treatments-xml/data/AE/04/87/AE0487C8F538FFD30986A25662C2FBB9.xml
2024-06-21 12:47:47 +02:00

187 lines
24 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F538FFD30986A25662C2FBB9" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus alluaudi Burgeon 1937" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="12" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Serrano, Artur R. M.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes &amp; CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5256</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>35</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
<mods:identifier type="ZooBank">CE0AC7CD-E55D-484C-9346-86F16C985DBE</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:AE0487C8F538FFD30986A25662C2FBB9" httpUri="http://treatment.plazi.org/id/AE0487C8F538FFD30986A25662C2FBB9" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection box="[151,614,330,357]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph blockId="11.[151,614,330,393]" box="[151,614,330,357]" pageId="11" pageNumber="12">
<heading bold="true" box="[151,614,330,357]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName ID-CoL="72SB6" authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[151,614,330,357]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis bold="true" box="[151,614,330,357]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[151,426,330,357]" italics="true" pageId="11" pageNumber="12">Macrocheilus alluaudi</emphasis>
<bibRefCitation author="Burgeon, L." box="[433,614,330,357]" pageId="11" pageNumber="12" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,361,367,393]" pageId="11" pageNumber="12" type="description">
<paragraph blockId="11.[151,614,330,393]" box="[151,361,367,393]" pageId="11" pageNumber="12">
<figureCitation box="[151,280,367,393]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="11" pageNumber="12">
Figs. 1f,
<quantity box="[251,280,367,393]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="11" pageNumber="12" unit="g" value="1.0">1g</quantity>
</figureCitation>
,
<figureCitation box="[293,322,367,393]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="11" pageNumber="12">2e</figureCitation>
,
<figureCitation box="[333,361,367,393]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="11" pageNumber="12">3d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[151,1437,438,1041]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[151,360,438,464]" pageId="11" pageNumber="12">Studied material.</emphasis>
<materialsCitation collectingDate="2017-11-28" collectorName="Bateke Plateau H. Krahnstover &amp; Collection P. Sch &amp; Herrenberg" country="Republic of the Congo" determinerName="A. Serrano" location="Reserve Naturelle des Gorilles" municipality="Lesio Louna" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1">
<specimenCount box="[369,451,439,465]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
: “
<collectingCountry box="[473,692,439,465]" name="Republic of the Congo" pageId="11" pageNumber="12">Republic of Congo</collectingCountry>
\
<location LSID="urn:lsid:plazi:treatment:AE0487C8F538FFD30986A25662C2FBB9:23726005F538FFD30BD4A2AB6509FE79" box="[709,1058,439,465]" country="Republic of the Congo" municipality="Lesio Louna" name="Reserve Naturelle des Gorilles" pageId="11" pageNumber="12">Réserve Naturelle des Gorilles</location>
de\
<collectingMunicipality box="[1110,1253,439,465]" pageId="11" pageNumber="12">Lésio Louna</collectingMunicipality>
17
<date box="[1300,1428,439,465]" pageId="11" pageNumber="12" value="2017-11-28">
<collectingDate box="[1300,1428,439,465]" pageId="11" pageNumber="12" value="2017-11-28">28.11.2017</collectingDate>
</date>
\
<collectorName box="[151,496,475,501]" pageId="11" pageNumber="12">Batéké Plateau H. Krahnstôver</collectorName>
leg.” (
<collectorName box="[571,771,475,501]" pageId="11" pageNumber="12">Collection P. Sch</collectorName>
̧le,
<collectorName box="[812,940,475,501]" pageId="11" pageNumber="12">Herrenberg</collectorName>
)\\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[983,1136,475,501]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[983,1136,475,501]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Burgeon" authorityYear="1937" box="[1150,1243,475,501]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis box="[1150,1243,475,501]" italics="true" pageId="11" pageNumber="12">alluaudi</emphasis>
