId="20" captionText="Fig. 12. Cochlostoma (Lovcenia) lanatum Zallot, Fehér &amp; Gittenberger sp. nov., holotype, Deleve Cave, Albania (HNHM 102806)." httpUri="https://zenodo.org/record/1469499/files/figure.png" pageId="20" pageNumber="21">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFEFFE1C0EFE4CEFCB8FE6E" box="[189,802,403,430]" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8BD0E64AFFFEFFE1C0EFE4CEFCB8FE6E" blockId="20.[189,802,403,430]" box="[189,802,403,430]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="AB7FC164FFFEFFE1C0EFE4CEFCB8FE6E" box="[189,802,403,430]" pageId="20" pageNumber="21">
<taxonomicName id="4C6F9DC9FFFEFFE1C0EFE4CEFD9DFE6D" authority="Zallot, Fehér &amp; Gittenberger, 2015" authorityName="Zallot, Fehér &amp; Gittenberger" authorityYear="2015" box="[189,519,403,430]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="sp.">
<emphasis id="B91B3A58FFFEFFE1C0EFE4CEFE45FE6D" box="[189,479,403,430]" italics="true" pageId="20" pageNumber="21">Cochlostoma (Lovcenia)</emphasis>
sp.
</taxonomicName>
–
<treatmentCitation id="0ACEC05BFFFEFFE1C276E4CEFC84FE6E" author="" box="[548,798,403,430]" page="80" pageId="20" pageNumber="21">
<bibRefCitation id="EFFE9BBBFFFEFFE1C276E4CEFD69FE6D" box="[548,755,403,430]" pageId="20" pageNumber="21" refId="ref9534" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article">
Zallot
<emphasis id="B91B3A58FFFEFFE1C220E4C9FD36FE6E" box="[626,684,403,429]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015
</bibRefCitation>
: 80
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFEFFE1C0EFE489FBD8FD83" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BD0E64AFFFEFFE1C0EFE489FEACFE2D" blockId="20.[189,310,468,494]" box="[189,310,468,494]" pageId="20" pageNumber="21">
<heading id="D0985126FFFEFFE1C0EFE489FEACFE2D" bold="true" box="[189,310,468,494]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B91B3A58FFFEFFE1C0EFE489FEACFE2D" bold="true" box="[189,310,468,494]" pageId="20" pageNumber="21">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFEFFE1C0EFE75EFBD8FD83" blockId="20.[189,1399,515,577]" pageId="20" pageNumber="21">Shell without spots or color bands, relatively large and slender, with sharp, regularly spaced ribs on the upper whorls; ribs less sharp and more narrowly spaced on the lower whorls.</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFEFFE1C0EFE735FF72FC06" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BD0E64AFFFEFFE1C0EFE735FE30FD41" blockId="20.[189,426,616,642]" box="[189,426,616,642]" pageId="20" pageNumber="21">
<heading id="D0985126FFFEFFE1C0EFE735FE30FD41" bold="true" box="[189,426,616,642]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B91B3A58FFFEFFE1C0EFE735FE30FD41" bold="true" box="[189,426,616,642]" pageId="20" pageNumber="21">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFEFFE1C0B8E7CAFDDCFCD8" blockId="20.[189,1398,663,795]" pageId="20" pageNumber="21">
<materialsCitation id="3B07EC17FFFEFFE1C0B8E7CAFDDBFCD8" ID-GBIF-Occurrence="1931541542" collectedFrom="limestone rock within cave" collectingDate="2016-06-28" collectionCode="HNHM" collectorName="Erőss &amp; Fehér &amp; Szekeres &amp; Grego" country="Albania" county="Malësia District" elevation="1300" latitude="42.3576" location="SW slope of Mali e Veleçikut, Shpellë e Deleve, NW of Razëm" longLatPrecision="6" longitude="19.5269" municipality="Shpelle" pageId="20" pageNumber="21" specimenCode="HNHM 102806" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D458E8FFFEFFE1C0B8E7CAFEC6FD72" box="[234,348,663,689]" pageId="20" pageNumber="21" type="holotype">
