175 lines
16 KiB
XML
175 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1174.107314" ID-Pensoft-Pub="1313-2970-1174-35" ID-Pensoft-UUID="635A059AEF505E17B338F6484B5F4DBF" ID-ZooBank="91E908240C45471A86BBA24ADC35E743" ModsDocID="1313-2970-1174-35" checkinTime="1691559557484" checkinUser="pensoft" docAuthor="Harris, Steven C. & Armitage, Brian J." docDate="2023" docId="B10FC2854AC751759DE9DAFFE554FF56" docLanguage="en" docName="ZooKeys 1174: 35-74" docOrigin="ZooKeys 1174" docPubDate="2023-08-08" docSource="http://dx.doi.org/10.3897/zookeys.1174.107314" docTitle="Metrichia cascada Harris & Armitage 2023, sp. nov." docType="treatment" docUuid="F7F609A6-F108-48BD-A3EE-1E17238D4002" docUuidSource="ZooBank" docVersion="1" id="635A059AEF505E17B338F6484B5F4DBF" lastPageNumber="35" masterDocId="635A059AEF505E17B338F6484B5F4DBF" masterDocTitle="The Trichoptera of Panama XXII. Sixteen new microcaddisfly species (Trichoptera, Hydroptilidae)" masterLastPageNumber="74" masterPageNumber="35" pageNumber="35" updateTime="1691559557484" updateUser="pensoft">
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<mods:mods id="C0103E52C0815C8B43C0895C7C1D2458" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="724ADF6BF1B5BEB2CC41F5A1C101A55E">
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<mods:title id="D55A9866A354E78DAAFDF680F364BEA0">The Trichoptera of Panama XXII. Sixteen new microcaddisfly species (Trichoptera, Hydroptilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7695A5702C3CC310322A72EF802CEF8A" type="personal">
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<mods:role id="69C1C12DA7B5BA46EE97375F5F3883AA">
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<mods:roleTerm id="09BF058DE23C7DEF0D7302858968D7E6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E9A14A5962F355BA5E4D35D11D1764B">Harris, Steven C.</mods:namePart>
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<mods:nameIdentifier id="60A836786E481B48C0DCB3789B999D61" type="ORCID">https://orcid.org/0000-0002-6432-7462</mods:nameIdentifier>
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<mods:affiliation id="7FD6F71264EF3ADD1CD6BB14DB38B443">Department of Biology and Environmental Sciences, Western Pennsylvania University, Clarion, PA 16214, USA & Museo de Peces de Agua Dulce e Invertebrados, Universidad Autonoma de Chiriqui, David, Panama</mods:affiliation>
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</mods:name>
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<mods:name id="6D72365720C16F14F65E96812F1160DD" type="personal">
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<mods:roleTerm id="1F1D435A5D3C32A8DC1D71C1CCD493A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D0277A2DDADB00DD4207B7C2C6EDCE3D">Armitage, Brian J.</mods:namePart>
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<mods:nameIdentifier id="FC55E17028488DD2BBCA2FE9077B29D5" type="ORCID">https://orcid.org/0000-0003-3182-1533</mods:nameIdentifier>
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<mods:affiliation id="1D7F646B09F3818D2E491D7CDD126543">Museo de Peces de Agua Dulce e Invertebrados, Universidad Autonoma de Chiriqui, David, Panama</mods:affiliation>
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<mods:nameIdentifier id="4E75FE23E27A64BB7179ABCBA29D99F3" type="email">tobikera89@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="47056CB9DE8E170EB526BFD033D2CEE2">text</mods:typeOfResource>
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<mods:titleInfo id="82DC0E4A04982F906F7AAEF74F5A6E05">
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<mods:title id="1FACEBBA3A9700E9F8638C9CD8678DAD">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="738E48825DD8A0BFEF8A5A6A60063C26">
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<mods:date id="48C9A470F81DEFFCE7A9E130A2492F93">2023</mods:date>
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<mods:detail id="D9249E3098019B304955CFB52F3A7938" type="pubDate">
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<mods:number id="5248561EE771BB8AC2E4809E1009920C">2023-08-08</mods:number>
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</mods:detail>
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<mods:detail id="F8F19769BAA69B1B88D182212D476915" type="volume">
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<mods:number id="8D265D288BEF029BA3980B3D960EAD1F">1174</mods:number>
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</mods:detail>
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<mods:extent id="D118B70A49F72F537B239C480D6D876E" unit="page">
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<mods:start id="FB6107ADB74D0FF6EB402304F0804163">35</mods:start>
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<mods:end id="6CB914E2DB57AF7E6FC71C8D9CECDB43">74</mods:end>
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<mods:location id="386368592E00C55E624E59D4FCC96E99">
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<mods:url id="EDDE0D7838C53A9AD6AE3D78C22652DB">http://dx.doi.org/10.3897/zookeys.1174.107314</mods:url>
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</mods:location>
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<mods:classification id="99A9CF6460ED13995EA4842729A35F4A">journal article</mods:classification>
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<mods:identifier id="2217B87A701896C1E052EAF1D0DF5142" type="DOI">http://dx.doi.org/10.3897/zookeys.1174.107314</mods:identifier>
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<mods:identifier id="FC94BAB5396EB832B67D511B4AB85262" type="Pensoft-Pub">1313-2970-1174-35</mods:identifier>
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<mods:identifier id="B6C816D16B12CB2F84B44569A085D64A" type="ZooBank">91E908240C45471A86BBA24ADC35E743</mods:identifier>
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<mods:identifier id="F6794E2034AF3A0684CD48EEDCDF8BD6" type="Pensoft-UUID">635A059AEF505E17B338F6484B5F4DBF</mods:identifier>