</taxonomicName>
\
<bibRefCitation author="Burgeon, L." box="[1258,1426,475,501]" pageId="11" pageNumber="12" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
” [h]\ “
<determinerName box="[209,325,511,537]" pageId="11" pageNumber="12">A.Serrano</determinerName>
det. 2021” [t] [h]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="discussion">
<paragraph blockId="11.[151,1437,438,1041]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,315,547,573]" pageId="11" pageNumber="12">Remarks.</emphasis>
The species was described as a variety of
<taxonomicName authority="(Dejean, 1831)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[786,1092,547,573]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="labrosus">
<emphasis box="[786,918,547,573]" italics="true" pageId="11" pageNumber="12">M. labrosus</emphasis>
(Dejean, 1831)
</taxonomicName>
and later it was elevated to the subspecies level by
<bibRefCitation author="Basilewsky, P." box="[375,586,583,609]" pageId="11" pageNumber="12" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky (1960)</bibRefCitation>
due to its well differentiated external morphological characters, such as the absence of a big yellow spot in each elytron (
<figureCitation box="[647,773,619,645]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="11" pageNumber="12">
Figs. 1f,
<quantity box="[745,773,619,645]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="11" pageNumber="12" unit="g" value="1.0">1g</quantity>
</figureCitation>
) and less dense punctuation on the head and pronotum. Lat- er, it was considered a distinct species without any explanation for the change in taxonomic category (
<bibRefCitation author="Lorenz, W." box="[1285,1431,654,681]" pageId="11" pageNumber="12" refId="ref25636" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world. (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
;
<bibRefCitation author="Hackel, M. &amp; Farkac, J." box="[151,409,691,717]" pageId="11" pageNumber="12" pagination="261 - 384" refId="ref25548" refString="Hackel, M. &amp; Farkac, J. (2013) A checklist of the subfamily Anthiinae Bonelli, 1813 of the World (Coleoptera: Carabidae). Studies and Reports. Taxonomical Series, 9 (2), 261 - 384." type="journal article" year="2013">Häckel &amp; Farkač 2013</bibRefCitation>
). Taking into account the abovementioned external morphological differences, in addition to the slight different shapes of the labrum and of the median lobe of both forms (cf.
<figureCitation box="[1067,1158,727,753]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="11" pageNumber="12">Figs. 2e</figureCitation>
,
<figureCitation box="[1169,1197,727,753]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="11" pageNumber="12">3d</figureCitation>
<emphasis box="[1204,1227,727,752]" italics="true" pageId="11" pageNumber="12">vs</emphasis>
6e, 7c), we follow the taxonomic treatment given by
<bibRefCitation author="Lorenz, W." box="[532,692,762,789]" pageId="11" pageNumber="12" refId="ref25636" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world. (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005)</bibRefCitation>
and
<bibRefCitation author="Hackel, M. &amp; Farkac, J." box="[745,1020,763,789]" pageId="11" pageNumber="12" pagination="261 - 384" refId="ref25548" refString="Hackel, M. &amp; Farkac, J. (2013) A checklist of the subfamily Anthiinae Bonelli, 1813 of the World (Coleoptera: Carabidae). Studies and Reports. Taxonomical Series, 9 (2), 261 - 384." type="journal article" year="2013">Häckel &amp; Farkač (2013)</bibRefCitation>
. Moreover, the pronotum hind angles in
<taxonomicName authorityName="Burgeon" authorityYear="1937" box="[181,313,799,825]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis box="[181,313,799,825]" italics="true" pageId="11" pageNumber="12">M. alluaudi</emphasis>
</taxonomicName>
are rounded while in
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[565,698,799,825]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="labrosus">
<emphasis box="[565,698,799,825]" italics="true" pageId="11" pageNumber="12">M. labrosus</emphasis>
</taxonomicName>
are dentiform (cf.
<figureCitation box="[914,1045,799,825]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="11" pageNumber="12">
Figs. 1f,
<quantity box="[1017,1045,799,825]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="11" pageNumber="12" unit="g" value="1.0">1g</quantity>
</figureCitation>
<emphasis box="[1053,1076,799,824]" italics="true" pageId="11" pageNumber="12">vs</emphasis>
<figureCitation box="[1084,1161,798,825]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="11" pageNumber="12">Fig. 5i</figureCitation>
). In addition, the elytral intervals of the former species have a thin polygonal microreticulation not visible in the latter species. The species is known from
<collectingCountry box="[322,714,871,897]" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">Democratic Republic of the Congo</collectingCountry>
(
<bibRefCitation author="Basilewsky, P." box="[728,923,871,897]" pageId="11" pageNumber="12" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky 1960</bibRefCitation>
) and
<collectingCountry box="[986,1240,871,897]" name="Republic of the Congo" pageId="11" pageNumber="12">Republic of the Congo</collectingCountry>
(
<bibRefCitation author="Lorenz, W. &amp; Banki, Y. &amp; Roskov, M. &amp; Doring, G. &amp; Ower, L. &amp; Vandepitte, D. &amp; Hobern, D. &amp; Remsen, P. &amp; Schalk, R. E. &amp; DeWalt, M. &amp; Keping, J. &amp; Miller, T. &amp; Orrell, R. &amp; Aalbu, R. &amp; Adlard, E. M. &amp; Adriaenssens, C. &amp; Aedo, E. &amp; Aescht, N. &amp; Akkari, S." box="[1254,1403,871,897]" pageId="11" pageNumber="12" refId="ref25680" refString="Lorenz, W. (2021) Carabcat database. In: O. Banki, Y. Roskov, M. Doring, G. Ower, L. Vandepitte, D. Hobern, D. Remsen, P. Schalk, R. E. DeWalt, M. Keping, J. Miller, T. Orrell, R. Aalbu, R. Adlard, E. M. Adriaenssens, C. Aedo, E. Aescht, N. Akkari, S. Alexander, et al., Catalogue of Life Checklist. Version 3. August 2021. https: // doi. org / 10.48580 / dfqf- 3 dk" type="book" year="2021">Lorenz 2021</bibRefCitation>
).
</paragraph>
<paragraph blockId="11.[151,1437,438,1041]" pageId="11" pageNumber="12">
<emphasis box="[199,364,907,933]" italics="true" pageId="11" pageNumber="12">Male genitalia</emphasis>
(
<figureCitation box="[379,463,907,933]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="11" pageNumber="12">Fig. 3d</figureCitation>
). Median lobe very slightly different from that of
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1023,1156,907,933]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="labrosus">
<emphasis box="[1023,1156,907,933]" italics="true" pageId="11" pageNumber="12">M. labrosus</emphasis>
</taxonomicName>
, having a lesser rounded apex and the stripes are membranous in its inner sides; the left paramere is very similar in both species (cf.
<figureCitation box="[1346,1436,943,969]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="11" pageNumber="12">Figs. 3d</figureCitation>
and
<figureCitation box="[199,228,979,1005]" captionStart="FIGURE 7" captionStartId="19.[152,255,1531,1555]" captionTargetBox="[163,1424,429,1485]" captionTargetId="figure-63@19.[151,1436,421,1492]" captionTargetPageId="19" captionText="FIGURE 7.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. dorsalis, ZAM (TAC); b, M. elegantulus, DOC? (MRAC); c, M. labrosus, RCA (PSC); d, M. longicollis, MOZ (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745297" httpUri="https://zenodo.org/record/7745297/files/figure.png" pageId="11" pageNumber="12">7c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,1001,1014,1041]" pageId="11" pageNumber="12" type="distribution">
<paragraph blockId="11.[151,1437,438,1041]" box="[199,1001,1014,1041]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,353,1014,1040]" pageId="11" pageNumber="12">Distribution.</emphasis>
<collectingCountry box="[359,590,1015,1041]" name="Republic of the Congo" pageId="11" pageNumber="12">Congo (Brazzaville)</collectingCountry>
,
<collectingCountry box="[601,997,1015,1041]" name="Democratic Republic of the Congo" pageId="11" pageNumber="12">Democratic Republic of the Congo</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>