187 lines
22 KiB
XML
187 lines
22 KiB
XML
<document id="423B74A73C1A5696C00AC292D204F2CE" ID-CLB-Dataset="21869" ID-DOI="10.5852/ejt.2016.236" ID-GBIF-Dataset="2dfee902-a7fc-472b-bf1d-27029b3504f3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3847840" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590436812908" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2016" docId="C146B323FFB6FFE0FD589E4FFE00F3A3" docLanguage="en" docName="ejt-2016-236.pdf.imf" docOrigin="European Journal of Taxonomy 236" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ptilototheca Herbert 2016, gen. nov." docType="treatment" docUuid="51C2C9E7-9514-43DF-A09B-3E391D3B61DD" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="3D7FCB5BFFBEFFE9FFC0994BFF82F57F" masterDocTitle="New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="9" updateTime="1698841246942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A919343EC9F6D6D2D74AA75FC5B55132">New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)</mods:title>
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<mods:namePart id="609B4A64942D349F4D333D978318A267">Herbert, David G.</mods:namePart>
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<mods:title id="E81CF55D31D88741737F1DE117DBF411">European Journal of Taxonomy</mods:title>
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<mods:part id="C6052D0873AF6EC0035ADEB3FB1338BA">
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<mods:date id="CE56B7B546642ECD45476146A8E002F6">2016</mods:date>
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<mods:number id="93D60194AC73C9120371B4AB2C66BCC4">2016-10-12</mods:number>
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<mods:number id="7588F069E98AECE20E3F9E2BC2BA78D6">236</mods:number>
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<treatment id="C146B323FFB6FFE0FD589E4FFE00F3A3" ID-DOI="http://doi.org/10.5281/zenodo.3854748" ID-GBIF-Taxon="164282653" ID-Zenodo-Dep="3854748" LSID="urn:lsid:zoobank.org:act:51C2C9E7-9514-43DF-A09B-3E391D3B61DD" httpUri="http://treatment.plazi.org/id/C146B323FFB6FFE0FD589E4FFE00F3A3" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<subSubSection id="01F551BEFFB6FFE1FD589E4FFB4BF23D" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="49500235FFB6FFE1FD589E4FFC1EF260" blockId="8.[363,1225,1796,1858]" box="[664,924,1796,1823]" pageId="8" pageNumber="9">
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<heading id="1218B559FFB6FFE1FD589E4FFC1EF260" box="[664,924,1796,1823]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="8EEF79B6FFB6FFE1FD589E4FFCABF260" ID-CoL="7PF62" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[664,809,1796,1823]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="7B9BDE27FFB6FFE1FD589E4FFCABF260" bold="true" box="[664,809,1796,1823]" italics="true" pageId="8" pageNumber="9">Ptilototheca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB6FFE1FCF19E4EFC1EF260" box="[817,924,1797,1823]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="49500235FFB6FFE1FEAB9E6CFB4BF23D" blockId="8.[363,1225,1796,1858]" box="[363,1225,1831,1858]" pageId="8" pageNumber="9">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB6FFE1FF7D9E22FD61F2CD" pageId="8" pageNumber="9" type="type_taxon">
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<paragraph id="49500235FFB6FFE1FF7D9E22FEDAF2FC" blockId="8.[189,344,1897,1923]" box="[189,344,1897,1923]" pageId="8" pageNumber="9">
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<heading id="1218B559FFB6FFE1FF7D9E22FEDAF2FC" bold="true" box="[189,344,1897,1923]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
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<emphasis id="7B9BDE27FFB6FFE1FF7D9E22FEDAF2FC" bold="true" box="[189,344,1897,1923]" pageId="8" pageNumber="9">
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<typeStatus id="9654BC97FFB6FFE1FF7D9E22FF78F2FC" box="[189,250,1897,1923]" pageId="8" pageNumber="9">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="49500235FFB6FFE1FF7D9EDCFD61F2CD" blockId="8.[189,739,1943,1970]" box="[189,739,1943,1970]" pageId="8" pageNumber="9">
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<taxonomicName id="8EEF79B6FFB6FFE1FF7D9EDCFDAEF2CE" ID-CoL="4Q8K5" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[189,556,1943,1969]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="soutpansbergensis" status="gen. et sp. nov.">
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<emphasis id="7B9BDE27FFB6FFE1FF7D9EDCFDAEF2CE" box="[189,556,1943,1969]" italics="true" pageId="8" pageNumber="9">Ptilototheca soutpansbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB6FFE1FDF39ED3FD61F2CD" box="[563,739,1944,1970]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB7FFE0FF7D9840FE09F4C2" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="49500235FFB7FFE0FF7D9840FEB4F45A" blockId="9.[189,310,267,293]" box="[189,310,267,293]" pageId="9" pageNumber="10">
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<heading id="1218B559FFB7FFE0FF7D9840FEB4F45A" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="7B9BDE27FFB7FFE0FF7D9840FEB4F45A" bold="true" box="[189,310,267,293]" pageId="9" pageNumber="10">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="49500235FFB7FFE0FF7D9872FE09F4C2" blockId="9.[189,1399,313,445]" pageId="9" pageNumber="10">Shell small, delicate and glossy, periphery below mid-whorl, base narrowly umbilicate; protoconch with close-set spiral lines of small punctations. Epiphallus very short, flagellum squat, vas deferens long, thickened and convoluted, penial verge present; spermatophore tail with well-developed, finely divided, branching spines.</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB7FFE0FF7D98AEFC2CF72F" pageId="9" pageNumber="10" type="etymology">
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<paragraph id="49500235FFB7FFE0FF7D98AEFEC6F480" blockId="9.[189,324,485,511]" box="[189,324,485,511]" pageId="9" pageNumber="10">
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<heading id="1218B559FFB7FFE0FF7D98AEFEC6F480" bold="true" box="[189,324,485,511]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="7B9BDE27FFB7FFE0FF7D98AEFEC6F480" bold="true" box="[189,324,485,511]" pageId="9" pageNumber="10">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="49500235FFB7FFE0FF7D9B5FFC2CF72F" blockId="9.[189,1399,531,592]" pageId="9" pageNumber="10">
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From
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<emphasis id="7B9BDE27FFB7FFE0FEC79B58FEC9F752" box="[263,331,531,557]" italics="true" pageId="9" pageNumber="10">ptilon</emphasis>
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,
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<emphasis id="7B9BDE27FFB7FFE0FE9C9B58FE36F752" box="[348,436,531,557]" italics="true" pageId="9" pageNumber="10">ptilotos</emphasis>
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(Gr.) a ‘feather, feathered’ and
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<emphasis id="7B9BDE27FFB7FFE0FCF69B58FCF4F752" box="[822,886,531,557]" italics="true" pageId="9" pageNumber="10">theca</emphasis>
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(L.) a ‘case or container’; referring to the plumose ornamentation of the spermatophore. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB7FFE0FF7D9B33FE00F3A3" pageId="9" pageNumber="10" type="discussion">
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<paragraph id="49500235FFB7FFE0FF7D9B33FEADF7ED" blockId="9.[189,303,632,658]" box="[189,303,632,658]" pageId="9" pageNumber="10">
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<heading id="1218B559FFB7FFE0FF7D9B33FEADF7ED" bold="true" box="[189,303,632,658]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="7B9BDE27FFB7FFE0FF7D9B33FEADF7ED" bold="true" box="[189,303,632,658]" pageId="9" pageNumber="10">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="49500235FFB7FFE0FF7D9BEDFE25F022" blockId="9.[189,1399,678,1373]" pageId="9" pageNumber="10">
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Within a southern African context, the spirally punctate protoconch of
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<taxonomicName id="8EEF79B6FFB7FFE0FBDE9BEDFB2FF7BF" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[1054,1197,678,704]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="7B9BDE27FFB7FFE0FBDE9BEDFB2FF7BF" box="[1054,1197,678,704]" italics="true" pageId="9" pageNumber="10">Ptilototheca</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FB799BECFAA5F7BE" box="[1209,1319,679,705]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
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might suggest an affinity with
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<taxonomicName id="8EEF79B6FFB7FFE0FE129B82FDC5F79C" authorityName="Ancey" authorityYear="1887" box="[466,583,713,739]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="7B9BDE27FFB7FFE0FE129B82FDC5F79C" box="[466,583,713,739]" italics="true" pageId="9" pageNumber="10">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FD8D9B81FDF1F79B" box="[589,627,714,740]" pageId="9" pageNumber="10" sensu="stricto">
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<emphasis id="7B9BDE27FFB7FFE0FD8D9B81FDEEF79C" box="[589,620,714,739]" italics="true" pageId="9" pageNumber="10">s.s</emphasis>
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.
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</taxonomicNameLabel>
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, but the distal genitalia of the present species are quite distinct and clearly discount any close relationship between the two. Although few other southern African heliciform urocyclids possess such a protoconch, the character is commonly present in semi-slug genera from further north in Africa (
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<bibRefCitation id="2D7E7FC4FFB7FFE0FE1E9A78FD11F632" author="Van Mol J. J." box="[478,659,819,845]" pageId="9" pageNumber="10" refId="ref14648" refString="Van Mol J. J. 1970. Revision des Urocyclidae (Mollusca, Gastropoda, Pulmonata) - anatomie - systematique - zoogeographie. Premiere partie. Annales du Musee Royal de l'Afrique Centrale, Sciences zoologiques 180, Royal Museum for Central Africa, Tervuren." type="book" year="1970">Van Mol 1970</bibRefCitation>
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). In terms of its distal genital anatomy,
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<taxonomicName id="8EEF79B6FFB7FFE0FBBE9A7FFAF5F632" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[1150,1399,819,845]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="soutpansbergensis" status="gen. et sp. nov.">
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<emphasis id="7B9BDE27FFB7FFE0FBBE9A7FFAF5F632" box="[1150,1399,819,845]" italics="true" pageId="9" pageNumber="10">P. soutpansbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FF7D9A1CFEF2F60E" box="[189,368,855,881]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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is unlike any other southern African heliciform urocyclid species. The extremely short epiphallus, short squat flagellum, ornately bristled spermatophore and long, thick, convoluted vas deferens are distinctive characters not seen in other species from the region. In addition, the presence of a penial verge and a papillate preputial region are also unusual features among local species, but these are also present in
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<taxonomicName id="8EEF79B6FFB7FFE0FE239AA8FD2AF682" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[483,680,995,1021]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="wolkbergensis" status="sp. nov.">
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<emphasis id="7B9BDE27FFB7FFE0FE239AA8FD2AF682" box="[483,680,995,1021]" italics="true" pageId="9" pageNumber="10">S. wolkbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FD719AAFFC8FF681" box="[689,781,996,1022]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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(see below). That species, however, lacks a spirally punctate protoconch, possesses an atrial diverticulum, has a short, slender, non-convoluted vas deferens, and an epiphallus and flagellum of more conventional length for
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<taxonomicName id="8EEF79B6FFB7FFE0FC0A9D62FBBDF13C" authorityName="Ancey" authorityYear="1887" box="[970,1087,1065,1091]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus" sensu="lato">
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<emphasis id="7B9BDE27FFB7FFE0FC0A9D62FBBDF13C" box="[970,1087,1065,1091]" italics="true" pageId="9" pageNumber="10">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FB899D62FBE9F13B" box="[1097,1131,1065,1092]" pageId="9" pageNumber="10" sensu="lato">
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<emphasis id="7B9BDE27FFB7FFE0FB899D62FBE6F13C" box="[1097,1124,1065,1091]" italics="true" pageId="9" pageNumber="10">s.l</emphasis>
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.
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</taxonomicNameLabel>
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Species of the central African semi-slug genus
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<taxonomicName id="8EEF79B6FFB7FFE0FE209D06FC9EF118" authority="Pilsbry, 1919" authorityName="Pilsbry" authorityYear="1919" box="[480,796,1100,1127]" class="Gastropoda" family="Urocyclidae" genus="Angustivestis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB7FFE0FE209D06FDF9F118" box="[480,635,1101,1127]" italics="true" pageId="9" pageNumber="10">Angustivestis</emphasis>
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Pilsbry, 1919
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</taxonomicName>
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also possess a penial verge and a papillate preputial region, as well as a long, thick, convoluted vas deferens, but again in that genus the protoconch is not spirally punctate, the epiphallus and flagellum not conspicuously shortened and there is usually an atrial diverticulum (
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<bibRefCitation id="2D7E7FC4FFB7FFE0FE659DFDFDD1F1AF" author="Van Mol J. J." box="[421,595,1206,1232]" pageId="9" pageNumber="10" refId="ref14648" refString="Van Mol J. J. 1970. Revision des Urocyclidae (Mollusca, Gastropoda, Pulmonata) - anatomie - systematique - zoogeographie. Premiere partie. Annales du Musee Royal de l'Afrique Centrale, Sciences zoologiques 180, Royal Museum for Central Africa, Tervuren." type="book" year="1970">Van Mol 1970</bibRefCitation>
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). Considered collectively, the conchological and anatomical features exhibited by
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<taxonomicName id="8EEF79B6FFB7FFE0FE979D91FDCEF18C" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[343,588,1241,1267]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="soutpansbergensis" status="gen. et sp. nov.">
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<emphasis id="7B9BDE27FFB7FFE0FE979D91FDCEF18C" box="[343,588,1241,1267]" italics="true" pageId="9" pageNumber="10">P. soutpansbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB7FFE0FD949D91FC87F18B" box="[596,773,1242,1268]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
strongly suggest that it is not closely related to other southern African heliciform urocyclids and that it belongs to a distinct lineage that may perhaps have closer affinities with central African genera. Its distribution in the extreme north of
|
||
<collectingCountry id="31F842A5FFB7FFE0FB7F9C6BFADAF045" box="[1215,1368,1312,1338]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
|
||
is consistent with this.
|
||
</paragraph>
|
||
<paragraph id="49500235FFB7FFE0FF7D9CCFFE00F3A3" blockId="9.[189,1399,1412,1756]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="8EEF79B6FFB7FFE0FF7D9CCFFD68F0E0" authority="Melvill & Ponsonby, 1909" authorityName="Melvill & Ponsonby" authorityYear="1909" box="[189,746,1412,1439]" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="pumilio">
|
||
<emphasis id="7B9BDE27FFB7FFE0FF7D9CCFFE22F0E1" box="[189,416,1412,1438]" italics="true" pageId="9" pageNumber="10">Helicarion pumilio</emphasis>
|
||
Melvill & Ponsonby, 1909
|
||
</taxonomicName>
|
||
, the type species of
|
||
<taxonomicName id="8EEF79B6FFB7FFE0FC249CCFFAF3F0E0" authority="Godwin-Austen, 1912" authorityName="Godwin-Austen" authorityYear="1912" box="[996,1393,1412,1439]" class="Gastropoda" family="Urocyclidae" genus="Andrarion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB7FFE0FC249CCFFBDCF0E1" box="[996,1118,1412,1438]" italics="true" pageId="9" pageNumber="10">Andrarion</emphasis>
|
||
<bibRefCitation id="2D7E7FC4FFB7FFE0FBA89CCFFAF3F0E0" author="Godwin-Austen H. H." box="[1128,1393,1412,1439]" pageId="9" pageNumber="10" pagination="122 - 139" refId="ref13983" refString="Godwin-Austen H. H. 1912. A review of the South-African land-Mollusca belonging to the family Zonitidae [Part I]. Annals and Magazine of Natural History Series 8, 9: 122 - 139. Available from http: // biodiversitylibrary. org / page / 18638879 [accessed 10 Aug. 2016]" type="journal article" year="1912">Godwin-Austen, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
, was also described from the Soutpansberg. This remains a poorly known and enigmatic species, but it differs from the present material in having a smaller, more depressed shell with a smooth protoconch and a rapidly expanding last whorl [confirmed by examination of the type material:
|
||
<typeStatus id="9654BC97FFB7FFE0FB079CA5FAB1F377" box="[1223,1331,1518,1544]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
|
||
(des.
|
||
<bibRefCitation id="2D7E7FC4FFB7FFE0FF7D9F5AFE2BF354" author="Connolly M." box="[189,425,1553,1579]" pageId="9" pageNumber="10" pagination="59 - 306" refId="ref13749" refString="Connolly M. 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306." type="journal article" year="1912">Connolly 1912: 101</bibRefCitation>
|
||
) NHMUK 1910.1.5.12 (maj. diameter
|
||
<quantity id="8E17AFD0FFB7FFE0FCB79F5AFC51F353" box="[887,979,1553,1580]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="10" unit="mm" value="6.8">6.8 mm</quantity>
|
||
) and three
|
||
<typeStatus id="9654BC97FFB7FFE0FB979F5AFA82F354" box="[1111,1280,1553,1579]" pageId="9" pageNumber="10" type="paralectotype">paralectotypes</typeStatus>
|
||
NHMUK 1937.12.30.4113–5]. The original specimens remain the only material available and I strongly suspect that they are simply juveniles of a larger species of
|
||
<taxonomicName id="8EEF79B6FFB7FFE0FCC19F13FBABF30D" authority="" box="[769,1065,1624,1650]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subGenus" subGenus="Kerkophorus">
|
||
<emphasis id="7B9BDE27FFB7FFE0FCC19F13FCF4F30D" box="[769,886,1624,1650]" italics="true" pageId="9" pageNumber="10">Sheldonia</emphasis>
|
||
(
|
||
<emphasis id="7B9BDE27FFB7FFE0FC459F13FB9CF30D" box="[901,1054,1624,1650]" italics="true" pageId="9" pageNumber="10">Kerkophorus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, at least one species of which occurs in the Afrotemperate forests of
|
||
<collectingRegion id="8B2BCCD7FFB7FFE0FD469F30FD77F3EA" box="[646,757,1659,1685]" country="South Africa" name="Limpopo" pageId="9" pageNumber="10">Limpopo</collectingRegion>
|
||
. This remains a matter requiring further investigation, but for the present it is sufficient to confirm that the material discussed herein is undoubtedly distinct from
|
||
<taxonomicName id="8EEF79B6FFB7FFE0FF3D9F89FEF9F3A4" authorityName="Melvill & Ponsonby" authorityYear="1909" box="[253,379,1729,1755]" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="pumilio">
|
||
<emphasis id="7B9BDE27FFB7FFE0FF3D9F89FEF9F3A4" box="[253,379,1729,1755]" italics="true" pageId="9" pageNumber="10">H. pumilio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |