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<document id="2C622BAC9F600515BFD9508F8B05B740" ID-CLB-Dataset="31368" ID-DOI="10.11646/zootaxa.4351.1.1" ID-GBIF-Dataset="c2a3a65e-90ff-4bba-af4d-91d82594f636" ID-ISSN="1175-5326" ID-Zenodo-Dep="1063746" ID-ZooBank="99BC6304-97CF-4463-B368-D2F102535DD9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511248295005" checkinUser="plazi" docAuthor="Wang, Lin-Fei, Zhou, Hong-Zhang &amp; Lü, Liang" docDate="2017" docId="B44687B22B78FFF099E2FE5FFDC1F874" docLanguage="en" docName="zootaxa.4351.1.1.pdf.imf" docOrigin="Zootaxa 4351 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anotylus opaciceps" docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="487FFFCA2B57FFC09975FFABFFDEFFA3" masterDocTitle="Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China" masterLastPageNumber="79" masterPageNumber="1" pageNumber="48" updateTime="1698502226059" updateUser="ExternalLinkService">
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<mods:title id="8347892568E344B2A90473C080DA6978">Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China</mods:title>
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<mods:namePart id="1392B1628EFB80880F60911DC6E8EFC4">Wang, Lin-Fei</mods:namePart>
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<mods:namePart id="9A5A7C78F0B3FECBB91817E21766A63B">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="B44687B22B78FFF099E2FE5FFDC1F874" ID-DOI="http://doi.org/10.5281/zenodo.6033735" ID-GBIF-Taxon="137467049" ID-Zenodo-Dep="6033735" LSID="urn:lsid:plazi:treatment:B44687B22B78FFF099E2FE5FFDC1F874" httpUri="http://treatment.plazi.org/id/B44687B22B78FFF099E2FE5FFDC1F874" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<subSubSection id="74F5652F2B78FFEF99E2FE5FFEDAFD93" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="3C5036A42B78FFEF99E2FE5FFDB9FDAC" blockId="47.[151,615,500,560]" box="[151,615,500,527]" pageId="47" pageNumber="48">
<heading id="671881C82B78FFEF99E2FE5FFDB9FDAC" bold="true" box="[151,615,500,527]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<taxonomicName id="FBEF4D272B78FFEF99E2FE5FFDB9FDAC" ID-CoL="66YBP" authority="(Bernhauer, 1938)" baseAuthorityName="Bernhauer" baseAuthorityYear="1938" box="[151,615,500,527]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B78FFEF99E2FE5FFDB9FDAC" bold="true" box="[151,615,500,527]" pageId="47" pageNumber="48">
<emphasis id="0E9BEAB62B78FFEF99E2FE5FFEA1FDAD" bold="true" box="[151,383,500,526]" italics="true" pageId="47" pageNumber="48">Anotylus opaciceps</emphasis>
(
<bibRefCitation id="587E4B552B78FFEF98FBFE5EFD81FDAC" author="Bernhauer" box="[398,607,501,527]" pageId="47" pageNumber="48" refString="Bernhauer, M. (1938) Zur Staphylinidenfauna von China u. Japan. Entomologisches Nachrichtenblatt, Troppau, 12, 17 - 39." type="journal article" year="1938">Bernhauer, 1938</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3C5036A42B78FFEF99E2FDBCFEDAFD93" blockId="47.[151,615,500,560]" box="[151,260,535,560]" pageId="47" pageNumber="48">(Figs. 13)</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B78FFEF99E2FDCBFAA3FD10" pageId="47" pageNumber="48" type="description">
<paragraph id="3C5036A42B78FFEF99E2FDCBFAA3FD10" blockId="47.[151,1436,608,691]" pageId="47" pageNumber="48">
<bibRefCitation id="587E4B552B78FFEF99E2FDCBFE97FDD5" author="Bernhauer" box="[151,329,608,630]" pageId="47" pageNumber="48" refString="Bernhauer, M. (1938) Zur Staphylinidenfauna von China u. Japan. Entomologisches Nachrichtenblatt, Troppau, 12, 17 - 39." type="journal article" year="1938">Bernhauer, 1938</bibRefCitation>
: 24 (
<taxonomicName id="FBEF4D272B78FFEF98FEFDCBFE3AFDD5" authorityName="Gravenhorst" authorityYear="1802" box="[395,484,608,630]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF98FEFDCBFE3AFDD5" box="[395,484,608,630]" italics="true" pageId="47" pageNumber="48">Oxytelus</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="FBEF4D272B78FFEF9B11FDCBFD63FDD5" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[612,701,608,630]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9B11FDCBFD63FDD5" box="[612,701,608,630]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
; Type locality: Nordwestl. China: Chinkiang); Herman, 1970: 419 (
<taxonomicName id="FBEF4D272B78FFEF99BAFDD5FEF6FD37" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[207,296,638,660]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF99BAFDD5FEF6FD37" box="[207,296,638,660]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
);
<bibRefCitation id="587E4B552B78FFEF9834FDD4FE04FD37" author="Herman" box="[321,474,638,660]" pageId="47" pageNumber="48" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the second millennium. Vol. 3, Oxyteline group. Bulletin of the American Museum of Natural History, 265, 1067 - 1806." type="book chapter" year="2001">Herman, 2001</bibRefCitation>
: 1378 (
<taxonomicName id="FBEF4D272B78FFEF9B59FDD5FD5BFD37" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[556,645,638,660]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9B59FDD5FD5BFD37" box="[556,645,638,660]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
; catalog);
<bibRefCitation id="587E4B552B78FFEF9B8DFDD5FC48FD37" author="Smetana" box="[760,918,638,660]" pageId="47" pageNumber="48" refString="Smetana, A. (2004) Subfamily Oxytelinae Fleming, 1821. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera: Hydrophiloidea - Histeroidea - Staphylinoidea. Vol. 2. Apollo Books, Stenstrup, pp. 511 - 535." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 515 (
<taxonomicName id="FBEF4D272B78FFEF9AAEFDD5FBEAFD37" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[987,1076,638,660]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9AAEFDD5FBEAFD37" box="[987,1076,638,660]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
; Palaearctic catalog; distribution);
<bibRefCitation id="587E4B552B78FFEF99B2FD36FE58FD10" author="Zheng" box="[199,390,669,691]" pageId="47" pageNumber="48" refString="Zheng, F. - K. &amp; Pu, S. - H. (2004) Study of the genus Anotylus C. G. Thomson (Coleoptera: Staphylindae, Oxytelinae). Journal of China West Normal University (Natural Sciences), 25, 370 - 374." type="journal article" year="2004">Zheng &amp; Pu, 2004</bibRefCitation>
: 372 (
<taxonomicName id="FBEF4D272B78FFEF98B1FD36FDC3FD10" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[452,541,669,691]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF98B1FD36FDC3FD10" box="[452,541,669,691]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
; China);
<bibRefCitation id="587E4B552B78FFEF9B09FD36FC56FD10" author="Schulke" box="[636,904,669,691]" pageId="47" pageNumber="48" refString="Schulke, M. &amp; Smetana, A. (2015) Oxytelinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. 1 (Hydrophiloidea-Staphylinoidea). Leiden &amp; Boston, Brill, pp. 757 - 795." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
: 772 (
<taxonomicName id="FBEF4D272B78FFEF9AB3FD36FBC1FD10" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[966,1055,669,691]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9AB3FD36FBC1FD10" box="[966,1055,669,691]" italics="true" pageId="47" pageNumber="48">Anotylus</emphasis>
</taxonomicName>
; Palaearctic catalog; distribution).
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B78FFEF99E2FD74FE07FC73" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="3C5036A42B78FFEF99E2FD74FE07FC73" blockId="47.[151,1436,735,1048]" pageId="47" pageNumber="48">
<emphasis id="0E9BEAB62B78FFEF99E2FD74FDE3FD5B" bold="true" box="[151,573,735,760]" pageId="47" pageNumber="48">Type material examined. Lectotype</emphasis>
(designed here): female, Nordweatl. China Chinkiang [= Zhenjiang, Jiangsu Prov.] Col. Reitter//
<taxonomicName id="FBEF4D272B78FFEF980EFCAFFDE3FCBE" authority="Bernh." authorityName="Bernh." box="[379,573,772,797]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">opaciceps Bernh.</taxonomicName>
Typus//
<taxonomicName id="FBEF4D272B78FFEF9BE8FCAEFCBFFCBE" authority="Bernh." authorityName="Bernh." box="[669,865,772,797]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B78FFEF9BE8FCAEFCD3FCBF" box="[669,781,773,796]" italics="true" pageId="47" pageNumber="48">opaciceps</emphasis>
Bernh.
</taxonomicName>
Typus (
<taxonomicName id="FBEF4D272B78FFEF9AC9FCAFFBC5FCBF" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[956,1051,772,796]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9AC9FCAFFBC5FCBF" box="[956,1051,772,796]" italics="true" pageId="47" pageNumber="48">anotylus</emphasis>
</taxonomicName>
)// Chicago NHMus M. Bernhauer Collection//
<taxonomicName id="FBEF4D272B78FFEF9857FC83FDFEFCE3" authority="Bn" authorityName="Bn" box="[290,544,808,832]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B78FFEF9857FC83FE25FC9C" box="[290,507,808,832]" italics="true" pageId="47" pageNumber="48">Anotylus opaciceps</emphasis>
Bn
</taxonomicName>
// FMNHINS 0 0 0 0 131 002// Photographed Kelsey Keaton 2014 Emu Catalog// SYNTYPE teste D. J. Clarke2014 GDI Imaging Project. (FMNH);
<emphasis id="0E9BEAB62B78FFEF9AD0FCE7FB94FCC6" bold="true" box="[933,1098,844,869]" pageId="47" pageNumber="48">Paralectotype</emphasis>
: 1 female, Nordweatl. China Chinkiang [= Zhenjiang, Jiangsu Prov.] Col. Reitter//
<taxonomicName id="FBEF4D272B78FFEF9A76FCDBFBF1FC2B" authority="Bernh. Cotypus" authorityName="Bernh. Cotypus" box="[771,1071,879,904]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B78FFEF9A76FCDBFCADFC24" box="[771,883,880,903]" italics="true" pageId="47" pageNumber="48">opaciceps</emphasis>
Bernh. Cotypus
</taxonomicName>
<taxonomicName id="FBEF4D272B78FFEF9D4CFCDBFB46FC2B" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[1081,1176,880,904]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B78FFEF9D4CFCDBFB46FC2B" box="[1081,1176,880,904]" italics="true" pageId="47" pageNumber="48">anotylus</emphasis>
</taxonomicName>
// Chicago NHMus M. Bernhauer Collection// SYNTYPE teste D. J. Clarke2014 GDI Imaging Project// FMNHINS 2840707 FIELD MUSEUM Pinned. (FMNH)
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B78FFF099B2FC77FCF3F810" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="3C5036A42B78FFEF99B2FC77FBFFFBB4" blockId="47.[151,1436,735,1048]" pageId="47" pageNumber="48">
<emphasis id="0E9BEAB62B78FFEF99B2FC77FEAAFC56" bold="true" box="[199,372,988,1013]" pageId="47" pageNumber="48">Redescription.</emphasis>
Body dark brown; mouthparts and legs yellow. Forebody slightly pubescent, punctate and striate, with coriaceous and depressed intervals. Body length [average]
<quantity id="FB179B412B78FFEF9AC0FC54FBC2FBB4" box="[949,1052,1023,1048]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="47" pageNumber="48" unit="mm" value="4.25">4.25 mm</quantity>
.
</paragraph>
<caption id="6890662C2B78FFEF99E2F8C2FE8BF81E" httpUri="https://zenodo.org/record/1063808/files/figure.png" pageId="47" pageNumber="48" targetBox="[151,1410,1103,1840]" targetPageId="47">
<paragraph id="3C5036A42B78FFEF99E2F8C2FE8BF81E" blockId="47.[151,1436,1897,1981]" pageId="47" pageNumber="48">
<emphasis id="0E9BEAB62B78FFEF99E2F8C2FEE6F8DC" bold="true" box="[151,312,1897,1919]" pageId="47" pageNumber="48">FIGURE 13-1.</emphasis>
<taxonomicName id="FBEF4D272B78FFEF9836F8C2FD0CF8DC" authority="(Bernhauer, 1938)" authorityName="Bernhauer" authorityYear="1938" baseAuthorityName="Bernhauer" baseAuthorityYear="1938" box="[323,722,1897,1919]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B78FFEF9836F8C2FDD5F8DC" box="[323,523,1897,1919]" italics="true" pageId="47" pageNumber="48">Anotylus opaciceps</emphasis>
(Bernhauer, 1938)
</taxonomicName>
A, habitus female; B, head and pronotum female; C, specimen on mounting card with labels; D, sternite VII female; E, sternite VIII female; F, spermatheca. Scales: A, B = 0.5 mm; D, E= 0.2 mm, F = 0.05 mm.
</paragraph>
</caption>
<caption id="6890662C2B67FFF099E2FB71FCC5FAAC" httpUri="https://zenodo.org/record/1063810/files/figure.png" pageId="48" pageNumber="49" targetBox="[204,1389,244,1168]" targetPageId="48">
<paragraph id="3C5036A42B67FFF099E2FB71FCC5FAAC" blockId="48.[151,1435,1242,1295]" pageId="48" pageNumber="49">
<emphasis id="0E9BEAB62B67FFF099E2FB71FEE8FB53" bold="true" box="[151,310,1242,1264]" pageId="48" pageNumber="49">FIGURE 13-2.</emphasis>
<taxonomicName id="FBEF4D272B67FFF09848FB71FD1BFB53" authority="(Bernhauer, 1938)" authorityName="Bernhauer" authorityYear="1938" baseAuthorityName="Bernhauer" baseAuthorityYear="1938" box="[317,709,1242,1264]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis id="0E9BEAB62B67FFF09848FB71FDDDFB53" box="[317,515,1242,1264]" italics="true" pageId="48" pageNumber="49">Anotylus opaciceps</emphasis>
(Bernhauer, 1938)
</taxonomicName>
A, head and pronotum female; B, sternite VII female; C, sternite VIII female; D, spermatheca. Scales: A– C = 0.5 mm; D = 0.05 mm.
</paragraph>
</caption>
<paragraph id="3C5036A42B67FFF099B2FA9DFEF8F985" blockId="48.[151,1437,1334,2007]" pageId="48" pageNumber="49">
<emphasis id="0E9BEAB62B67FFF099B2FA9DFEC5FAED" box="[199,283,1334,1358]" italics="true" pageId="48" pageNumber="49">Female</emphasis>
: Head (Fig. 13-1B, 13-2A) slightly broader, but narrower than pronotum. Vertex extremely convex, with a pair of raising parts glabrous and curved, with deep sulcus running parallel in middle; all depressed parts coriaceous. Supra-antennal ridge with margin incrassate. Temples as long as eyes. Clypeus nearly square, coriaceous, about 1/2 of total head length, with anterior margin roundly produced. Epistomal suture with lateral portions straight and parallel, running backward to level of anterior margin of eyes. Occipital suture nearly straight. Antenna filiform, segment IV globular, V–X short and transverse, with ultimate article stilliform. Neck without smooth area.
</paragraph>
<paragraph id="3C5036A42B67FFF099B2F998FB67F8E4" blockId="48.[151,1437,1334,2007]" pageId="48" pageNumber="49">Pronotum (Fig. 13-1B, 13-2A) transverse, broadest at near anterior 1/4; disc slightly convex, mid-longitudinal sulcus broad and shallow, narrowing and deepening to base, with two shallow, straight paramedial sulci, and bounded by two obtuse glabrous longitudinal carinae carrying big punctures, and two shallow para-lateral depressions close to lateral margins. Anterior margin slightly and broadly bi-emarginate, with anterior angles slightly protruded. Lateral margin weakly crenate in posterior 1/4. Lateral and posterior margins running nearly continuously, posterior angles less distinguishable. Scutellum with crest-shaped impression with median lobe sharply pointed. Elytra with fine and moderately dense striae and punctures, not pubescent; lateral margin curved and posterior margins truncate. Abdomen coriaceous, covered with punctures and pubescence.</paragraph>
<paragraph id="3C5036A42B67FFF099B2F8F9FCF3F810" blockId="48.[151,1437,1334,2007]" pageId="48" pageNumber="49">Sternite VII (Fig. 13-1D, 13-2B) with posterior margin straight, and bearing long setae laterally. Sternite VIII (Fig. 13-1E, 13-2C) with posterior margin broadly rounded and protruded. Spermatheca (Fig. 13-1F, 13-2D) with base bulb-like, with apical part clavate and slightly curved.</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B67FFF099B2F815FDC1F874" box="[199,543,1982,2007]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="3C5036A42B67FFF099B2F815FDC1F874" blockId="48.[151,1437,1334,2007]" box="[199,543,1982,2007]" pageId="48" pageNumber="49">
<emphasis id="0E9BEAB62B67FFF099B2F815FEBEF874" bold="true" box="[199,352,1982,2007]" pageId="48" pageNumber="49">Distribution.</emphasis>
<collectingCountry id="44F876342B67FFF09812F815FE74F874" box="[359,426,1982,2007]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
(
<collectingRegion id="FE2BF8462B67FFF098CCF815FDCDF874" box="[441,531,1982,2007]" country="China" name="Jiangsu" pageId="48" pageNumber="49">Jiangsu</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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