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<mods:title id="41D8551E8447B59CB2257F105051911D">Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China</mods:title>
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<mods:namePart id="75EC4C550DEE6D98666479EF2469C1D1">Wang, Lin-Fei</mods:namePart>
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<mods:namePart id="F9BD73C84E7C36FE7636F0178140CE52">Zhou, Hong-Zhang</mods:namePart>
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<mods:namePart id="CFEFADEA951C3C5D0DFBC12A9A93F6C9">Lü, Liang</mods:namePart>
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<treatment id="B44687B22B51FFC699E2FEDBFA42F9DD" ID-DOI="http://doi.org/10.5281/zenodo.6033705" ID-GBIF-Taxon="137467048" ID-Zenodo-Dep="6033705" LSID="urn:lsid:plazi:treatment:B44687B22B51FFC699E2FEDBFA42F9DD" httpUri="http://treatment.plazi.org/id/B44687B22B51FFC699E2FEDBFA42F9DD" lastPageNumber="7" pageId="6" pageNumber="7">
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<heading id="671881C82B51FFC699E2FEDBFBCCFE29" bold="true" box="[151,1042,367,394]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="0E9BEAB62B51FFC699E2FEDBFBCCFE29" bold="true" box="[151,1042,367,394]" pageId="6" pageNumber="7">
Key to species groups of the genus
<taxonomicName id="FBEF4D272B51FFC69B3DFEC4FCA4FE29" ID-CoL="8VRGY" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[584,890,367,394]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B51FFC69B3DFEC4FD6AFE2A" bold="true" box="[584,692,367,393]" italics="true" pageId="6" pageNumber="7">Anotylus</emphasis>
<bibRefCitation id="587E4B552B51FFC69BCEFEDBFCA4FE29" author="Thomson" box="[699,890,368,394]" pageId="6" pageNumber="7" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. I. Berlingska Boktryckeriet, Lund, 290 pp." type="book" year="1859">Thomson, 1859</bibRefCitation>
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from China
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1. Anterior portion of the head with circular impression, pronotum with lateral portion expanded, aliform, tergites IX with ventral struts enlarged to a broad, rounded plate, almost as large as the main parts of these plates (
<figureCitation id="A4D42A212B51FFC69D34FE78FB4AFE4B" box="[1089,1172,467,488]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Fig. 2-O</figureCitation>
) (body size
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).......................................................................................
<taxonomicName id="FBEF4D272B51FFC69D82FE5AFA83FDA5" ID-CoL="4BKJ5" authorityName="Fauvel" authorityYear="1895" box="[1271,1373,497,518]" class="Insecta" family="Staphylinidae" genus="Oxytelopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="0E9BEAB62B51FFC69D82FE5AFA83FDA5" box="[1271,1373,497,518]" italics="true" pageId="6" pageNumber="7">cimicoides</emphasis>
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group
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<keyLead id="871E95912B51FFC699E2FDA5FA42FD80" box="[151,1436,526,547]" pageId="6" pageNumber="7">- Without the above mentioned characteristics................................................................2</keyLead>
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<keyStep id="871B2E012B51FFC699E2FD80FA45FD3A" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FD80FA42FDFD" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FD80FA42FDFD" pageId="6" pageNumber="7">
2. Pronotum lateral margins strongly crenate, without a continuous edge or border along margins, antenna IV stretched, longer than width (
<figureCitation id="A4D42A212B51FFC69821FDE2FE7DFDFD" box="[340,419,585,606]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Fig. 2-P</figureCitation>
) (body size
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)....................................................
<taxonomicName id="FBEF4D272B51FFC69C64FDE2FA83FDFD" ID-CoL="6ZFX5" authorityName="Fabricius" authorityYear="1775" box="[1297,1373,585,606]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="0E9BEAB62B51FFC69C64FDE2FA83FDFD" box="[1297,1373,585,606]" italics="true" pageId="6" pageNumber="7">rugosus</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FDCDFA45FD3A" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FDCDFA45FD3A" pageId="6" pageNumber="7">- Pronotum lateral margins not crenate, or partly crenate, or completely crenate, with a continuous edge or border along lateral margins.............................................................................................3</keyLead>
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3. Male head with greatly enlarged temples, with hind angles expanded posteriorly, scutellum with elongate lateral grooves, equal in length to mid-longitudinal sulcus (
<figureCitation id="A4D42A212B51FFC69B31FD15FD48FD70" box="[580,662,702,723]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Fig. 2-R</figureCitation>
) (body size 2.0
<quantity id="FB179B412B51FFC69A42FD15FC5FFD70" box="[823,897,702,723]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="mm" value="5.5">5.5 mm</quantity>
)................................
<taxonomicName id="FBEF4D272B51FFC69C71FD15FA83FD70" ID-CoL="678HF" baseAuthorityName="Eppelsheim" baseAuthorityYear="1877" box="[1284,1373,702,723]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gibbulus">
<emphasis id="0E9BEAB62B51FFC69C71FD15FA83FD70" box="[1284,1373,702,723]" italics="true" pageId="6" pageNumber="7">gibbulus</emphasis>
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group
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<paragraph id="3C5036A42B51FFC699E2FD77FA45FCAD" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FD77FA45FCAD" pageId="6" pageNumber="7">- Male head with developed temples but hind angles never expanded posteriorly, scutellum with short lateral grooves and their tips rounded.......................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="871B2E012B51FFC699E2FCBDFA45FC20" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FCBDFA42FCC5" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FCBDFA42FCC5" pageId="6" pageNumber="7">
4. Head and pronotum with surface sculptures composed of wrinkled striae and rides, clypeus squared and limited by a complete sharp suture. Tergite X with “rhomboid fusion” (
<bibRefCitation id="587E4B552B51FFC69BC4FC9FFC84FCEA" author="Makranczy" box="[689,858,820,841]" pageId="6" pageNumber="7" refString="Makranczy, Gy. (2006) Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici, 98, 29 - 119." type="journal article" year="2006">Makranczy, 2006</bibRefCitation>
,
<bibRefCitation id="587E4B552B51FFC69A11FC9FFC48FCEA" author="Makranczy" box="[868,918,820,841]" pageId="6" pageNumber="7" refString="Makranczy, Gy. (2011) Four new Neotropical species of Anotylus with an interesting sexual dimorphism (Coleoptera, Staphylinidae: Oxytelinae). Annales historico-naturales Musei Nationalis Hungarici, 103, 43 - 64." type="journal article" year="2011">2011</bibRefCitation>
), i.e. basolateral parts of tergite X separated and fused to tergites IX (body size 3.04.0 mm)........................................................
<taxonomicName id="FBEF4D272B51FFC69D9FFCFAFA83FCC5" ID-CoL="5V8FW" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[1258,1373,849,870]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="0E9BEAB62B51FFC69D9FFCFAFA83FCC5" box="[1258,1373,849,870]" italics="true" pageId="6" pageNumber="7">exasperatus</emphasis>
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group
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</paragraph>
<paragraph id="3C5036A42B51FFC699E2FCC5FA45FC20" blockId="6.[151,1436,438,1662]" box="[151,1435,878,899]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FCC5FA45FC20" box="[151,1435,878,899]" pageId="6" pageNumber="7">- Not like above description...............................................................................5</keyLead>
</paragraph>
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<keyStep id="871B2E012B51FFC699E2FC27FA42FC5A" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FC27FA42FC78" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FC27FA42FC78" pageId="6" pageNumber="7">
5. Head with post-ocular furrow absent, vertex with two longtitudinal furrows from clypeal area to occipital groove, pronotum lateral margins completely crenate, scutellum with median lobe of trilobed impression open behind (
<figureCitation id="A4D42A212B51FFC69DD2FC02FB25FC1D" box="[1191,1275,937,958]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Fig. 2-Q</figureCitation>
) (body size
<quantity id="FB179B412B51FFC69C04FC02FEF2FC78" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="5.4" metricValueMin="2.9" pageId="6" pageNumber="7" unit="mm" value="4.15" valueMax="5.4" valueMin="2.9">2.9 5.4 mm</quantity>
)..............................................................................
<taxonomicName id="FBEF4D272B51FFC69D9CFC6DFA80FC78" ID-CoL="66WG3" box="[1257,1374,966,987]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="0E9BEAB62B51FFC69D9CFC6DFA80FC78" box="[1257,1374,966,987]" italics="true" pageId="6" pageNumber="7">crassicornis</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FC4FFA42FC5A" blockId="6.[151,1436,438,1662]" box="[151,1436,996,1017]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FC4FFA42FC5A" box="[151,1436,996,1017]" pageId="6" pageNumber="7">- Not like above description...............................................................................6</keyLead>
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<paragraph id="3C5036A42B51FFC699E2FBAAFA42FBB5" blockId="6.[151,1436,438,1662]" box="[151,1436,1025,1046]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FBAAFA42FBB5" box="[151,1436,1025,1046]" pageId="6" pageNumber="7">6. Forebody more shinning, coriaceous, without fine longtitudinal striae............................................7</keyLead>
</paragraph>
<paragraph id="3C5036A42B51FFC699E2FBB5FA45FB90" blockId="6.[151,1436,438,1662]" box="[151,1435,1054,1075]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FBB5FA45FB90" box="[151,1435,1054,1075]" pageId="6" pageNumber="7">- Forebody not very much shining or dull, coarse, with fine longtitudinal striae.....................................10</keyLead>
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</keyStep>
<keyStep id="871B2E012B51FFC699E2FB97FA45FBCD" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FB97FA42FBF2" blockId="6.[151,1436,438,1662]" box="[151,1436,1084,1105]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FB97FA42FBF2" box="[151,1436,1084,1105]" pageId="6" pageNumber="7">
7. Head without occipital furrow (body size
<quantity id="FB179B412B51FFC69B04FB97FD38FBF2" box="[625,742,1084,1105]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.8" metricValueMin="3.6" pageId="6" pageNumber="7" unit="mm" value="4.2" valueMax="4.8" valueMin="3.6">3.64.8 mm</quantity>
)................................................
<taxonomicName id="FBEF4D272B51FFC69C5DFB97FA83FBF2" ID-CoL="66YCS" authorityName="Kraatz" authorityYear="1859" box="[1320,1373,1084,1105]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="0E9BEAB62B51FFC69C5DFB97FA83FBF2" box="[1320,1373,1084,1105]" italics="true" pageId="6" pageNumber="7">rufus</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FBF2FA45FBCD" blockId="6.[151,1436,438,1662]" box="[151,1435,1113,1134]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FBF2FA45FBCD" box="[151,1435,1113,1134]" pageId="6" pageNumber="7">- Head with at least part of occipital furrow..................................................................8</keyLead>
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<keyStep id="871B2E012B51FFC699E2FBDDFA42FB65" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FBDDFA42FB0A" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FBDDFA42FB0A" pageId="6" pageNumber="7">
8. Male clypeus depressed, anterior margin with two tubercle-like or horn-like projections, head convex, forebody sparely punc- tured, temples slightly dilated (body size 1.53.0 mm)..............................................
<taxonomicName id="FBEF4D272B51FFC69C6CFB3FFA83FB0A" ID-CoL="66WHQ" authorityName="Kraatz" authorityYear="1859" box="[1305,1373,1172,1193]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="micans">
<emphasis id="0E9BEAB62B51FFC69C6CFB3FFA83FB0A" box="[1305,1373,1172,1193]" italics="true" pageId="6" pageNumber="7">micans</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FB1AFA42FB65" blockId="6.[151,1436,438,1662]" box="[151,1436,1201,1222]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FB1AFA42FB65" box="[151,1436,1201,1222]" pageId="6" pageNumber="7">- Male clypeus not depressed, or depressed but anterior margin without obvious tubercle-like or horn-like projections.......9</keyLead>
</paragraph>
</keyStep>
<keyStep id="871B2E012B51FFC699E2FB65FA42FAFA" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FB65FA42FAA2" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FB65FA42FAA2" pageId="6" pageNumber="7">
9. Head with complete occipital furrow, eyes large, pronotum with mid-longitudinal sulcus deep and straight, paramedial sulci short and shallow, aedeagal paramere arm-like (body size 2.44.0 mm)..............................
<taxonomicName id="FBEF4D272B51FFC69D8FFB47FA83FAA2" ID-CoL="4BL4N" authorityName="Wollaston" authorityYear="1871" box="[1274,1373,1260,1281]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidifrons">
<emphasis id="0E9BEAB62B51FFC69D8FFB47FA83FAA2" box="[1274,1373,1260,1281]" italics="true" pageId="6" pageNumber="7">nitidifrons</emphasis>
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group
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</paragraph>
<paragraph id="3C5036A42B51FFC699E2FAA2FA42FAFA" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FAA2FA42FAFA" pageId="6" pageNumber="7">
- Parameres broad and contorted clockwise at apex 1/2, other characteristics not distinct, but with the following character com- bination: forebody with large and round punctuation, head as broad as pronotum (males), pronotum with paramedial sulci shal- low and broad, unobvious.................................................
<taxonomicName id="FBEF4D272B51FFC69D4EFAEFFADAFAFA" ID-CoL="5V8F2" authority="Bernhauer" authorityName="Bernhauer" box="[1083,1284,1348,1369]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="amicus">
<emphasis id="0E9BEAB62B51FFC69D4EFAEFFB44FAFA" box="[1083,1178,1348,1369]" italics="true" pageId="6" pageNumber="7">A. amicus</emphasis>
Bernhauer
</taxonomicName>
(
<taxonomicName id="FBEF4D272B51FFC69C67FAEFFA88FAFA" ID-CoL="5V8F2" box="[1298,1366,1348,1369]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="amicus">
<emphasis id="0E9BEAB62B51FFC69C67FAEFFA88FAFA" box="[1298,1366,1348,1369]" italics="true" pageId="6" pageNumber="7">amicus</emphasis>
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group)
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</paragraph>
</keyStep>
<keyStep id="871B2E012B51FFC699E2FACAFA45FA30" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FACAFA45FAD5" blockId="6.[151,1436,438,1662]" box="[151,1435,1377,1398]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FACAFA45FAD5" box="[151,1435,1377,1398]" pageId="6" pageNumber="7">10. Epistomal suture complete............................................................................. 11</keyLead>
</paragraph>
<paragraph id="3C5036A42B51FFC699E2FAD5FA45FA30" blockId="6.[151,1436,438,1662]" box="[151,1435,1406,1427]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FAD5FA45FA30" box="[151,1435,1406,1427]" pageId="6" pageNumber="7">- Epistomal suture not complete.......................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="871B2E012B51FFC699E2FA37FA42FA6D" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2FA37FA42FA12" blockId="6.[151,1436,438,1662]" box="[151,1436,1436,1457]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FA37FA42FA12" box="[151,1436,1436,1457]" pageId="6" pageNumber="7">
11. Pronotum lateral margins crenate (body size 2.03.0 mm)...........................................
<taxonomicName id="FBEF4D272B51FFC69C71FA37FA83FA12" ID-CoL="66YB8" authorityName="Gravenhorst" authorityYear="1802" box="[1284,1373,1436,1457]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="0E9BEAB62B51FFC69C71FA37FA83FA12" box="[1284,1373,1436,1457]" italics="true" pageId="6" pageNumber="7">nitidulus</emphasis>
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group
</keyLead>
</paragraph>
<paragraph id="3C5036A42B51FFC699E2FA12FA42FA6D" blockId="6.[151,1436,438,1662]" box="[151,1436,1465,1486]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FA12FA42FA6D" box="[151,1436,1465,1486]" pageId="6" pageNumber="7">
- Pronotum lateral margins not crenate (body size 1.0
<quantity id="FB179B412B51FFC69BA9FA12FCF6FA6D" box="[732,808,1465,1486]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="2.8">2.8 mm</quantity>
)...................................
<taxonomicName id="FBEF4D272B51FFC69DA0FA12FA83FA6D" ID-CoL="6ZFVD" authorityName="Block" authorityYear="1799" box="[1237,1373,1465,1486]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tetracarinatus">
<emphasis id="0E9BEAB62B51FFC69DA0FA12FA83FA6D" box="[1237,1373,1465,1486]" italics="true" pageId="6" pageNumber="7">tetracarinatus</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FA7DFA42FA48" blockId="6.[151,1436,438,1662]" box="[151,1436,1494,1515]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FA7DFA42FA48" box="[151,1436,1494,1515]" pageId="6" pageNumber="7">
12. Sternite VII with anterior angles obviously produced (
<figureCitation id="A4D42A212B51FFC69B92FA7DFCE1FA48" box="[743,831,1494,1515]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Figs. 2-S</figureCitation>
) (body size
<quantity id="FB179B412B51FFC69AC3FA7DFBF5FA48" box="[950,1067,1494,1515]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="6" pageNumber="7" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
).................
<taxonomicName id="FBEF4D272B51FFC69C77FA7DFA80FA48" ID-CoL="66WGP" authorityName="Erichson" authorityYear="1840" box="[1282,1374,1494,1515]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="intricatus">
<emphasis id="0E9BEAB62B51FFC69C77FA7DFA80FA48" box="[1282,1374,1494,1515]" italics="true" pageId="6" pageNumber="7">intricatus</emphasis>
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<paragraph id="3C5036A42B51FFC699E2FA5FFA45F9AA" blockId="6.[151,1436,438,1662]" box="[151,1435,1524,1545]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2FA5FFA45F9AA" box="[151,1435,1524,1545]" pageId="6" pageNumber="7">- Sternite VII with anterior angles not produced, at least not as sharp-angles....................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="871B2E012B51FFC699E2F9BAFA42F9DD" pageId="6" pageNumber="7">
<paragraph id="3C5036A42B51FFC699E2F9BAFA42F9E0" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2F9BAFA42F9E0" pageId="6" pageNumber="7">
13. Scutellum with median lobe of trilobed impression open behind (
<figureCitation id="A4D42A212B51FFC69A21F9BAFC6CF985" box="[852,946,1553,1574]" captionStart="FIGURE 2" captionStartId="9.[151,250,1840,1862]" captionTargetBox="[268,1337,209,1814]" captionTargetId="figure@9.[251,1343,193,1828]" captionTargetPageId="9" captionText="FIGURE 2. Head and prontoum (A N). A, amicus group (A. granadillae); B, cimicoides group (A. chinensis); C, complanatus group (A. complanatoides); D, crassicornis group (A. flavipennis [from Makranczy, 2013]); E, gibbulus group (A. megacephalus); F, tetracarinatus group (A. tetracarinatus); G, intricatus group (A. intricatus); H, micans group (A. uncifer); I, nitidulus group (A. nitidulus); J, rufus group (A. rufus); K, rugosus group (A. rugosus); L, sculpturatus group (A. cognatus); M, exasperatus group (A. fraternus); N, A. nitidifrons group (A. nitouensis); O, tergites IX with enlarged platelike ventral strut (Anotylus sp.); P. antenna segment IV; Q R, scutellum; S, male Sternite VIII (A. intricatus). Scales: D = 0.55 mm; rest, 0.5 mm." httpUri="https://zenodo.org/record/1063750/files/figure.png" pageId="6" pageNumber="7">Figs. 2-Q</figureCitation>
), head and pronotum with much more microsculp- ture, abdomen with thick setae, body lager (body size 2.54.0 mm)...............................
<taxonomicName id="FBEF4D272B51FFC69D96F985FA80F9E0" ID-CoL="66WFQ" authorityName="Erichson" authorityYear="1839" box="[1251,1374,1582,1603]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="0E9BEAB62B51FFC69D96F985FA80F9E0" box="[1251,1374,1582,1603]" italics="true" pageId="6" pageNumber="7">complanatus</emphasis>
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group
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</paragraph>
<paragraph id="3C5036A42B51FFC699E2F9E7FA42F9DD" blockId="6.[151,1436,438,1662]" pageId="6" pageNumber="7">
<keyLead id="871E95912B51FFC699E2F9E7FA42F9DD" pageId="6" pageNumber="7">
- Scutellum with median lobe of trilobed impression closed behind, head and pronotum with less microsculpture, abdomen with fine setae (body size
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)...........................................................
<taxonomicName id="FBEF4D272B51FFC69D95F9C2FA80F9DD" ID-CoL="75H5T" authorityName="Gravenhorst" authorityYear="1806" box="[1248,1374,1641,1662]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="0E9BEAB62B51FFC69D95F9C2FA80F9DD" box="[1248,1374,1641,1662]" italics="true" pageId="6" pageNumber="7">sculpturatus</emphasis>
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