<emphasis id="B91B3A58FFFEFFE1C0B8E7CAFEC6FD72" bold="true" box="[234,348,663,689]" pageId="20" pageNumber="21">Holotype</emphasis>
</typeStatus>
<collectingCountry id="F378A6DAFFFEFFE1C0EFE7E7FEDCFD17" box="[189,326,698,724]" name="Albania" pageId="20" pageNumber="21">ALBANIA</collectingCountry>
:
<specimenCount id="9D692DC3FFFEFFE1C107E7E7FEF1FD17" box="[341,363,698,724]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
,
<collectingCounty id="62B19EC6FFFEFFE1C12DE7E4FDD9FD10" box="[383,579,697,723]" pageId="20" pageNumber="21">Malësia District</collectingCounty>
,
<location id="8EB0B091FFFEFFE1C201E7E7FAE8FD17" LSID="urn:lsid:plazi:treatment:03C6575CFFFEFFE0C121E456FBDCFA40:8EB0B091FFFEFFE1C201E7E7FAE8FD17" box="[595,1394,697,724]" country="Albania" county="Malësia District" latitude="42.3576" longLatPrecision="6" longitude="19.5269" municipality="Shpelle" name="SW slope of Mali e Veleçikut, Shpellë e Deleve, NW of Razëm" pageId="20" pageNumber="21">SW slope of Mali e Veleçikut, Shpellë e Deleve, NW of Razëm</location>
,
<elevation id="00420179FFFEFFE1C0EFE780FEC9FD34" box="[189,339,733,759]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="21" unit="m" value="1300.0">
<quantity id="4C974BAFFFFEFFE1C0EFE780FE8FFD34" box="[189,277,733,759]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="21" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
,
<collectedFrom id="3873A5B7FFFEFFE1C132E780FD07FD34" box="[352,669,733,759]" pageId="20" pageNumber="21">limestone rock within cave</collectedFrom>
,
<geoCoordinate id="EE5B808DFFFEFFE1C2FAE781FCABFD34" box="[680,817,732,759]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="42.3576">42.3576° N</geoCoordinate>
,
<geoCoordinate id="EE5B808DFFFEFFE1C36EE781FC5BFD34" box="[828,961,732,759]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="19.5269">19.5269° E</geoCoordinate>
,
<date id="FFD1C08AFFFEFFE1C39FE780FBFFFD34" box="[973,1125,733,759]" pageId="20" pageNumber="21" value="2016-06-28">
<collectingDate id="EF953962FFFEFFE1C39FE780FBFFFD34" box="[973,1125,733,759]" pageId="20" pageNumber="21" value="2016-06-28">28 Jun. 2016</collectingDate>
</date>
,
<collectorName id="269A839CFFFEFFE1C422E780FB2EFD34" box="[1136,1204,733,759]" pageId="20" pageNumber="21">Erőss</collectorName>
,
<collectorName id="269A839CFFFEFFE1C4EDE780FA9EFD34" box="[1215,1284,733,759]" pageId="20" pageNumber="21">Fehér</collectorName>
,
<collectorName id="269A839CFFFEFFE1C55CE780FAECFD34" box="[1294,1398,733,759]" pageId="20" pageNumber="21">Szekeres</collectorName>
and
<collectorName id="269A839CFFFEFFE1C0BDE65DFEA2FCD9" box="[239,312,768,794]" pageId="20" pageNumber="21">Grego</collectorName>
leg. (
<specimenCode id="DBC94E31FFFEFFE1C128E65DFDA3FCD8" box="[378,569,768,795]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="20" pageNumber="21">HNHM 102806</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD0E64AFFFEFFE1C0B8E61FFF72FC06" blockId="20.[189,1399,834,965]" pageId="20" pageNumber="21">
<materialsCitation id="3B07EC17FFFEFFE1C0B8E61FFD2AFC60" ID-GBIF-Occurrence="1931541274" collectingDate="52-11-16" collectionCode="HNHM, NHMW" pageId="20" pageNumber="21" specimenCode="NHMW 111652, HNHM 99861">
<emphasis id="B91B3A58FFFEFFE1C0B8E61FFEFDFC9F" bold="true" box="[234,359,834,860]" pageId="20" pageNumber="21">Paratypes</emphasis>
<collectingCountry id="F378A6DAFFFEFFE1C0EFE638FEDCFCBC" box="[189,326,869,895]" name="Albania" pageId="20" pageNumber="21">ALBANIA</collectingCountry>
: 129 spec., same collection data as for holotype (
<specimenCode id="DBC94E31FFFEFFE1C3E4E638FBE6FCBC" box="[950,1148,868,895]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">
NHMW
<date id="FFD1C08AFFFEFFE1C476E639FBE6FCBC" box="[1060,1148,868,895]" pageId="20" pageNumber="21" value="52-11-16">111652</date>
</specimenCode>
/7+ 12fr.+19a+ 6 ja; GR/38+2fr;
<specimenCode id="DBC94E31FFFEFFE1C11DE6D5FE64FC61" box="[335,510,904,930]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="20" pageNumber="21">HNHM 99861</specimenCode>
/6+13fr.+ 26a);
</materialsCitation>
<materialsCitation id="3B07EC17FFFEFFE1C2EBE6D5FF72FC06" ID-GBIF-Occurrence="1931541270" collectingDate="2011-10-16" collectorName="Gittenberger" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D692DC3FFFEFFE1C2EBE6D5FD73FC61" box="[697,745,904,930]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
, same locality as holotype,
<date id="FFD1C08AFFFEFFE1C465E6D5FB4FFC61" box="[1079,1237,904,930]" pageId="20" pageNumber="21" value="2011-10-16">
<collectingDate id="EF953962FFFEFFE1C465E6D5FB4FFC61" box="[1079,1237,904,930]" pageId="20" pageNumber="21" value="2011-10-16">16 Oct. 2011</collectingDate>
</date>
,
<collectorName id="269A839CFFFEFFE1C4B1E6D5FAEDFC61" box="[1251,1399,904,930]" pageId="20" pageNumber="21">Gittenberger</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFEFFE0C0EFE6B0FC66FD36" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="8BD0E64AFFFEFFE1C0EFE6B0FEEFFBC4" blockId="20.[189,373,1005,1031]" box="[189,373,1005,1031]" pageId="20" pageNumber="21">
<heading id="D0985126FFFEFFE1C0EFE6B0FEEFFBC4" bold="true" box="[189,373,1005,1031]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B91B3A58FFFEFFE1C0EFE6B0FEEFFBC4" bold="true" box="[189,373,1005,1031]" pageId="20" pageNumber="21">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFEFFE1C0EFE146FBBCFBF6" blockId="20.[189,1062,1051,1078]" box="[189,1062,1051,1078]" pageId="20" pageNumber="21">
<typeStatus id="54D458E8FFFEFFE1C0EFE146FEB5FBF6" box="[189,303,1051,1077]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
: H
<quantity id="4C974BAFFFFEFFE1C104E146FE58FBF6" box="[342,450,1051,1078]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="mm" value="10.5">10.5 mm</quantity>
, W
<quantity id="4C974BAFFFFEFFE1C1BDE146FDD6FBF6" box="[495,588,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="20" pageNumber="21" unit="mm" value="4.6">4.6 mm</quantity>
, Wbw
<quantity id="4C974BAFFFFEFFE1C2CCE146FD61FBF6" box="[670,763,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="mm" value="3.8">3.8 mm</quantity>
, Ha
<quantity id="4C974BAFFFFEFFE1C362E146FC17FBF6" box="[816,909,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="20" pageNumber="21" unit="mm" value="3.4">3.4 mm</quantity>
, Wa
<quantity id="4C974BAFFFFEFFE1C397E146FBB8FBF6" box="[965,1058,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="20" pageNumber="21" unit="mm" value="3.1">3.1 mm</quantity>
.
</paragraph>
<paragraph id="8BD0E64AFFFEFFE1C0EFE100FAEDFBB4" blockId="20.[189,1399,1116,1143]" box="[189,1399,1116,1143]" pageId="20" pageNumber="21">
<typeStatus id="54D458E8FFFEFFE1C0EFE100FEB5FBB4" box="[189,303,1117,1143]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
(N=12): H
<quantity id="4C974BAFFFFEFFE1C1ECE101FDD4FBB4" box="[446,590,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.5" metricValueMin="8.4" pageId="20" pageNumber="21" unit="mm" value="8.95" valueMax="9.5" valueMin="8.4">8.4–9.5 mm</quantity>
, W
<quantity id="4C974BAFFFFEFFE1C22BE101FC93FBB4" box="[633,777,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="20" pageNumber="21" unit="mm" value="4.15" valueMax="4.4" valueMin="3.9">3.9–4.4 mm</quantity>
, Wbw
<quantity id="4C974BAFFFFEFFE1C30BE101FC73FBB4" box="[857,1001,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="20" pageNumber="21" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2–3.5 mm</quantity>
, Ha
<quantity id="4C974BAFFFFEFFE1C44EE101FB36FBB4" box="[1052,1196,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="20" pageNumber="21" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
, Wa
<quantity id="4C974BAFFFFEFFE1C4B1E101FAE8FBB4" box="[1251,1394,1116,1143]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="20" pageNumber="21" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">2.5–3.2 mm</quantity>
.
</paragraph>
<caption id="DF10B6C2FFFEFFE1C0EFE2F1FE6FF828" httpUri="https://zenodo.org/record/1469499/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1964,1990]" targetBox="[189,1398,1182,1935]" targetPageId="20">
<paragraph id="8BD0E64AFFFEFFE1C0EFE2F1FE6FF828" blockId="20.[189,1398,1964,2027]" pageId="20" pageNumber="21">
<emphasis id="B91B3A58FFFEFFE1C0EFE2F1FE82F805" bold="true" box="[189,280,1964,1990]" pageId="20" pageNumber="21">Fig. 12.</emphasis>
<taxonomicName id="4C6F9DC9FFFEFFE1C14CE2F1FC63F805" authority="Zallot, Feher &amp; Gittenberger" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[286,1017,1964,1991]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="lanatum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFEFFE1C14CE2F1FD3FF805" box="[286,677,1964,1991]" italics="true" pageId="20" pageNumber="21">Cochlostoma (Lovcenia) lanatum</emphasis>
Zallot, Fehér &amp; Gittenberger
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFEFFE1C3ADE2F0FBC1F804" box="[1023,1115,1965,1991]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Deleve Cave, Albania (HNHM 102806).
</paragraph>
</caption>
<paragraph id="8BD0E64AFFFFFFE0C0EFE456FED4FEE6" blockId="21.[189,334,267,290]" box="[189,334,267,293]" pageId="21" pageNumber="22">
<emphasis id="B91B3A58FFFFFFE0C0EFE456FED4FEE6" bold="true" box="[189,334,267,293]" pageId="21" pageNumber="22">Description</emphasis>
</paragraph>
<paragraph id="8BD0E64AFFFFFFE0C0EFE467FBAEFDE4" blockId="21.[189,1399,312,552]" pageId="21" pageNumber="22">SHELL. Rather large and slender, with 8½–9¼ whorls (H/W ratio 2.84). The apical 2.2 whorls form the protoconch, which is smooth for only the initial 0.2 whorl and finely ribbed with close riblets further on. The shell is more or less light horn-brown, without spots. The teleoconch is sculptured with rather prominent, sharp ribs, which are widely spaced on the upper whorls and increasingly more narrowly spaced and less sharp towards the aperture. The aperture has a moderately developed lip, enlarged at the columellar lobe, which is abruptly bent inwards to cover the umbilicus. While approaching the aperture, the body whorl abruptly enlarges, forming a rather prominent external lobe.</paragraph>
<paragraph id="8BD0E64AFFFFFFE0C0EFE712FC66FD36" blockId="21.[189,1399,590,757]" pageId="21" pageNumber="22">FEMALE GENITALIA. Bursa copulatrix large, with a pedunculus connected proximally; its proximal lobe smaller that the distal one. The short seminal receptacle is confined to the ventral side of the body, under (ventrally) the bursa copulatrix. The loops of the visceral oviduct are situated over the apex of the seminal receptacle. The junction of the uterine gland and the copulatory duct is far away from the connection between the pedunculus of the bursa and the distal oviduct.</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFFFFE0C0EFE640FD92FC4B" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BD0E64AFFFFFFE0C0EFE640FECCFCF4" blockId="21.[189,342,797,823]" box="[189,342,797,823]" pageId="21" pageNumber="22">
<heading id="D0985126FFFFFFE0C0EFE640FECCFCF4" bold="true" box="[189,342,797,823]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B91B3A58FFFFFFE0C0EFE640FECCFCF4" bold="true" box="[189,342,797,823]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFFFFE0C0EFE616FD92FC4B" blockId="21.[189,1398,843,904]" pageId="21" pageNumber="22">This species is only known from its type locality, the Deleve Cave on the southwestern slope of Mount Veleçik in northern Albania.</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFFFFE0C0EFE6EDFA8EFB45" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="8BD0E64AFFFFFFE0C0EFE6EDFE32FC09" blockId="21.[189,1398,944,1158]" box="[189,424,944,970]" pageId="21" pageNumber="22">
<heading id="D0985126FFFFFFE0C0EFE6EDFE32FC09" bold="true" box="[189,424,944,970]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B91B3A58FFFFFFE0C0EFE6EDFE32FC09" bold="true" box="[189,424,944,970]" pageId="21" pageNumber="22">Habitat preference</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFFFFE0C0EFE683FA8EFB45" blockId="21.[189,1398,944,1158]" pageId="21" pageNumber="22">
Like the majority of known species of
<taxonomicName id="4C6F9DC9FFFFFFE0C22EE683FC8DFC3B" box="[636,791,990,1016]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B91B3A58FFFFFFE0C22EE683FC8DFC3B" box="[636,791,990,1016]" italics="true" pageId="21" pageNumber="22">Cochlostoma</emphasis>
</taxonomicName>
, this is an obligate rock-dwelling snail. It was found in the twilight zone inside the Deleve Cave. Though little is known about its ecological requirements, it may be restricted to the epigean environment, like
<taxonomicName id="4C6F9DC9FFFFFFE0C36BE178FC54FBFC" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1906" box="[825,974,1061,1087]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="erika" subGenus="Lovcenia">
<emphasis id="B91B3A58FFFFFFE0C36BE178FC54FBFC" box="[825,974,1061,1087]" italics="true" pageId="21" pageNumber="22">C. (L.) erika</emphasis>
</taxonomicName>
(i.e., subtroglophile in the sense of
<bibRefCitation id="EFFE9BBBFFFFFFE0C0EFE115FEA2FBA1" box="[189,312,1096,1122]" pageId="21" pageNumber="22" refId="ref9015" refString="Sket B. 2008. Can we agree on an ecological classification of subterranean animals? Journal of Natural History 42 (21 - 22): 1549 - 1563. https: // doi. org / 10.1080 / 00222930801995762" type="journal article">Sket 2008</bibRefCitation>
). At the Deleve Cave, its niche partitioning with
<taxonomicName id="4C6F9DC9FFFFFFE0C33CE115FE98FB46" authority="(A. J. Wagner, 1901)" authorityName="A.J. Wagner" authorityYear="1901" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1901" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="roseoli" subGenus="Auritus">
<emphasis id="B91B3A58FFFFFFE0C33CE115FB4AFBA1" box="[878,1232,1096,1122]" italics="true" pageId="21" pageNumber="22">Cochlostoma (Auritus) roseoli</emphasis>
(A.J. Wagner, 1901)
</taxonomicName>
is similar to that of
<taxonomicName id="4C6F9DC9FFFFFFE0C1BCE131FDE4FB46" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1906" box="[494,638,1131,1158]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="erika" subGenus="Lovcenia">
<emphasis id="B91B3A58FFFFFFE0C1BCE131FDE4FB46" box="[494,638,1131,1158]" italics="true" pageId="21" pageNumber="22">C. (L.) erika</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F9DC9FFFFFFE0C2EAE131FCF1FB46" baseAuthorityName="Rossmassler" baseAuthorityYear="1837" box="[696,875,1132,1158]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="auritum" subGenus="Auritus">
<emphasis id="B91B3A58FFFFFFE0C2EAE131FCF1FB46" box="[696,875,1132,1158]" italics="true" pageId="21" pageNumber="22">C. (A.) auritum</emphasis>
</taxonomicName>
at the Popovo Cave in
<collectingCountry id="F378A6DAFFFFFFE0C42EE131FA95FB45" box="[1148,1295,1132,1158]" name="Serbia" pageId="21" pageNumber="22">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFFFFE0C0EFE1F0FBDCFA40" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BD0E64AFFFFFFE0C0EFE1F0FEB5FB04" blockId="21.[189,303,1197,1223]" box="[189,303,1197,1223]" pageId="21" pageNumber="22">
<heading id="D0985126FFFFFFE0C0EFE1F0FEB5FB04" bold="true" box="[189,303,1197,1223]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B91B3A58FFFFFFE0C0EFE1F0FEB5FB04" bold="true" box="[189,303,1197,1223]" pageId="21" pageNumber="22">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFFFFE0C0EFE186FBDCFA40" blockId="21.[189,1399,1243,1411]" pageId="21" pageNumber="22">
<bibRefCitation id="EFFE9BBBFFFFFFE0C0EFE186FE06FB35" box="[189,412,1243,1270]" pageId="21" pageNumber="22" refId="ref9534" refString="Zallot E., Groenenberg D. S. J., De Mattia W., Feher Z. &amp; Gittenberger E. 2015. Genera, subgenera and species of the Cochlostomatidae (Gastropoda, Caenogastropoda, Cochlostomatidae). Basteria 78 (4 - 6): 63 - 88." type="journal article">
Zallot
<emphasis id="B91B3A58FFFFFFE0C159E181FEDCFB36" box="[267,326,1243,1269]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2015)
</bibRefCitation>
assigned this species to
<taxonomicName id="4C6F9DC9FFFFFFE0C2ECE186FC7BFB35" authority="(Lovcenia)" authorityName="Lovcenia" box="[702,993,1243,1270]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B91B3A58FFFFFFE0C2ECE186FC7BFB35" box="[702,993,1243,1270]" italics="true" pageId="21" pageNumber="22">Cochlostoma (Lovcenia)</emphasis>
</taxonomicName>
, but with only one male specimen available, its systematic status remained difficult to judge. The shell morphology, in particular the protoconch sculpture, showed affinity to that of
<taxonomicName id="4C6F9DC9FFFFFFE0C347E07FFCE3FAFF" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1906" box="[789,889,1314,1340]" class="Gastropoda" family="Diplommatinidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="erika">
<emphasis id="B91B3A58FFFFFFE0C347E07FFCE3FAFF" box="[789,889,1314,1340]" italics="true" pageId="21" pageNumber="22">C. erika</emphasis>
</taxonomicName>
, whereas the molecular data suggested a status as a separate species. Since then, we have been able to collect and dissect female specimens. The structure of the female genitalia supports regarding it as a species on its own.
</paragraph>
</subSubSection>
</treatment>
</document>
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