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</mods:mods>
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<treatment id="B10FC2854AC751759DE9DAFFE554FF56" LSID="urn:lsid:zoobank.org:act:F7F609A6-F108-48BD-A3EE-1E17238D4002" httpUri="http://treatment.plazi.org/id/B10FC2854AC751759DE9DAFFE554FF56" lastPageNumber="35" pageId="0" pageNumber="35">
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<subSubSection id="613F91B3E9D1F4C97F10061CC07ABF5E" pageId="0" pageNumber="35" type="nomenclature">
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<paragraph id="878F8B0C20B4981151E5538F374366C4" pageId="0" pageNumber="35">
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<taxonomicName id="A235333E05172908FBAEFEB6EBE43843" LSID="https://zoobank.org/F7F609A6-F108-48BD-A3EE-1E17238D4002" authority="Harris & Armitage, 2023" authorityName="Harris & Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Metrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrichia cascada" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cascada" status="sp. nov.">Metrichia cascada</taxonomicName>
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<taxonomicNameLabel id="E5B5A9AC974C701AABAACA146DB80207" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="2D54F121A792D6E32770CCEE16F47D70" pageId="0" pageNumber="35" type="description">
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<paragraph id="83D1534EBCFBC1A783D35DEE31219421" pageId="0" pageNumber="35">
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<figureCitation id="29C71F8CF27C3F0B18BCE17014BA9304" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Metrichia cascada sp. nov., male holotype, genitalia A left lateral B dorsal C ventral D phallus, dorsal E phallus apex, left lateral." figureDoi="10.3897/zookeys.1174.107314.figure10" httpUri="https://binary.pensoft.net/fig/890135" pageId="0" pageNumber="35">Fig. 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EEBFA4ED476250360EA01CFF86D4382B" pageId="0" pageNumber="35" type="type locality">
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<paragraph id="4E9F20A7DAFCBD2E85786632A4801E99" pageId="0" pageNumber="35">Type locality.</paragraph>
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<paragraph id="818651CA161CE1109641465E1AD78325" pageId="0" pageNumber="35">
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<emphasis id="3057C9DC90211E687463BC157F947199" bold="true" pageId="0" pageNumber="35">Panama: Chiriqui Province</emphasis>
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: Cuenca 108; Boquete District;
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<normalizedToken id="7B24DEE09D06F0F32CB97E86F4E3F908" originalValue="Volcán">Volcan</normalizedToken>
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<normalizedToken id="C7016D3E3F8A5F66563FAADE0742F194" originalValue="Barú">Baru</normalizedToken>
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NP;
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<normalizedToken id="8203A7B6716AA5B042A21693E04BA7FE" originalValue="Río">Rio</normalizedToken>
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del Guayabo; PSPSCB-PNVB-C108-2017-018;
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<geoCoordinate id="478CF4477D34F55C27FC66CD65C5B3AD" degrees="8.84939" direction="north" orientation="latitude" precision="1" value="8.84939">8.84939°N</geoCoordinate>
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,
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<geoCoordinate id="017E16DCA237BFD4F4D4E06B687A4EC8" degrees="82.49349" direction="west" orientation="longitude" precision="1" value="-82.49349">82.49349°W</geoCoordinate>
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; 1947 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="B82C70B1528B22E4DBD4240C91E48309" pageId="0" pageNumber="35" type="type specimen">
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<paragraph id="AE900A3E533BD92D7AE39E763880152E" pageId="0" pageNumber="35">Type specimen.</paragraph>
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<paragraph id="E5F69CFCEF3641AECD9649AC2B07C2E8" pageId="0" pageNumber="35">
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<emphasis id="A5EBBCC32F0C3A8CC7F7B8D9C2A0AA14" bold="true" pageId="0" pageNumber="35">
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<emphasis id="661407FA6E1CF1547E3248402146E331" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
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: male, Panama: Chiriqui Province
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</emphasis>
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: Cuenca 108; Boquete District;
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<normalizedToken id="D7CB40F7E4B2003A1474EE662BAEB5C2" originalValue="Volcán">Volcan</normalizedToken>
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<normalizedToken id="425AD955BB4E2A49136A56DD709A38DE" originalValue="Barú">Baru</normalizedToken>
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NP;
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<normalizedToken id="3C89EC0B46DAD6F3CB021AC90DA9D715" originalValue="Río">Rio</normalizedToken>
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del Guayabo; PSPSCB-PNVB-C108-2017-018;
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<geoCoordinate id="474705EB5B310DC10958BA5BB021C814" degrees="8.84939" direction="north" orientation="latitude" precision="1" value="8.84939">8.84939°N</geoCoordinate>
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,
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<geoCoordinate id="110F55A189EDA46B6226EC5FD57266F1" degrees="82.49349" direction="west" orientation="longitude" precision="1" value="-82.49349">82.49349°W</geoCoordinate>
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; 1947 m a.s.l.; Malaise trap; E.
