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<mods:title id="06A5E18D54CD8A626E998E82FD0EE83C">Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China</mods:title>
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<mods:namePart id="D73C23510E36F52E003B994CE3E448D0">Wang, Lin-Fei</mods:namePart>
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<mods:namePart id="2FBBD163416CB9208B6A05231F3AF840">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="B44687B22B15FF8599E2FF3CFCF3FB02" ID-DOI="http://doi.org/10.5281/zenodo.6033747" ID-GBIF-Taxon="137467067" ID-Zenodo-Dep="6033747" LSID="urn:lsid:plazi:treatment:B44687B22B15FF8599E2FF3CFCF3FB02" httpUri="http://treatment.plazi.org/id/B44687B22B15FF8599E2FF3CFCF3FB02" lastPageId="69" lastPageNumber="70" pageId="66" pageNumber="67">
<subSubSection id="74F5652F2B15FF8299E2FF3CFEDAFF70" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="3C5036A42B15FF8299E2FF3CFDC6FF11" blockId="66.[151,536,151,211]" box="[151,536,151,178]" pageId="66" pageNumber="67">
<heading id="671881C82B15FF8299E2FF3CFDC6FF11" bold="true" box="[151,536,151,178]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
<taxonomicName id="FBEF4D272B15FF8299E2FF3CFDC6FF11" ID-CoL="66WHZ" authority="(Lohse, 1963)" baseAuthorityName="Lohse" baseAuthorityYear="1963" box="[151,536,151,178]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="0E9BEAB62B15FF8299E2FF3CFDC6FF11" bold="true" box="[151,536,151,178]" pageId="66" pageNumber="67">
<emphasis id="0E9BEAB62B15FF8299E2FF3CFEB7FF12" bold="true" box="[151,361,151,177]" italics="true" pageId="66" pageNumber="67">Anotylus mutator</emphasis>
(
<bibRefCitation id="587E4B552B15FF82980CFF33FDCEFF11" author="Lohse" box="[377,528,152,178]" pageId="66" pageNumber="67" refString="Lohse, G. A. (1963) Neue Staphyliniden aus Mitteleuropa und dem Alpengebiet. Entomologische Blatter fur Biologie und Systematik der Kafer, 59 (3), 168 - 178." type="journal article" year="1963">Lohse, 1963</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3C5036A42B15FF8299E2FF11FEDAFF70" blockId="66.[151,536,151,211]" box="[151,260,186,211]" pageId="66" pageNumber="67">(Figs. 19)</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B15FF8299E2FEA9FEEBFE11" pageId="66" pageNumber="67" type="description">
<paragraph id="3C5036A42B15FF8299E2FEA9FEEBFE11" blockId="66.[151,1436,258,434]" pageId="66" pageNumber="67">
<bibRefCitation id="587E4B552B15FF8299E2FEA9FEC7FEBB" author="Lohse" box="[151,281,258,280]" pageId="66" pageNumber="67" refString="Lohse, G. A. (1963) Neue Staphyliniden aus Mitteleuropa und dem Alpengebiet. Entomologische Blatter fur Biologie und Systematik der Kafer, 59 (3), 168 - 178." type="journal article" year="1963">Lohse, 1963</bibRefCitation>
: 174 (
<taxonomicName id="FBEF4D272B15FF82982FFEA9FE6CFEBB" authorityName="Gravenhorst" authorityYear="1802" box="[346,434,258,280]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF82982FFEA9FE6CFEBB" box="[346,434,258,280]" italics="true" pageId="66" pageNumber="67">Oxytelus</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="FBEF4D272B15FF829B53FEA9FDA0FEBB" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[550,638,258,280]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829B53FEA9FDA0FEBB" box="[550,638,258,280]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; Type locality: Garlstorfer Forst, Kreis Harburg, Niedersachsen);
<bibRefCitation id="587E4B552B15FF829C51FEA9FA47FEBB" author="Toth" box="[1316,1433,258,280]" pageId="66" pageNumber="67" refString="Toth, L. (1974) Oxytelus mutator Lohse a Karpatmedence faunajaban (Coleoptera: Staphylinidae). Folia Entomologica Hungarica, 27 (2), 221 - 226." type="journal article" year="1974">Tóth, 1974</bibRefCitation>
: 222, 223 (
<taxonomicName id="FBEF4D272B15FF829840FE8AFE53FE94" authorityName="Gravenhorst" authorityYear="1802" box="[309,397,289,311]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829840FE8AFE53FE94" box="[309,397,289,311]" italics="true" pageId="66" pageNumber="67">Oxytelus</emphasis>
</taxonomicName>
; characters; Hungary);
<bibRefCitation id="587E4B552B15FF829BF3FE8AFCD4FE94" author="Lohse" box="[646,778,289,311]" pageId="66" pageNumber="67" refString="Lohse, G. A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. &amp; Lohse, G. (Eds.), Die Kafer Mitteleuropas. Vol. 4. Krefeld, Goecke &amp; Evers, 4, 1 - 264." type="journal article" year="1964">Lohse, 1964</bibRefCitation>
: 85 (
<taxonomicName id="FBEF4D272B15FF829A36FE8AFC42FE94" authorityName="Gravenhorst" authorityYear="1802" box="[835,924,289,311]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829A36FE8AFC42FE94" box="[835,924,289,311]" italics="true" pageId="66" pageNumber="67">Oxytelus</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="FBEF4D272B15FF829D61FE8AFBB3FE94" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[1044,1133,289,311]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829D61FE8AFBB3FE94" box="[1044,1133,289,311]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; characters; central Europe);
<bibRefCitation id="587E4B552B15FF8299B2FEEBFE5FFEF5" author="Hammond" box="[199,385,320,342]" pageId="66" pageNumber="67" refString="Hammond, P. M. (1976 a) A review of the genus Anotylus C. G. Thomson (Coleoptera: Staphylinidae). Bulletin of the British Museum (Natural History) (Entomology), 33, 139 - 187." type="journal article" year="1976" yearSuffix="a">Hammond, 1976a</bibRefCitation>
: 159 (
<taxonomicName id="FBEF4D272B15FF8298B7FEEBFDC5FEF5" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[450,539,320,342]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF8298B7FEEBFDC5FEF5" box="[450,539,320,342]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
);
<bibRefCitation id="587E4B552B15FF829B44FEEBFD6EFEF5" author="Lucht" box="[561,688,320,342]" pageId="66" pageNumber="67" refString="Lucht, W. H. (1987) Die Kafer Mitteleuropas. Katalog. Goecke and Evers, Krefeld, 342 pp." type="book" year="1987">Lucht, 1987</bibRefCitation>
: 93 (
<taxonomicName id="FBEF4D272B15FF829B91FEEBFCE3FEF5" authorityName="Gravenhorst" authorityYear="1802" box="[740,829,320,342]" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829B91FEEBFCE3FEF5" box="[740,829,320,342]" italics="true" pageId="66" pageNumber="67">Oxytelus</emphasis>
</taxonomicName>
; checklist; Germany; Poland; Austria; Denmark);
<bibRefCitation id="587E4B552B15FF829C31FEEAFF23FED7" author="Herman" pageId="66" pageNumber="67" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the second millennium. Vol. 3, Oxyteline group. Bulletin of the American Museum of Natural History, 265, 1067 - 1806." type="book chapter" year="2001">Herman, 2001</bibRefCitation>
: 1372 (
<taxonomicName id="FBEF4D272B15FF82983AFEF5FE76FED7" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[335,424,350,372]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF82983AFEF5FE76FED7" box="[335,424,350,372]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; catalog);
<bibRefCitation id="587E4B552B15FF829B6FFEF5FD66FED7" author="Smetana" box="[538,696,350,372]" pageId="66" pageNumber="67" refString="Smetana, A. (2004) Subfamily Oxytelinae Fleming, 1821. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera: Hydrophiloidea - Histeroidea - Staphylinoidea. Vol. 2. Apollo Books, Stenstrup, pp. 511 - 535." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 514 (
<taxonomicName id="FBEF4D272B15FF829B89FEF5FC8AFED7" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[764,852,350,372]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829B89FEF5FC8AFED7" box="[764,852,350,372]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; Palaearctic catalog; distribution);
<bibRefCitation id="587E4B552B15FF829DB3FEF5FF23FE30" author="Schulke" pageId="66" pageNumber="67" refString="Schulke, M. &amp; Smetana, A. (2015) Oxytelinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. 1 (Hydrophiloidea-Staphylinoidea). Leiden &amp; Boston, Brill, pp. 757 - 795." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
: 771 (
<taxonomicName id="FBEF4D272B15FF82984EFED6FE4AFE30" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[315,404,381,403]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF82984EFED6FE4AFE30" box="[315,404,381,403]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; Palaearctic catalog; distribution); Semionenkov et al., 2015: 33 (
<taxonomicName id="FBEF4D272B15FF829D5BFED6FB59FE30" authorityName="Wang &amp; Zhou &amp; Lü" authorityYear="2017" box="[1070,1159,381,403]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E9BEAB62B15FF829D5BFED6FB59FE30" box="[1070,1159,381,403]" italics="true" pageId="66" pageNumber="67">Anotylus</emphasis>
</taxonomicName>
; West of the European part of Russia).
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B15FF8299E2FE75FC10FD24" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="3C5036A42B15FF8299E2FE75FC10FD24" blockId="66.[151,1436,478,719]" pageId="66" pageNumber="67">
<emphasis id="0E9BEAB62B15FF8299E2FE75FDCDFE54" bold="true" box="[151,531,478,503]" pageId="66" pageNumber="67">Material examined. EUROPE:</emphasis>
<specimenCount id="2AE9FD2D2B15FF829B54FE74FDABFE54" box="[545,629,478,503]" pageId="66" pageNumber="67" type="male">1 male</specimenCount>
, Bruck N. Ö [Nieder-Österreich], Bernhauer// Chicago NHMus M. Bernhauer Collection//
<taxonomicName id="FBEF4D272B15FF8298EAFDA8FC5FFDB8" authority="(Lohse) P. M. Hammond" authorityName="(Lohse) P. M. Hammond" box="[415,897,514,539]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mutotar">
<emphasis id="0E9BEAB62B15FF8298EAFDA8FDBCFDB9" box="[415,610,515,539]" italics="true" pageId="66" pageNumber="67">Anotylus mutotar</emphasis>
(Lohse) P. M. Hammond
</taxonomicName>
det. 1977// FMNHINS 2821689 Field Museum (
<collectionCode id="5AFEAE612B15FF8299EAFD8CFF29FD9C" box="[159,247,551,575]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="66" pageNumber="67">FMNH</collectionCode>
);
<specimenCount id="2AE9FD2D2B15FF829878FD8CFEA8FD9C" box="[269,374,550,575]" pageId="66" pageNumber="67" type="female">1 female</specimenCount>
, Hung. Trencsén Nagy-Sziklás [leg.] Kocsi//
<collectionCode id="5AFEAE612B15FF829AF8FD8CFC3FFD9C" box="[909,993,551,575]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="66" pageNumber="67">CNHM</collectionCode>
1955 Karl Brancsik Colln ex Eduard Knirsch// FMNHINS 2821690 Field Museum (
<collectionCode id="5AFEAE612B15FF829BDCFDE0FCDFFDC0" box="[681,769,587,611]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="66" pageNumber="67">FMNH</collectionCode>
);
<collectingCountry id="44F876342B15FF829A61FDE1FCBDFDC0" box="[788,867,586,611]" name="Russia" pageId="66" pageNumber="67">
<emphasis id="0E9BEAB62B15FF829A61FDE1FCBDFDC0" bold="true" box="[788,867,586,611]" pageId="66" pageNumber="67">Russia</emphasis>
</collectingCountry>
:
<specimenCount id="2AE9FD2D2B15FF829A07FDE0FC09FDC0" box="[882,983,586,611]" pageId="66" pageNumber="67" type="female">1 female</specimenCount>
, Transsilv. Kronstadt// Chicago NHMus M. Bernhauer Collection // FMNHINS 2821691 Field Museum (
<collectionCode id="5AFEAE612B15FF829A1EFDC4FC1CFD24" box="[875,962,623,647]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="66" pageNumber="67">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B15FF8599B2FD39FF35FB96" lastPageId="69" lastPageNumber="70" pageId="66" pageNumber="67" type="description">
<paragraph id="3C5036A42B15FF8299B2FD39FB21FD6D" blockId="66.[151,1436,478,719]" pageId="66" pageNumber="67">
<emphasis id="0E9BEAB62B15FF8299B2FD39FEAAFD08" bold="true" box="[199,372,658,683]" pageId="66" pageNumber="67">Redescription.</emphasis>
Body brownness, shiny; elytra yellow brown, with base slightly black; legs yellow. Forebody pubescent, moderately punctate and striate. Body length [average] male,
<quantity id="FB179B412B15FF829AB2FD1CFBF0FD6D" box="[967,1070,695,719]" metricMagnitude="-3" metricUnit="m" metricValue="4.98" pageId="66" pageNumber="67" unit="mm" value="4.98">4.98 mm</quantity>
; female,
<quantity id="FB179B412B15FF829DE1FD1DFB22FD6D" box="[1172,1276,694,719]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="66" pageNumber="67" unit="mm" value="3.65">3.65 mm</quantity>
.
</paragraph>
<caption id="6890662C2B15FF8299E2F8F1FE8CF848" httpUri="https://zenodo.org/record/1063842/files/figure.png" pageId="66" pageNumber="67" targetBox="[163,1409,802,1817]" targetPageId="66">
<paragraph id="3C5036A42B15FF8299E2F8F1FE8CF848" blockId="66.[151,1436,1882,2027]" pageId="66" pageNumber="67">
<emphasis id="0E9BEAB62B15FF8299E2F8F1FEEDF8D3" bold="true" box="[151,307,1882,1904]" pageId="66" pageNumber="67">FIGURE 19-1.</emphasis>
<taxonomicName id="FBEF4D272B15FF82984CF8F1FDA5F8D3" authority="(Lohse, 1963)" authorityName="Lohse" authorityYear="1963" baseAuthorityName="Lohse" baseAuthorityYear="1963" box="[313,635,1882,1904]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="0E9BEAB62B15FF82984CF8F1FE36F8D3" box="[313,488,1882,1904]" italics="true" pageId="66" pageNumber="67">Anotylus mutator</emphasis>
(Lohse, 1963)
</taxonomicName>
A, forebody male; B, habitus female; C, head male; D, pronotum male; E, head and pronotum female; F, sternite VII male; G, sternite VIII male; H, sternite VIII female; I, median lobe of aedeagus (ventral view); J, median lobe of aedeagus (dorsal view); K, median lobe of aedeagus (lateral view); L, left paramere (in situ) of aedeagus (lateral view); M, parameres (in situ) of aedeagus (vertral view); N, spermatheca. Scales: A–E= 0.5 mm; N = 0.05 mm; rest, 0.2 mm.
</paragraph>
</caption>
<caption id="6890662C2B14FF8399E2F81AFB2AF845" httpUri="https://zenodo.org/record/1063844/files/figure.png" pageId="67" pageNumber="68" targetBox="[203,1380,257,1857]" targetPageId="67">
<paragraph id="3C5036A42B14FF8399E2F81AFB2AF845" blockId="67.[151,1436,1969,2022]" pageId="67" pageNumber="68">
<emphasis id="0E9BEAB62B14FF8399E2F81AFEE8F864" bold="true" box="[151,310,1969,1991]" pageId="67" pageNumber="68">FIGURE 19-2.</emphasis>
<taxonomicName id="FBEF4D272B14FF83984AF81AFD56F864" authority="(Lohse, 1963)" authorityName="Lohse" authorityYear="1963" baseAuthorityName="Lohse" baseAuthorityYear="1963" box="[319,648,1969,1991]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="0E9BEAB62B14FF83984AF81AFE2EF864" box="[319,496,1969,1991]" italics="true" pageId="67" pageNumber="68">Anotylus mutator</emphasis>
(Lohse, 1963)
</taxonomicName>
A, head and pronotum male; B, head and pronotum female; C, sternite VII male; D, sternite VIII male; E, sternite VII female; F, sternite VIII female. Scale: A, B= 0.5 mm; rest, 0.2 mm.
</paragraph>
</caption>
<caption id="6890662C2B13FF8499E2F883FC49F8D8" httpUri="https://zenodo.org/record/1063846/files/figure.png" pageId="68" pageNumber="69" targetBox="[220,1384,202,1805]" targetPageId="68">
<paragraph id="3C5036A42B13FF8499E2F883FC49F8D8" blockId="68.[151,1436,1832,1915]" pageId="68" pageNumber="69">
<emphasis id="0E9BEAB62B13FF8499E2F883FEEAF89D" bold="true" box="[151,308,1832,1854]" pageId="68" pageNumber="69">FIGURE 19-3.</emphasis>
<taxonomicName id="FBEF4D272B13FF84984FF883FDA5F89D" authority="(Lohse, 1963)" authorityName="Lohse" authorityYear="1963" baseAuthorityName="Lohse" baseAuthorityYear="1963" box="[314,635,1832,1854]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="0E9BEAB62B13FF84984FF883FE36F89D" box="[314,488,1832,1854]" italics="true" pageId="68" pageNumber="69">Anotylus mutator</emphasis>
(Lohse, 1963)
</taxonomicName>
A, median lobe of aedeagus (ventral view); B, median lobe of aedeagus (dorsal view); C, median lobe of aedeagus (lateral view); D, parameres (in situ) of aedeagus (ventral view); E, left paramere (in situ) of aedeagus (lateral view); F, spermatheca. Scales: F = 0.05 mm; rest, 0.2 mm.
</paragraph>
</caption>
<paragraph id="3C5036A42B13FF8599B2F803FBDCFEE3" blockId="68.[151,1436,1960,2021]" lastBlockId="69.[151,1437,151,1185]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<emphasis id="0E9BEAB62B13FF8499B2F803FEDFF863" box="[199,257,1960,1984]" italics="true" pageId="68" pageNumber="69">Male</emphasis>
: Head (Fig. 19-1C, 19-2A) slightly broader, but narrower than pronotum. Disc extremely convex, with moderate-sized round punctures and longitudinal striae, denser at sides; mid-longitudinal sulcus short and shallow, with deeper impression on each side before base. Supra-antennal ridge with margin incrassate, and running backward to near base. Temples not dilated, approximately as long as eyes. Clypeus nearly square, about 1/3 of total head length, convex and coriaceous, with spare small punctures; anterior margin slightly straight and not produced. Epistomal suture with lateral portions straight and parallel, running backward to level of anterior margin of eyes; separated by coriaceous ground sculpture from the rest of the surface.
</paragraph>
<paragraph id="3C5036A42B12FF8599B2FEE7FC7AFE24" blockId="69.[151,1437,151,1185]" pageId="69" pageNumber="70">Occipital suture slightly curved forward in middle. Antenna filiform, article IV globular, V–X short and transverse, with ultimate article stilliform. Neck without smooth area.</paragraph>
<paragraph id="3C5036A42B12FF8599B2FE3EFEAEFD53" blockId="69.[151,1437,151,1185]" pageId="69" pageNumber="70">Pronotum (Fig. 19-1D, 19-2A) transverse, broadest at near anterior 1/3, with moderate-sized round punctures and dense striae, denser at sides; disc slightly convex, median sulcus straight and deep, with two shallow and broad paramedial sulci, bounded by two obtuse longitudinal carina, with shallow depression close to lateral margins. Anterior margin slightly bi-emarginate. Anterior angles slightly produced. Posterior angles obvious; lateral margin slightly crenate on posterior 1/4. Scutellum with crest-shaped impression, with sharply pointed median lobe. Elytra densely striate and punctate, not pubescent; lateral margin curved and posterior margins truncate and slightly slant. Abdomen coriaceous, covered with punctures and pubescence. Sternite VII (Fig. 19-1F, 19-2C) with posterior margin broadly but slightly emarginate, bearing long setae laterally, in middle with two small flat tubercles, not beyond posterior margin; Sternite VIII (Fig. 19-1G, 19-2D) with posterior margin deeply emarginate and carrying long setae laterally.</paragraph>
<paragraph id="3C5036A42B12FF8599B2FD57FAF0FCDC" blockId="69.[151,1437,151,1185]" pageId="69" pageNumber="70">
Aedeagus (Fig.
<date id="485110642B12FF85980AFD57FE66FCB7" box="[383,440,764,788]" pageId="69" pageNumber="70" value="1919-01-01">19-1I</date>
–K, 19-3A–C) with median lobe oval, inflated at base and shortly narrowing to apex; apical orifice large and ventral orifice crescentic; internal sac of median lobe with complicated sclerites. Parameres (Fig. 19-1L, M, 19-3D, E) closely situated to each other; with basic part flattened and enlarged, apex part narrow and elongate; with rounded process at middle of ventral edge of each paramere, furnished with long seta.
</paragraph>
<paragraph id="3C5036A42B12FF8599B2FC27FF35FB96" blockId="69.[151,1437,151,1185]" pageId="69" pageNumber="70">
<emphasis id="0E9BEAB62B12FF8599B2FC27FEC5FC07" box="[199,283,908,932]" italics="true" pageId="69" pageNumber="70">Female</emphasis>
. Head (Fig. 19-1E, 19-2B) smaller than male, narrower than pronotum (Fig. 19-1E, 19-2B). Temples shorter than eyes. Sternite VII (Fig. 19-2E) with posterior margin nearly straight. Sternite VIII (Fig. 19-1H, 19-2F) with posterior margin broadly shallowly bi-emarginate, obtusely angulate in middle and slightly beyond lateral angles posteriorly. Spermatheca (Fig. 19-1N, 19-3F) with base bulb-like, with apical part clavate and slightly curved.
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B12FF8599B2FB94FDAAFBF4" box="[199,628,1087,1112]" pageId="69" pageNumber="70" type="distribution">
<paragraph id="3C5036A42B12FF8599B2FB94FDAAFBF4" blockId="69.[151,1437,151,1185]" box="[199,628,1087,1112]" pageId="69" pageNumber="70">
<emphasis id="0E9BEAB62B12FF8599B2FB94FEBEFBFB" bold="true" box="[199,352,1087,1112]" pageId="69" pageNumber="70">Distribution.</emphasis>
<collectingCountry id="44F876342B12FF859812FB94FE6AFBFB" box="[359,436,1087,1112]" name="Russia" pageId="69" pageNumber="70">Russia</collectingCountry>
,
<collectingCountry id="44F876342B12FF8598B5FB94FDCFFBFB" box="[448,529,1087,1112]" name="Turkey" pageId="69" pageNumber="70">Turkey</collectingCountry>
, Europe.
</paragraph>
</subSubSection>
<subSubSection id="74F5652F2B12FF8599B2FBCFFCF3FB02" pageId="69" pageNumber="70" type="discussion">
<paragraph id="3C5036A42B12FF8599B2FBCFFCF3FB02" blockId="69.[151,1437,151,1185]" pageId="69" pageNumber="70">
<emphasis id="0E9BEAB62B12FF8599B2FBCFFEEAFBDE" bold="true" box="[199,308,1124,1149]" pageId="69" pageNumber="70">Remarks</emphasis>
. Similar to
<taxonomicName id="FBEF4D272B12FF8598B3FBCFFDAAFBDF" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[454,628,1124,1148]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="0E9BEAB62B12FF8598B3FBCFFDAAFBDF" box="[454,628,1124,1148]" italics="true" pageId="69" pageNumber="70">A. sculpturatus</emphasis>
</taxonomicName>
, distinguished mainly by sternite VII, parameres and internal sac of aedeagus, so we redescribed and illustrated in this passage.
</paragraph>
</subSubSection>
</treatment>
</document>
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