treatments-xml/data/03/C6/00/03C600089624A548CE85FE590E29F8BA.xml
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<document ID-DOI="10.11646/zootaxa.5264.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836989" ID-ZooBank="9C61EF9D-1085-4EC6-B8A1-DCA65F768D76" approvalRequired="5" approvalRequired_for_textStreams="4" approvalRequired_for_treatments="1" checkinTime="1681730074228" checkinUser="plazi" docAuthor="Kostova, Rumyana &amp; Bekchiev, Rostislav" docDate="2023" docId="03C600089624A548CE85FE590E29F8BA" docLanguage="en" docName="zootaxa.5264.4.3.pdf" docOrigin="Zootaxa 5264 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus cardioderus subsp. irenis Csiki 1912" docType="treatment" docVersion="2" lastPageNumber="511" masterDocId="FFFF78709622A54ECE12FF830B3FFFBF" masterDocTitle="Trechus (s. str.) “ subnotatus ” species group (Coleoptera: Carabidae) in Bulgaria" masterLastPageNumber="525" masterPageNumber="505" pageNumber="511" updateTime="1681731023114" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Trechus (s. str.) “ subnotatus ” species group (Coleoptera: Carabidae) in Bulgaria</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kostova, Rumyana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bekchiev, Rostislav</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>505</mods:start>
<mods:end>525</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.4.3</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.4.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836989</mods:identifier>
<mods:identifier type="ZooBank">9C61EF9D-1085-4EC6-B8A1-DCA65F768D76</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA" httpUri="http://treatment.plazi.org/id/03C600089624A548CE85FE590E29F8BA" lastPageNumber="511" pageId="6" pageNumber="511">
<subSubSection box="[151,832,474,501]" pageId="6" pageNumber="511" type="nomenclature">
<paragraph blockId="6.[151,832,474,501]" box="[151,832,474,501]" pageId="6" pageNumber="511">
<heading bold="true" box="[151,832,474,501]" fontSize="11" level="1" pageId="6" pageNumber="511" reason="1">
<emphasis bold="true" box="[151,832,474,501]" pageId="6" pageNumber="511">
4.
<taxonomicName ID-CoL="57ZNZ" authority="Csiki, 1912" authorityName="Csiki" authorityYear="1912" box="[181,652,474,501]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="irenis">
<emphasis bold="true" box="[181,505,474,501]" italics="true" pageId="6" pageNumber="511">Trechus cardioderus irenis</emphasis>
<bibRefCitation author="Csiki, E." box="[513,652,474,500]" pageId="6" pageNumber="511" pagination="18 - 20" refId="ref12487" refString="Csiki, E. (1912) Uj barlangi carabidak. Rovartani Lapok, 19 (2 - 3), 18 - 20." type="journal article" year="1912">Csiki, 1912</bibRefCitation>
</taxonomicName>
(status novus)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="511" type="materials_examined">
<paragraph blockId="6.[151,1437,546,1797]" box="[151,1286,546,575]" pageId="6" pageNumber="511">
<emphasis bold="true" box="[151,327,546,573]" pageId="6" pageNumber="511">Type material:</emphasis>
<materialsCitation box="[334,1286,546,575]" collectionCode="MNHN" collectorName="Mihok &amp; Jeannel" country="Romania" location="Romania" pageId="6" pageNumber="511" specimenCount="1" typeStatus="syntype">
<emphasis bold="true" box="[334,430,546,572]" pageId="6" pageNumber="511">
<typeStatus box="[334,430,546,572]" pageId="6" pageNumber="511">Syntype</typeStatus>
</emphasis>
:
<emphasis bold="true" box="[445,554,546,572]" pageId="6" pageNumber="511">
<collectingCountry box="[445,554,546,572]" name="Romania" pageId="6" pageNumber="511">Romania</collectingCountry>
</emphasis>
: 1³, Antr. Godinest, Hunyad,
<collectorName box="[898,973,547,573]" pageId="6" pageNumber="511">Mihôk</collectorName>
leg. (coll.
<collectorName box="[1093,1180,547,573]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[1191,1278,547,573]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph blockId="6.[151,1437,546,1797]" pageId="6" pageNumber="511">
<emphasis bold="true" box="[199,505,582,608]" pageId="6" pageNumber="511">Other material examined:</emphasis>
<materialsCitation collectingDate="1996-07-25" collectingDateMax="2020-07-08" collectingDateMin="1996-07-25" collectionCode="NMNHS, R, BFUS" collectorName="B. Gueorguiev &amp; P. Mitov &amp; P. Stoev &amp; B. Petrov &amp; Bekchiev &amp; Kostova" country="Bulgaria" county="Rhodope Mts" latitude="41.51438" location="Stara" longLatPrecision="1" longitude="25.16167" municipality="Rila Mts" pageId="6" pageNumber="511" specimenCount="1">
<emphasis bold="true" box="[509,838,582,609]" pageId="6" pageNumber="511">
<collectingCountry box="[509,618,582,608]" name="Bulgaria" pageId="6" pageNumber="511">Bulgaria</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CC61FDC5098BFDDF" box="[627,692,582,608]" country="Bulgaria" county="Rhodope Mts" latitude="41.51438" longLatPrecision="1" longitude="25.16167" municipality="Rila Mts" name="Stara" pageId="6" pageNumber="511">Stara</location>
planina
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CD05FDC40879FDDE" box="[791,838,583,609]" country="Bulgaria" county="Rhodope Mts" latitude="41.51438" longLatPrecision="1" longitude="25.16167" municipality="Rila Mts" name="Mts" pageId="6" pageNumber="511">Mts</location>
</emphasis>
: 1³,west of
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CDC6FDC40F06FDDE" box="[980,1081,583,609]" country="Bulgaria" county="Rhodope Mts" latitude="41.51438" longLatPrecision="1" longitude="25.16167" municipality="Rila Mts" name="Krachela" pageId="6" pageNumber="511">Krachela</location>
peak,
<geoCoordinate box="[1150,1289,583,609]" degrees="42.740061" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.740063">N42.740061</geoCoordinate>
<geoCoordinate box="[1293,1432,582,609]" degrees="25.036336" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="25.036337">E25.036336</geoCoordinate>
,
<date box="[151,280,618,645]" pageId="6" pageNumber="511" value="1996-07-25">
<collectingDate box="[151,280,618,645]" pageId="6" pageNumber="511" value="1996-07-25">25.07.1996</collectingDate>
</date>
,
<collectorName box="[295,464,619,645]" pageId="6" pageNumber="511">B. Guéorguiev</collectorName>
leg. (
<collectionCode box="[536,644,619,645]" pageId="6" pageNumber="511">NMNHS</collectionCode>
); 1³, below Tazha chalet,
<geoCoordinate box="[962,1102,618,645]" degrees="42.751484" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.751484">N42.751484</geoCoordinate>
<geoCoordinate box="[1113,1251,618,645]" degrees="25.001135" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="25.001135">E25.001135</geoCoordinate>
,
<date box="[1266,1395,619,645]" pageId="6" pageNumber="511" value="1996-07-27">
<collectingDate box="[1266,1395,619,645]" pageId="6" pageNumber="511" value="1996-07-27">27.07.1996</collectingDate>
</date>
,
<collectorName pageId="6" pageNumber="511">B. Guéorguiev</collectorName>
leg. (
<collectionCode box="[352,460,655,681]" pageId="6" pageNumber="511">NMNHS</collectionCode>
); 1³, Toplya Vill, “Golyama Zhelezna” cave, vault of cave,
<date box="[1164,1292,655,681]" pageId="6" pageNumber="511" value="1997-09-28">
<collectingDate box="[1164,1292,655,681]" pageId="6" pageNumber="511" value="1997-09-28">28.09.1997</collectingDate>
</date>
,
<collectorName box="[1307,1404,655,681]" pageId="6" pageNumber="511">P. Mitov</collectorName>
&amp;
<collectorName box="[151,318,691,717]" pageId="6" pageNumber="511">B. Guéorguiev</collectorName>
leg. (
<collectionCode box="[386,494,691,717]" pageId="6" pageNumber="511">NMNHS</collectionCode>
); 1³ Rusalka hut,
<geoCoordinate box="[712,851,691,717]" degrees="42.689396" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.689396">N42.689396</geoCoordinate>
<geoCoordinate box="[861,1000,690,717]" degrees="25.056614" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="25.056614">E25.056614</geoCoordinate>
,
<date box="[1013,1142,691,717]" pageId="6" pageNumber="511" value="1997-12-10">
<collectingDate box="[1013,1142,691,717]" pageId="6" pageNumber="511" value="1997-12-10">12.10.1997</collectingDate>
</date>
,
<collectorName box="[1156,1249,691,717]" pageId="6" pageNumber="511">P. Stoev</collectorName>
leg. (
<collectionCode box="[1316,1424,691,717]" pageId="6" pageNumber="511">NMNHS</collectionCode>
); 3³³, Divchovoto Vill.,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CFBEFD540942FD4E" box="[428,637,727,753]" country="Bulgaria" county="Rhodope Mts" latitude="41.51438" longLatPrecision="1" longitude="25.16167" municipality="Rila Mts" name="Gradeshnitsa cave" pageId="6" pageNumber="511">Gradeshnitsa cave</location>
,
<geoCoordinate box="[650,790,727,753]" degrees="42.982760" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.98276">N42.982760</geoCoordinate>
<geoCoordinate box="[798,937,726,753]" degrees="24.263574" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="24.263575">E24.263574</geoCoordinate>
,
<date box="[950,1079,727,753]" pageId="6" pageNumber="511" value="1999-03-07">
<collectingDate box="[950,1079,727,753]" pageId="6" pageNumber="511" value="1999-03-07">03.07.1999</collectingDate>
</date>
,
<collectorName box="[1092,1199,727,753]" pageId="6" pageNumber="511">B. Petrov</collectorName>
leg. (
<collectionCode box="[1266,1374,727,753]" pageId="6" pageNumber="511">NMNHS</collectionCode>
); 1³, Vasilyovo Vill.,
<geoCoordinate box="[338,463,763,789]" degrees="42.88206" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.88206">N42.88206</geoCoordinate>
<geoCoordinate box="[473,597,763,789]" degrees="24.37411" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="24.37411">E24.37411</geoCoordinate>
,
<date box="[611,739,763,789]" pageId="6" pageNumber="511" value="2012-07-27">
<collectingDate box="[611,739,763,789]" pageId="6" pageNumber="511" value="2012-07-27">27.07.2012</collectingDate>
</date>
,
<collectionCode box="[754,776,763,789]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[789,895,763,789]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[964,1040,763,789]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
); 1³, Vasilyovo Vill, “Kozev dol” place,
<geoCoordinate box="[223,347,798,825]" degrees="41.51438" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="41.51438">N41.51438</geoCoordinate>
<geoCoordinate box="[353,478,798,825]" degrees="25.16167" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="25.16167">E25.16167</geoCoordinate>
,
<date box="[487,616,798,825]" pageId="6" pageNumber="511" value="2015-07-05">
<collectingDate box="[487,616,798,825]" pageId="6" pageNumber="511" value="2015-07-05">07.05.2015</collectingDate>
</date>
,
<collectionCode box="[626,648,799,825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[657,763,799,825]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[822,898,799,825]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
); 4³³, Chiflik Vill.,
<geoCoordinate box="[1129,1254,798,825]" degrees="42.82665" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.82665">N42.82665</geoCoordinate>
<geoCoordinate box="[1260,1385,798,825]" degrees="24.54894" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="24.54894">E24.54894</geoCoordinate>
, 24
<date box="[151,280,835,861]" pageId="6" pageNumber="511" value="2017-04-27">
<collectingDate box="[151,280,835,861]" pageId="6" pageNumber="511" value="2017-04-27">27.04.2017</collectingDate>
</date>
,
<collectionCode box="[290,312,835,861]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[322,428,835,861]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[490,566,835,861]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
); 1³, Pravets, path to “Kamicheto” place,
<geoCoordinate box="[1037,1161,835,861]" degrees="42.87944" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.87944">N42.87944</geoCoordinate>
<geoCoordinate box="[1168,1293,835,861]" degrees="23.93628" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="23.93628">E23.93628</geoCoordinate>
,
<date box="[1303,1432,835,861]" pageId="6" pageNumber="511" value="2020-04-08">
<collectingDate box="[1303,1432,835,861]" pageId="6" pageNumber="511" value="2020-04-08">04.08.2020</collectingDate>
</date>
,
<collectionCode box="[151,173,871,897]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[186,292,871,897]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[360,436,871,897]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
); 1³, Kalugerovo Vill., Chekotinski Monastery,
<geoCoordinate box="[995,1120,871,897]" degrees="42.98888" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.98888">N42.98888</geoCoordinate>
<geoCoordinate box="[1130,1255,871,897]" degrees="23.87380" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="23.8738">E23.87380</geoCoordinate>
,
<date box="[1268,1397,871,897]" pageId="6" pageNumber="511" value="2020-07-08">
<collectingDate box="[1268,1397,871,897]" pageId="6" pageNumber="511" value="2020-07-08">07.08.2020</collectingDate>
</date>
,
<collectionCode box="[1411,1433,871,897]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[151,244,907,933]" pageId="6" pageNumber="511">Kostova</collectorName>
leg. (
<collectionCode box="[306,382,907,933]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
); 1³, Kalugerovo Vill., road to Chekotinski Monastery,
<geoCoordinate box="[1005,1130,907,933]" degrees="42.98888" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.98888">N42.98888</geoCoordinate>
<geoCoordinate box="[1136,1261,907,933]" degrees="23.87380" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="23.8738">E23.87380</geoCoordinate>
,
<date box="[1271,1400,907,933]" pageId="6" pageNumber="511" value="2020-07-08">
<collectingDate box="[1271,1400,907,933]" pageId="6" pageNumber="511" value="2020-07-08">07.08.2020</collectingDate>
</date>
,
<collectionCode box="[1411,1433,907,933]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[151,257,943,969]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[319,395,943,969]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
);
<emphasis bold="true" box="[413,612,942,969]" pageId="6" pageNumber="511">Sredna gora Mts</emphasis>
: 1³, Bratiya Vill.,
<date box="[824,952,943,969]" pageId="6" pageNumber="511" value="2018-12-08">
<collectingDate box="[824,952,943,969]" pageId="6" pageNumber="511" value="2018-12-08">12.08.2018</collectingDate>
</date>
,
<collectionCode box="[963,985,943,969]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[995,1101,943,969]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[1162,1238,943,969]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
);
<emphasis bold="true" box="[1257,1360,942,969]" pageId="6" pageNumber="511">
<collectingMunicipality box="[1257,1360,942,969]" pageId="6" pageNumber="511">Rila Mts</collectingMunicipality>
</emphasis>
: 3³³, Shipochane Vill.,
<geoCoordinate box="[353,477,979,1005]" degrees="42.36716" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="42.36716">N42.36716</geoCoordinate>
<geoCoordinate box="[486,611,979,1005]" degrees="23.62139" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="23.62139">E23.62139</geoCoordinate>
,
<date box="[622,751,979,1005]" pageId="6" pageNumber="511" value="2019-06-17">
<collectingDate box="[622,751,979,1005]" pageId="6" pageNumber="511" value="2019-06-17">17.06.2019</collectingDate>
</date>
,
<collectionCode box="[763,785,979,1005]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[797,903,979,1005]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[967,1043,979,1005]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
);
<emphasis bold="true" box="[1063,1234,979,1005]" pageId="6" pageNumber="511">
<collectingCounty box="[1063,1229,979,1005]" pageId="6" pageNumber="511">Rhodope Mts</collectingCounty>
:
</emphasis>
2 ³³, Temenuga Vill.,
<geoCoordinate box="[214,339,1014,1041]" degrees="41.51438" direction="north" orientation="latitude" pageId="6" pageNumber="511" precision="1" value="41.51438">N41.51438</geoCoordinate>
<geoCoordinate box="[346,471,1014,1041]" degrees="25.16167" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="1" value="25.16167">E25.16167</geoCoordinate>
,
<date box="[482,611,1015,1041]" pageId="6" pageNumber="511" value="2019-06-26">
<collectingDate box="[482,611,1015,1041]" pageId="6" pageNumber="511" value="2019-06-26">26.06.2019</collectingDate>
</date>
,
<collectionCode box="[622,644,1015,1041]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="511">R</collectionCode>
.
<collectorName box="[655,761,1015,1041]" pageId="6" pageNumber="511">Bekchiev</collectorName>
leg. (
<collectionCode box="[824,897,1015,1041]" country="Bulgaria" httpUri="http://grbio.org/cool/7wb6-ejfj" name="University of Sofia, Biology Faculty" pageId="6" pageNumber="511">BFUS</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph blockId="6.[151,1437,546,1797]" box="[199,1436,1050,1077]" pageId="6" pageNumber="511">
<emphasis bold="true" box="[199,718,1050,1076]" pageId="6" pageNumber="511">Additional comparative material examined:</emphasis>
<emphasis box="[724,1176,1051,1077]" italics="true" pageId="6" pageNumber="511">dissected aedeagi (on microscope slides)</emphasis>
:
<taxonomicName authorityName="Putzeys" authorityYear="1870" box="[1189,1349,1051,1077]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="cardioderus">
<emphasis box="[1189,1349,1051,1077]" italics="true" pageId="6" pageNumber="511">T. cardioderus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1355,1408,1051,1077]" pageId="6" pageNumber="511" sensu="stricto">s.str.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="6.[151,1437,546,1797]" pageId="6" pageNumber="511">
<materialsCitation box="[151,838,1086,1115]" collectionCode="MNHN" collectorName="Jeannel" country="Romania" location="Ponorici" pageId="6" pageNumber="511" specimenCount="1" stateProvince="Hunedoara">
<emphasis bold="true" box="[151,269,1086,1112]" pageId="6" pageNumber="511">
<collectingCountry box="[151,264,1086,1112]" name="Romania" pageId="6" pageNumber="511">Romania</collectingCountry>
:
</emphasis>
1³,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CF5BFBBC0A94FBE6" box="[329,427,1087,1113]" country="Romania" name="Ponorici" pageId="6" pageNumber="511" stateProvince="Hunedoara">Ponorici</location>
,
<collectingRegion box="[440,563,1087,1113]" country="Romania" name="Hunedoara" pageId="6" pageNumber="511">Hunedoarâ</collectingRegion>
(coll.
<collectorName box="[639,726,1087,1113]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[739,830,1087,1113]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<taxonomicName box="[850,1127,1087,1113]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="pilisensis">
<emphasis box="[850,1127,1087,1113]" italics="true" pageId="6" pageNumber="511">T. cardioderus pilisensis</emphasis>
</taxonomicName>
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Austria" location="Graz" pageId="6" pageNumber="511" specimenCount="1">
<emphasis bold="true" box="[1155,1255,1086,1112]" pageId="6" pageNumber="511">
<collectingCountry box="[1155,1250,1086,1112]" name="Austria" pageId="6" pageNumber="511">Austria</collectingCountry>
:
</emphasis>
1³,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CB31FBBC0E66FBE6" box="[1315,1369,1087,1113]" country="Austria" name="Graz" pageId="6" pageNumber="511">Graz</location>
(coll.
<collectorName box="[151,238,1123,1149]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[252,343,1123,1149]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[364,834,1123,1151]" collectionCode="MNHN" collectorName="Jeannel" country="Austria" location="Reichenburg" pageId="6" pageNumber="511" specimenCount="1">
1³,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CF8DFBE00911FBC2" box="[415,558,1123,1149]" country="Austria" name="Reichenburg" pageId="6" pageNumber="511">Reichenburg</location>
(coll.
<collectorName box="[635,722,1123,1149]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[735,826,1123,1149]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<emphasis box="[847,1048,1123,1149]" italics="true" pageId="6" pageNumber="511">pinned specimens</emphasis>
:
<taxonomicName authorityName="Putzeys" authorityYear="1870" box="[1065,1229,1123,1149]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="cardioderus">
<emphasis box="[1065,1229,1123,1149]" italics="true" pageId="6" pageNumber="511">T. cardioderus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1239,1292,1123,1149]" pageId="6" pageNumber="511" sensu="stricto">s.str.</taxonomicNameLabel>
<materialsCitation collectionCode="MNHN" collectorName="Abaliget &amp; Jeannel" country="Hungary" location="Montes Mecsek" pageId="6" pageNumber="511" specimenCount="1">
<emphasis bold="true" box="[1320,1436,1123,1149]" pageId="6" pageNumber="511">
<collectingCountry box="[1320,1432,1123,1149]" name="Hungary" pageId="6" pageNumber="511">Hungary</collectingCountry>
:
</emphasis>
4³³,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CECEFB040AAFFB1E" box="[220,400,1159,1185]" country="Hungary" name="Montes Mecsek" pageId="6" pageNumber="511">Montes Mecsek</location>
, grotte
<collectorName box="[479,578,1159,1185]" pageId="6" pageNumber="511">Abaliget</collectorName>
, Dr. Gebhardt leg. (coll.
<collectorName box="[851,938,1159,1185]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[948,1039,1159,1185]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Romania" location="Baile Herculane" pageId="6" pageNumber="511" specimenCount="2" specimenCount-female="2">
<emphasis bold="true" box="[1056,1175,1158,1184]" pageId="6" pageNumber="511">
<collectingCountry box="[1056,1169,1158,1184]" name="Romania" pageId="6" pageNumber="511">Romania</collectingCountry>
:
</emphasis>
<specimenCount box="[1180,1238,1159,1185]" pageId="6" pageNumber="511" type="female">2♀♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CAF3FB040EA7FB1E" box="[1249,1432,1159,1185]" country="Romania" name="Baile Herculane" pageId="6" pageNumber="511">Bǎile Herculane</location>
, Banat (coll.
<collectorName box="[296,383,1195,1221]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[397,488,1195,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation box="[510,1077,1195,1223]" collectionCode="MNHN" collectorName="Jeannel" country="Romania" location="Ponorici" pageId="6" pageNumber="511" specimenCount="1" stateProvince="Hunedoara">
1³,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CC20FB2809ABFB7A" box="[562,660,1195,1221]" country="Romania" name="Ponorici" pageId="6" pageNumber="511" stateProvince="Hunedoara">Ponorici</location>
,
<collectingRegion box="[675,798,1195,1221]" country="Romania" name="Hunedoara" pageId="6" pageNumber="511">Hunedoarâ</collectingRegion>
(coll.
<collectorName box="[876,963,1195,1221]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[978,1069,1195,1221]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="MNHN" collectorName="Jeannel" country="Romania" elevation="1500" location="Carpathes" municipality="Negoi" pageId="6" pageNumber="511" specimenCount="1" stateProvince="Hunedoara">
1³,
<collectingMunicipality box="[1143,1214,1195,1221]" pageId="6" pageNumber="511">Negoi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03C600089624A548CE85FE590E29F8BA:8EB0E7C59624A548CADFFB280E7FFB7A" box="[1229,1344,1195,1221]" country="Romania" municipality="Negoi" name="Carpathes" pageId="6" pageNumber="511" stateProvince="Hunedoara">Carpathes</location>
,
<quantity box="[1358,1436,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="511" unit="m" value="1500.0">
<elevation box="[1358,1436,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="511" unit="m" value="1500.0">1500m</elevation>
</quantity>
(coll.
<collectorName box="[217,304,1231,1257]" pageId="6" pageNumber="511">Jeannel</collectorName>
,
<collectionCode box="[314,401,1231,1257]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph blockId="6.[151,1437,546,1797]" pageId="6" pageNumber="511">
<emphasis bold="true" box="[199,275,1267,1293]" pageId="6" pageNumber="511">Notes:</emphasis>
Relatively small: the body length of males is
<quantity box="[797,936,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.7" metricValueMin="3.8" pageId="6" pageNumber="511" unit="mm" value="4.25" valueMax="4.7" valueMin="3.8">3.84.7 mm</quantity>
. The bodys dorsal surface is reddish-brown and unicoloured. Fully reduced wings. The legs, antennae, and palpi are lighter—yellowish brown. Median lobe of the aedeagus: in lateral view, it is strongly arched at the dorsal side and almost straight in the distal two-thirds at the ventral side. The apex is short and curved abruptly downward. In dorsal view, the apex is slightly narrowed distally, and the apical lamella has a straight tip (
<figureCitation box="[607,676,1411,1437]" captionStart="FIGURE 4" captionStartId="7.[151,250,1851,1875]" captionTargetBox="[277,1310,181,1827]" captionTargetId="figure-19@7.[277,1310,181,1827]" captionTargetPageId="7" captionText="FIGURE 4. Variability in the median lobe of the aedeagus of Trechus cardioderus irenis Csiki in lateral and dorsal view (a,b—Rila Mts., Shipochane Vill.; c,d—Rila Mts., Shipochane Vill.; e,f—Central Stara Planina Mts., below Tazha chalet; g,h— Central Stara Planina Mts., West of Krachela Peak; I,j—West Stara Planina Mts. Kalugerovo Vill., Chekotinski Monastery; k,l—Central Stara Planina Mts., Vasilyovo Vill.; m,n—Central Stara Planina Mts., Chiflik Vill.; o,p—Sredna gora Mts., Bratiya Vill.). Scale: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7837002" httpUri="https://zenodo.org/record/7837002/files/figure.png" pageId="6" pageNumber="511">Fig. 4</figureCitation>
). The copulatory piece is strongly curved upward at the apical part. The subspecies first was discribed as a separate species by
<bibRefCitation author="Csiki, E." box="[815,948,1447,1473]" pageId="6" pageNumber="511" pagination="18 - 20" refId="ref12487" refString="Csiki, E. (1912) Uj barlangi carabidak. Rovartani Lapok, 19 (2 - 3), 18 - 20." type="journal article" year="1912">Csiki, 1912</bibRefCitation>
, after in 1927, Jeannel synonymised it with
<emphasis box="[151,465,1483,1509]" italics="true" pageId="6" pageNumber="511">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1870" box="[151,461,1483,1509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="cardioderus">T. cardioderus cardioderus</taxonomicName>
.
</emphasis>
We designated it as a subspecies of
<taxonomicName authorityName="Putzeys" authorityYear="1870" box="[885,1048,1483,1509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="cardioderus">
<emphasis box="[885,1048,1483,1509]" italics="true" pageId="6" pageNumber="511">T. cardioderus</emphasis>
</taxonomicName>
after comparing the
<typeStatus box="[1288,1375,1483,1509]" pageId="6" pageNumber="511">syntype</typeStatus>
of
<taxonomicName authorityName="Csiki" authorityYear="1912" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="irenis">
<emphasis italics="true" pageId="6" pageNumber="511">T. irenis</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1870" box="[269,573,1519,1545]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="cardioderus">
<emphasis box="[269,573,1519,1545]" italics="true" pageId="6" pageNumber="511">T. cardioderus cardioderus</emphasis>
</taxonomicName>
in the collection of Jeannel (
<collectionCode box="[900,991,1519,1545]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
), as well as with the Bulgarian material of the subspecies. The male genitalia of the studied Bulgarian material are very similar in lateral view to those of the
<typeStatus box="[192,279,1591,1617]" pageId="6" pageNumber="511">syntype</typeStatus>
in Jeannels collection (
<collectionCode box="[546,635,1591,1617]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
) and slightly differ in the shape of the apex from those of
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1870" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="cardioderus">
<emphasis italics="true" pageId="6" pageNumber="511">T. cardioderus cardioderus</emphasis>
</taxonomicName>
examined at
<collectionCode box="[432,521,1627,1653]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
. In
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1870" box="[560,860,1627,1653]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="cardioderus" subSpecies="cardioderus">
<emphasis box="[560,860,1627,1653]" italics="true" pageId="6" pageNumber="511">T. cardioderus cardioderus</emphasis>
</taxonomicName>
specimens from the collection of Jeannel (
<collectionCode box="[1333,1424,1627,1653]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
), the apex in lateral view is curved more abruptly straight downward, quite similarly to his illustration (
<bibRefCitation author="Jeannel, R." pageId="6" pageNumber="511" refId="ref13152" refString="Jeannel, R. (1927) Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres. Deuxieme Livraison. Socieitei entomologique de France, Paris, 592 pp." type="book" year="1927">Jeannel, 1927: 455</bibRefCitation>
). In the
<taxonomicName authorityName="Csiki" authorityYear="1912" box="[284,373,1699,1724]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="irenis">
<emphasis box="[284,373,1699,1724]" italics="true" pageId="6" pageNumber="511">T. irenis</emphasis>
</taxonomicName>
<typeStatus box="[379,466,1699,1725]" pageId="6" pageNumber="511">syntype</typeStatus>
from the collection of Jeannel (
<collectionCode box="[815,906,1699,1725]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="511">MNHN</collectionCode>
), the apex in lateral view is also curved abruptly downward but much shorter and under a very obtuse angle, as in
<figureCitation box="[877,945,1735,1761]" captionStart="FIGURE 4" captionStartId="7.[151,250,1851,1875]" captionTargetBox="[277,1310,181,1827]" captionTargetId="figure-19@7.[277,1310,181,1827]" captionTargetPageId="7" captionText="FIGURE 4. Variability in the median lobe of the aedeagus of Trechus cardioderus irenis Csiki in lateral and dorsal view (a,b—Rila Mts., Shipochane Vill.; c,d—Rila Mts., Shipochane Vill.; e,f—Central Stara Planina Mts., below Tazha chalet; g,h— Central Stara Planina Mts., West of Krachela Peak; I,j—West Stara Planina Mts. Kalugerovo Vill., Chekotinski Monastery; k,l—Central Stara Planina Mts., Vasilyovo Vill.; m,n—Central Stara Planina Mts., Chiflik Vill.; o,p—Sredna gora Mts., Bratiya Vill.). Scale: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7837002" httpUri="https://zenodo.org/record/7837002/files/figure.png" pageId="6" pageNumber="511">Fig. 4</figureCitation>
. As the differences between both groups are clear but only in the median lobe apex, we assigned
<taxonomicName authorityName="Csiki" authorityYear="1912" box="[737,827,1771,1796]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="irenis">
<emphasis box="[737,827,1771,1796]" italics="true" pageId="6" pageNumber="511">T. irenis</emphasis>
</taxonomicName>
the status of subspecies of
<taxonomicName authorityName="Putzeys" authorityYear="1870" box="[1133,1295,1771,1797]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="cardioderus">
<emphasis box="[1133,1295,1771,1797]" italics="true" pageId="6" pageNumber="511">T. cardioderus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>