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<normalizedToken id="1F296B213D779D01BB2432300A6A319E" originalValue="Álvarez">Alvarez</normalizedToken>
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, E.
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<normalizedToken id="FF529107793616C72DD4B8CB71AA6742" originalValue="Pérez">Perez</normalizedToken>
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, T.
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<normalizedToken id="07617141CCB9630E5C30D1B370D93359" originalValue="Ríos">Rios</normalizedToken>
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; 5-8.vi.2017; MIUP-008-T-2023 (in alcohol).
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</paragraph>
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</subSubSection>
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<subSubSection id="B8CAC1BEF16E978700F9587E1A87342D" pageId="0" pageNumber="35" type="diagnosis">
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<paragraph id="FF14F2175A072595DF31E9260C4EDD4B" pageId="0" pageNumber="35">Diagnosis.</paragraph>
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<paragraph id="3016DE0E1261E8C84DB897E417FFED02" pageId="0" pageNumber="35">
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Similar in many respects to both
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<taxonomicName id="48F5499F488DC10C65299AB0A1C6F987" lsidName="M. pakitza" pageId="0" pageNumber="35" rank="species" species="pakitza">
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<emphasis id="8C1B2E3E3E3D5544F5BB06C322BBB5F8" italics="true" pageId="0" pageNumber="35">M. pakitza</emphasis>
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</taxonomicName>
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Flint & Bueno-Soria and
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<taxonomicName id="2A3F602D892026EFBD0B4BC5A915C44C" lsidName="M. diosa" pageId="0" pageNumber="35" rank="species" species="diosa">
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<emphasis id="7738F6E23A85B0E9ED9F7805D6543514" italics="true" pageId="0" pageNumber="35">M. diosa</emphasis>
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</taxonomicName>
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Flint & Bueno-Soria, both from Peru,
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<taxonomicName id="C2BED568223C8FB7170EBF1D26B84A5A" lsidName="M. cascada" pageId="0" pageNumber="35" rank="species" species="cascada">
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<emphasis id="6A3D68C1221E5CB921C7D18C7CF39E5F" italics="true" pageId="0" pageNumber="35">M. cascada</emphasis>
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</taxonomicName>
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sp. nov. is distinguished from these species by the sharply narrowed posterior of the inferior appendage in lateral view, which is also deeply incised posteroventrally. There is also a spinose projection from the basal anterior margin and a knob-like process from the posterior margin of the inferior appendage when viewed in dorsal or ventral views in
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<taxonomicName id="C6818120BA2FB6E8197C577B4D43DE69" lsidName="M. cascada" pageId="0" pageNumber="35" rank="species" species="cascada">
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<emphasis id="BD7EF1BBC0C3B770CA5B177FDE4C0DD4" italics="true" pageId="0" pageNumber="35">M. cascada</emphasis>
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</taxonomicName>
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sp. nov., which is not seen in the related species.
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</paragraph>
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<caption id="D03CFF5E7044653D3BC16787EEF5995B" doi="10.3897/zookeys.1174.107314.figure10" httpUri="https://binary.pensoft.net/fig/890135" pageId="0" pageNumber="35" start="Figure 10" startId="F10">
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<paragraph id="A40D3570A2405BD6191908F390EBF6AE" pageId="0" pageNumber="35">
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<emphasis id="1FE12443EF57255925027AAE3E090678" bold="true" pageId="0" pageNumber="35">Figure 10.</emphasis>
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<taxonomicName id="3E2A206CC3EC4DB8C86103280952ED79" authorityName="Harris & Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Metrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrichia cascada" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cascada">
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<emphasis id="932C5CD88FDBC1097236F83AA8D0FC12" italics="true" pageId="0" pageNumber="35">Metrichia cascada</emphasis>
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</taxonomicName>
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sp. nov., male holotype, genitalia
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<emphasis id="2E825B85DDF34D72DC117F20388B061B" bold="true" pageId="0" pageNumber="35">A</emphasis>
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left lateral
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<emphasis id="28A9343F8B2487D4F740ADCD3E12E326" bold="true" pageId="0" pageNumber="35">B</emphasis>
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dorsal
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<emphasis id="409191DBDE2CD469A5EA8ABDD230B543" bold="true" pageId="0" pageNumber="35">C</emphasis>
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ventral
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<emphasis id="AB6E3220473C50BB9357C3163A01D3F6" bold="true" pageId="0" pageNumber="35">D</emphasis>
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phallus, dorsal
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<emphasis id="4CA30C0B3ED512E00F4CA5217D4A7D51" bold="true" pageId="0" pageNumber="35">E</emphasis>
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phallus apex, left lateral.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="095C8CA680C66AD0D277F04160E7511A" pageId="0" pageNumber="35" type="description">
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<paragraph id="51BB30A506F55079EBEB95363AAF88BE" pageId="0" pageNumber="35">Description.</paragraph>
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<paragraph id="7041CF33BD9C6ED088D95F053A2AED74" pageId="0" pageNumber="35">
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<emphasis id="DB68549EA7F7503C67214B6E12B2A609" bold="true" pageId="0" pageNumber="35">Male.</emphasis>
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Total length 1.9 mm, antenna short with 17 segments, middle segments flattened and widening, wings and body brown in alcohol, abdominal terga without modifications.
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<emphasis id="987E1DAB51A88D258A89AFD82CCDD641" bold="true" pageId="0" pageNumber="35">
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G
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<emphasis id="BA4F9ECBF64081823B942600A5F4B1CC" italics="true" pageId="0" pageNumber="35">enitalia</emphasis>
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.
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</emphasis>
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Abdominal segment VII annular without ventromesal process. Segment VIII triangular, tapering ventrally; in ventral view very short, broad emargination posteriorly; in dorsal view truncate. Segment IX triangular, tapering anteriorly into segment VII, posterior margin sinuate; in ventral view slightly incised mesally on posterior margin, anterior margin rounded; narrow in dorsal view. Preanal appendage (cercus) ovate laterally and dorsally. Dorsolateral hook in lateral view narrow over length, tapering apically and curving ventrad; in dorsal view narrow over length, apices rounded and tapering inward. Segment X lobate in lateral view, rounded distally; in dorsal view broadly triangular, apex truncate and membranous. Inferior appendage in lateral view wide basally and rounded, tapering distally, incised posteroventrally, upper portion long and pointed apically, lower portion short and acute, process with numerous dark peg-like setae from basal inner margin; in ventral view wide basally, tapering distally, subapical knob on inner margin; in dorsal view wide basally, tapering distally, subapical spike on inner margin, renal-shaped process bearing short pegs on inner margin near base. Phallus in dorsal view wide basally, narrowing apically, distal portion with pair of spines, subapical spine larger than apical spine, and straight, apical spine short and curved; in lateral view, subapical spine twice as long and thicker than apical spine, curving slightly under phallic shaft, apical spine short and dorsal in position.
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</paragraph>
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</subSubSection>
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<subSubSection id="29278486BDA9FAAF6E4FA1E56F03152B" pageId="0" pageNumber="35" type="distribution">
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<paragraph id="2D2804C1C96D82E46E9C00C7CB04EF13" pageId="0" pageNumber="35">Distribution.</paragraph>
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<paragraph id="A71CAAC594156C1916C3D71A491FC523" pageId="0" pageNumber="35">Panama.</paragraph>
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</subSubSection>
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<subSubSection id="7F28821D51158030E45341CF15D366FA" pageId="0" pageNumber="35" type="etymology">
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<paragraph id="940F26C4910648ACF6A0056328CF7304" pageId="0" pageNumber="35">Etymology.</paragraph>
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<paragraph id="60E9EB286B24345D6161537865FDA88E" pageId="0" pageNumber="35">
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The species name
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<taxonomicName id="80BDFBEB88D0988BCF753D49D8E681EF" authorityName="Harris & Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Metrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrichia cascada" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cascada">
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<emphasis id="A281F6FCD2457DAC4544260541AF22F2" italics="true" pageId="0" pageNumber="35">Metrichia cascada</emphasis>
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</taxonomicName>
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(cascade or waterfall) derives from Spanish, referring to the cascade on the
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<normalizedToken id="8FBBDBDC5517E8954C4C63ECEA17F012" originalValue="Río">Rio</normalizedToken>
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del Guayabo where the species was collected.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |