498 lines
61 KiB
XML
498 lines
61 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F537FFDE0986A64263CEFAFD" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus philippeorum Serrano 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="5" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Serrano, Artur R. M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
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<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
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<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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||
<mods:date>2023</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2023-03-16</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>5256</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>35</mods:end>
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||
</mods:extent>
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||
</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
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<mods:identifier type="ZooBank">CE0AC7CD-E55D-484C-9346-86F16C985DBE</mods:identifier>
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</mods:mods>
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||
<treatment LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD" httpUri="http://treatment.plazi.org/id/AE0487C8F537FFDE0986A64263CEFAFD" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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||
<subSubSection box="[151,562,1374,1401]" pageId="4" pageNumber="5" type="nomenclature">
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||
<paragraph blockId="4.[151,562,1374,1437]" box="[151,562,1374,1401]" pageId="4" pageNumber="5">
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||
<heading bold="true" box="[151,562,1374,1401]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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||
<emphasis bold="true" box="[151,562,1374,1401]" pageId="4" pageNumber="5">
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||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[151,488,1374,1401]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
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<emphasis bold="true" box="[151,488,1374,1401]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[495,562,1375,1401]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,364,1411,1437]" pageId="4" pageNumber="5" type="description">
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||
<paragraph blockId="4.[151,562,1374,1437]" box="[151,364,1411,1437]" pageId="4" pageNumber="5">
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||
<figureCitation box="[151,241,1411,1437]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="4" pageNumber="5">Figs. 1c</figureCitation>
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,
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<figureCitation box="[254,282,1411,1437]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="4" pageNumber="5">2c</figureCitation>
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,
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<figureCitation box="[294,324,1411,1437]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="4" pageNumber="5">3b</figureCitation>
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,
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<figureCitation box="[336,364,1411,1437]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="4" pageNumber="5">4b</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="5" type="materials_examined">
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||
<paragraph blockId="4.[151,1437,1482,2013]" pageId="4" pageNumber="5">
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||
<emphasis bold="true" box="[151,324,1482,1509]" pageId="4" pageNumber="5">Type material.</emphasis>
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||
<materialsCitation collectingDate="2005-08-24" collectingDateMax="2005-08-28" collectingDateMin="2005-08-24" collectorName="Phillippe Moretto" country="Burkina Faso" determinerName="A. Serrano" elevation="232" latitude="11.283334" location="Pama" longLatPrecision="21" longitude="0.7163889" pageId="4" pageNumber="5" specimenCount="2" typeStatus="holotype">
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||
<typeStatus box="[331,435,1483,1509]" pageId="4" pageNumber="5">Holotype</typeStatus>
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male: “
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||
<collectingCountry box="[520,675,1483,1509]" name="Burkina Faso" pageId="4" pageNumber="5">Burkina Faso</collectingCountry>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0BBFA6D763C7FA4D" box="[686,748,1483,1509]" country="Burkina Faso" latitude="11.283334" longLatPrecision="21" longitude="0.7163889" name="Pama" pageId="4" pageNumber="5">Pama</location>
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||
\
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||
<date box="[762,952,1482,1509]" pageId="4" pageNumber="5" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">
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||
<collectingDate box="[762,952,1482,1509]" pageId="4" pageNumber="5" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">24–28.VIII.2005</collectingDate>
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||
</date>
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||
,\
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||
<geoCoordinate box="[972,1124,1483,1511]" degrees="11" direction="north" minutes="17" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="00" value="11.283334">11º 17′ 00′′ N</geoCoordinate>
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||
,
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<geoCoordinate box="[1135,1246,1483,1511]" degrees="0" direction="east" minutes="42" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="59" value="0.7163889">0º 42′59E</geoCoordinate>
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||
,
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||
<quantity box="[1257,1320,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="4" pageNumber="5" unit="m" value="232.0">
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||
<elevation box="[1257,1320,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="4" pageNumber="5" unit="m" value="232.0">232m</elevation>
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||
</quantity>
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||
\
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||
<collectorName pageId="4" pageNumber="5">Phillippe Moretto</collectorName>
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leg.” \\ “Macr-12” (yellow label)\\ “
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||
<typeStatus box="[655,762,1519,1545]" pageId="4" pageNumber="5">Holotype</typeStatus>
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||
male\
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||
<determinerName box="[839,962,1519,1545]" pageId="4" pageNumber="5">A. Serrano</determinerName>
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||
det. 2022” [t] [h] [red card]
|
||
</materialsCitation>
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||
.
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||
<materialsCitation collectingDate="2005-08-24" collectingDateMax="2005-08-28" collectingDateMin="2005-08-24" collectorName="Phillippe Moretto" country="Burkina Faso" elevation="232" location="Pama" longitude="0.7163889" pageId="4" pageNumber="5" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
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||
<typeStatus box="[1297,1410,1519,1545]" pageId="4" pageNumber="5">Paratypes</typeStatus>
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||
:
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||
<specimenCount pageId="4" pageNumber="5" type="male">3 males</specimenCount>
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||
: “
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||
<collectingCountry box="[242,398,1555,1581]" name="Burkina Faso" pageId="4" pageNumber="5">Burkina Faso</collectingCountry>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0888A50F60FCF985" box="[409,471,1555,1581]" country="Burkina Faso" longitude="0.7163889" name="Pama" pageId="4" pageNumber="5">Pama</location>
|
||
\
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||
<date box="[487,676,1554,1581]" pageId="4" pageNumber="5" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">
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||
<collectingDate box="[487,676,1554,1581]" pageId="4" pageNumber="5" value="2005-08-24" valueMax="2005-08-28" valueMin="2005-08-24">24–28.VIII.2005</collectingDate>
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||
</date>
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||
,\ 11º 17′ 00″′N,
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||
<geoCoordinate box="[862,975,1555,1583]" degrees="0" direction="east" minutes="42" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="59" value="0.7163889">0º 42′59E</geoCoordinate>
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||
,
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||
<quantity box="[986,1050,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="4" pageNumber="5" unit="m" value="232.0">
|
||
<elevation box="[986,1050,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="4" pageNumber="5" unit="m" value="232.0">232m</elevation>
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||
</quantity>
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||
\
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<collectorName box="[1065,1264,1555,1581]" pageId="4" pageNumber="5">Phillippe Moretto</collectorName>
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leg.” \\ “Macr- 13”\\ “Macr-14”, “Macr-15” (yellow labels)\\
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||
</materialsCitation>
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||
“
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<materialsCitation collectingDate="2001-07-24" collectorName="A. Kudrna Jr." country="Benin" determinerName="A. Serrano" location="Bensekou" pageId="4" pageNumber="5" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus box="[683,784,1591,1617]" pageId="4" pageNumber="5">Paratype</typeStatus>
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male\
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<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[863,1175,1591,1617]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
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<emphasis box="[863,1175,1591,1617]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1183,1247,1591,1617]" pageId="4" pageNumber="5">
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<taxonomicNameLabel box="[1183,1247,1591,1617]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
\
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<determinerName box="[1266,1387,1591,1617]" pageId="4" pageNumber="5">A. Serrano</determinerName>
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det. 2022” [t] [h] [red card];
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<specimenCount box="[436,541,1627,1653]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
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: “NE
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<collectingCountry box="[612,678,1627,1653]" name="Benin" pageId="4" pageNumber="5">Benin</collectingCountry>
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\
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<locationDeviation box="[696,824,1627,1653]" location="Kandi" pageId="4" pageNumber="5">E of Kandi</locationDeviation>
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\
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<quantity box="[841,901,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="km" value="7.0">7 km</quantity>
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||
E of
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<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0AD8A547651CF9DD" box="[969,1079,1627,1653]" country="Benin" name="Bensekou" pageId="4" pageNumber="5">Bensekou</location>
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\ 23–
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<date box="[1138,1265,1627,1653]" pageId="4" pageNumber="5" value="2001-07-24">
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<collectingDate box="[1138,1265,1627,1653]" pageId="4" pageNumber="5" value="2001-07-24">24.07.2001</collectingDate>
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</date>
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\
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<collectorName box="[1281,1437,1627,1653]" pageId="4" pageNumber="5">A. Kudrna Jr.</collectorName>
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LGT” \\ “Macr-16” (yellow label)\\
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</materialsCitation>
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“
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<materialsCitation collectingDate="2001-06-25" collectorName="A. Kudrna Lgt. & Th. Assmann" country="Benin" determinerName="A. Serrano" location="Bensekou" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<typeStatus box="[564,665,1663,1689]" pageId="4" pageNumber="5">Paratype</typeStatus>
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||
female\
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<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[762,1073,1663,1689]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
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||
<emphasis box="[762,1073,1663,1689]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
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||
</taxonomicName>
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||
<emphasis bold="true" box="[1080,1143,1663,1689]" pageId="4" pageNumber="5">
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||
<taxonomicNameLabel box="[1080,1143,1663,1689]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
|
||
\
|
||
<determinerName box="[1160,1279,1663,1689]" pageId="4" pageNumber="5">A. Serrano</determinerName>
|
||
det. 2022” [t] [h] [red card];
|
||
<specimenCount box="[317,398,1699,1725]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
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: “NE
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<collectingCountry box="[466,532,1699,1725]" name="Benin" pageId="4" pageNumber="5">Benin</collectingCountry>
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||
\
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||
<locationDeviation box="[549,673,1699,1725]" location="Kandi" pageId="4" pageNumber="5">E of Kandi</locationDeviation>
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||
\
|
||
<quantity box="[688,746,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="5" unit="km" value="4.0">4 km</quantity>
|
||
W of
|
||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0A23A5BF628BF915" box="[818,928,1699,1725]" country="Benin" name="Bensekou" pageId="4" pageNumber="5">Bensekou</location>
|
||
\ 24–
|
||
<date box="[985,1111,1698,1725]" pageId="4" pageNumber="5" value="2001-06-25">
|
||
<collectingDate box="[985,1111,1698,1725]" pageId="4" pageNumber="5" value="2001-06-25">25.06.2001</collectingDate>
|
||
</date>
|
||
\
|
||
<collectorName box="[1126,1300,1699,1725]" pageId="4" pageNumber="5">A. Kudrna Lgt.</collectorName>
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||
” \\ “Collection\
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||
<collectorName box="[211,361,1735,1761]" pageId="4" pageNumber="5">Th. Assmann</collectorName>
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||
\ Bleckede, CAB” (yellow label)\\
|
||
</materialsCitation>
|
||
“
|
||
<materialsCitation collectingDate="2008-06-18" country="Senegal" county="Nianing" determinerName="A. Serrano" location="Afrique" municipality="Petit Cote" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus box="[769,870,1735,1761]" pageId="4" pageNumber="5">Paratype</typeStatus>
|
||
male\
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[950,1263,1735,1761]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
|
||
<emphasis box="[950,1263,1735,1761]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1272,1337,1735,1761]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel box="[1272,1337,1735,1761]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
\
|
||
<determinerName pageId="4" pageNumber="5">A. Serrano</determinerName>
|
||
det. 2022” [t] [h] [red card];
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||
<specimenCount box="[537,618,1771,1797]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
|
||
: “
|
||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0B6EA5F763F2F8AD" box="[639,729,1771,1797]" country="Senegal" county="Nianing" municipality="Petit Cote" name="Afrique" pageId="4" pageNumber="5">Afrique</location>
|
||
de L`Ouest\
|
||
<collectingCountry box="[872,960,1771,1797]" name="Senegal" pageId="4" pageNumber="5">Senegal</collectingCountry>
|
||
<collectingMunicipality box="[968,1081,1771,1797]" pageId="4" pageNumber="5">Petit Cote</collectingMunicipality>
|
||
\
|
||
<collectingCounty box="[1089,1179,1771,1797]" pageId="4" pageNumber="5">Nianing</collectingCounty>
|
||
<date box="[1187,1314,1771,1797]" pageId="4" pageNumber="5" value="2008-06-18">
|
||
<collectingDate box="[1187,1314,1771,1797]" pageId="4" pageNumber="5" value="2008-06-18">18.VI.2008</collectingDate>
|
||
</date>
|
||
IRD\Ultra Violet\Alain Coache\\ “
|
||
<typeStatus box="[415,516,1807,1833]" pageId="4" pageNumber="5">Paratype</typeStatus>
|
||
male\
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[594,906,1807,1833]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
|
||
<emphasis box="[594,906,1807,1833]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[914,978,1807,1833]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel box="[914,978,1807,1833]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
\
|
||
<determinerName box="[996,1116,1807,1833]" pageId="4" pageNumber="5">A. Serrano</determinerName>
|
||
det. 2022” [t] [h] [red card]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation box="[156,1336,1843,1869]" collectingDate="2009-09-20" country="Senegal" county="Mbour" location="Afrique" municipality="Petit Cote" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount box="[156,231,1843,1869]" pageId="4" pageNumber="5" type="male">1male</specimenCount>
|
||
: “
|
||
<location LSID="urn:lsid:plazi:treatment:AE0487C8F537FFDE0986A64263CEFAFD:23726005F537FFDC0811A42F6071F8E5" box="[256,346,1843,1869]" country="Senegal" county="Mbour" municipality="Petit Cote" name="Afrique" pageId="4" pageNumber="5">Afrique</location>
|
||
de L`Ouest\
|
||
<collectingCountry box="[494,582,1843,1869]" name="Senegal" pageId="4" pageNumber="5">Senegal</collectingCountry>
|
||
<collectingMunicipality box="[593,708,1843,1869]" pageId="4" pageNumber="5">Petit Cote</collectingMunicipality>
|
||
\
|
||
<collectingCounty box="[716,792,1843,1869]" pageId="4" pageNumber="5">Mbour</collectingCounty>
|
||
<date box="[803,931,1843,1869]" pageId="4" pageNumber="5" value="2009-09-20">
|
||
<collectingDate box="[803,931,1843,1869]" pageId="4" pageNumber="5" value="2009-09-20">20.IX.2009</collectingDate>
|
||
</date>
|
||
IRD\Ultra Violet\Alain Coache\\ “
|
||
</materialsCitation>
|
||
<typeStatus box="[1336,1437,1843,1869]" pageId="4" pageNumber="5">Paratype</typeStatus>
|
||
male\
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[220,531,1879,1905]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
|
||
<emphasis box="[220,531,1879,1905]" italics="true" pageId="4" pageNumber="5">Macrocheilus philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[537,600,1879,1905]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel box="[537,600,1879,1905]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
\A. Serrano det. 2022” [t] [h] [red card].
|
||
<materialsCitation box="[1060,1367,1879,1905]" collectionCode="PSC" country="Senegal" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus box="[1060,1164,1879,1905]" pageId="4" pageNumber="5">Holotype</typeStatus>
|
||
deposited in
|
||
<collectionCode box="[1314,1367,1879,1905]" pageId="4" pageNumber="5">PSC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectionCode="ACC, ASC, PSC, TAC" country="Senegal" pageId="4" pageNumber="5" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus pageId="4" pageNumber="5">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode box="[359,419,1915,1941]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13787" name="Oak Hill Park Museum" pageId="4" pageNumber="5" type="Herbarium">ACC</collectionCode>
|
||
,
|
||
<collectionCode box="[429,486,1915,1941]" country="USA" lsid="urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="4" pageNumber="5" type="Herbarium">ASC</collectionCode>
|
||
,
|
||
<collectionCode box="[497,547,1915,1941]" pageId="4" pageNumber="5">PSC</collectionCode>
|
||
and
|
||
<collectionCode box="[601,658,1915,1941]" country="USA" lsid="urn:lsid:biocol.org:col:14176" name="Tarleton State University, Biological Sciences Department" pageId="4" pageNumber="5" type="Herbarium">TAC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph blockId="4.[151,1437,1482,2013]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<emphasis bold="true" box="[199,465,1950,1976]" pageId="4" pageNumber="5">Differential diagnosis.</emphasis>
|
||
Head, including mouthparts, antennae, pronotum, scutellum, legs, and abdomen orange-brownish or reddish-brown; elytra fuscous black or dark brown, lateral and apical regions orange-brownish, a pair of large longitudinal humeral orange bands and a sub transverse one of the same colour in the last third; shiny and pubescent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
|
||
<paragraph blockId="5.[151,1437,151,2013]" box="[199,1407,222,249]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[199,345,222,248]" pageId="5" pageNumber="6">Description.</emphasis>
|
||
Length of
|
||
<typeStatus box="[468,570,223,249]" pageId="5" pageNumber="6">holotype</typeStatus>
|
||
:
|
||
<quantity box="[581,683,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="5" pageNumber="6" unit="mm" value="11.4">11.4 mm</quantity>
|
||
. Length of
|
||
<typeStatus box="[810,922,223,249]" pageId="5" pageNumber="6">paratypes</typeStatus>
|
||
:
|
||
<quantity box="[932,1094,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" metricValueMax="1.29" metricValueMin="1.13" pageId="5" pageNumber="6" unit="mm" value="12.1" valueMax="12.9" valueMin="11.3">11.3–12.9 mm</quantity>
|
||
(males),
|
||
<quantity box="[1199,1298,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="6" unit="mm" value="11.5">11.5 mm</quantity>
|
||
(female).
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
|
||
<emphasis box="[199,260,259,285]" italics="true" pageId="5" pageNumber="6">Head</emphasis>
|
||
(
|
||
<figureCitation box="[277,358,259,285]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="5" pageNumber="6">Fig. 1c</figureCitation>
|
||
). Slightly wider than long (males: length
|
||
<quantity box="[835,1001,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.1799999999999997" metricValueMax="2.34" metricValueMin="2.02" pageId="5" pageNumber="6" unit="mm" value="2.1799999999999997" valueMax="2.34" valueMin="2.02">2.02–2.34 mm</quantity>
|
||
, width:
|
||
<quantity box="[1094,1261,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" metricValueMax="2.82" metricValueMin="2.5" pageId="5" pageNumber="6" unit="mm" value="2.66" valueMax="2.82" valueMin="2.5">2.50–2.82 mm</quantity>
|
||
; female: length
|
||
<quantity box="[151,254,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="5" pageNumber="6" unit="mm" value="2.34">2.34 mm</quantity>
|
||
; width:
|
||
<quantity box="[344,447,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.56" pageId="5" pageNumber="6" unit="mm" value="2.56">2.56 mm</quantity>
|
||
). Surface shining, coarsely and deeply punctured, less dense in middle of the frons, each puncture with a long, decumbent and fine brown-yellowish seta, without microsculpture; antero-lateral impressions well marked, frons excavated behind clypeus; last maxillary palpomere securiform, pubescent; eyes prominent, temples short with rows of large setae along posterior margin of eyes. Clypeus slightly punctured, each puncture with a long, decumbent and fine brown-yellowish seta, anterior half with a row of four large and one to two minor submarginal setiferous punctures. Labrum smooth, orange and shiny, with a medium thin blackish band, 1.1–1.2 times wider than long (males: length
|
||
<quantity box="[662,828,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="11.2" metricValueMin="8.6" pageId="5" pageNumber="6" unit="mm" value="0.99" valueMax="1.12" valueMin="0.86">0.86–1.12 mm</quantity>
|
||
, width
|
||
<quantity box="[911,1077,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.22" metricValueMin="1.06" pageId="5" pageNumber="6" unit="mm" value="1.14" valueMax="1.22" valueMin="1.06">1.06–1.22 mm</quantity>
|
||
; female: length
|
||
<quantity box="[1258,1360,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="6" unit="mm" value="0.98">0.98 mm</quantity>
|
||
, width
|
||
<quantity box="[151,255,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="5" pageNumber="6" unit="mm" value="1.17">1.17 mm</quantity>
|
||
), posterior half subparallel-sided, anterior half almost triangular, lateral margins converging and the apex rounded; a large seta at the widest point near the rim, at the apex 4 setae, two exactly on the margin from the top, and two beneath the margin just on the top, not visible in dorsal view; posterior lateral half with a longitudinal groove directed inward and up to widest point with a fringe of short setae underneath (
|
||
<figureCitation box="[1058,1142,655,681]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="5" pageNumber="6">Fig. 2c</figureCitation>
|
||
). One pair of supraorbital setae on each side (
|
||
<figureCitation box="[366,446,691,717]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="5" pageNumber="6">Fig. 1c</figureCitation>
|
||
). Mentum with a central triangular tooth, slightly shorter than the side ones, with several setae near the base; on the tooth itself three bristles near the apex. Antennae with the first three antennomeres less densely pubescent with rather long setae, from the fourth antennomere densely pubescent with shorter setae, first and third antennomeres almost cylindrical, second and fourth sub-pyriform, the remaining depressed, slightly longer than wide, with a thin median longitudinal blackish smooth area; last antennomere more or less acuminate.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
|
||
<emphasis box="[199,280,871,897]" italics="true" pageId="5" pageNumber="6">Thorax</emphasis>
|
||
. Pronotum slightly cordiform (
|
||
<figureCitation box="[632,712,871,897]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="5" pageNumber="6">Fig. 1c</figureCitation>
|
||
), transverse, about 1.2–1.4 times wider than long (males: length
|
||
<quantity box="[151,317,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.095" metricValueMax="2.24" metricValueMin="1.95" pageId="5" pageNumber="6" unit="mm" value="2.095" valueMax="2.24" valueMin="1.95">1.95–2.24 mm</quantity>
|
||
, width:
|
||
<quantity box="[410,577,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.7199999999999998" metricValueMax="3.01" metricValueMin="2.43" pageId="5" pageNumber="6" unit="mm" value="2.7199999999999998" valueMax="3.01" valueMin="2.43">2.43–3.01 mm</quantity>
|
||
; female: length
|
||
<quantity box="[761,864,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" pageId="5" pageNumber="6" unit="mm" value="2.08">2.08 mm</quantity>
|
||
, width
|
||
<quantity box="[949,1054,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" pageId="5" pageNumber="6" unit="mm" value="2.66">2.66 mm</quantity>
|
||
), maximum width more or less at the middle length; surface covered with coarsely punctures, more densely in the anterior, lateral and posterior areas, bearing each one a long and fine brown-yellowish seta, intervals smooth; anterior angles rounded, anterior margin arcuate; posterior margin slightly arcuate in the middle, strongly oblique sinuate towards the posterior angles which are obtuse-sharped and bear a long seta; posterior margin not margined, anterior margin margined in the middle, lateral margins with complete and distinct margination from the anterior angles and reaching the posterior angles, slightly sinuate just anterior to posterior angles, forming a short tooth just before the angles, a long seta in front of the widest part of pronotum, anterior angles with some large bristles; anterior transverse sulcus not distinct, pronotal base with a pair of longitudinal faint foveae separated by a median sulcus in a flat and depressed area that reaches the posterior margin. Scutellum triangular, strongly setiferous punctured. Prosternum coarsely punctured, without microsculpture.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
|
||
<emphasis box="[199,269,1303,1329]" italics="true" pageId="5" pageNumber="6">Elytra</emphasis>
|
||
(
|
||
<figureCitation box="[286,366,1303,1329]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="5" pageNumber="6">Fig. 1c</figureCitation>
|
||
) 1.8–1.9 times longer than wide (males: length
|
||
<quantity box="[918,1085,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="7.59" metricValueMax="8.38" metricValueMin="6.8" pageId="5" pageNumber="6" unit="mm" value="7.59" valueMax="8.38" valueMin="6.8">6.80–8.38 mm</quantity>
|
||
, width
|
||
<quantity box="[1171,1339,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.115" metricValueMax="4.55" metricValueMin="3.68" pageId="5" pageNumber="6" unit="mm" value="4.115" valueMax="4.55" valueMin="3.68">3.68–4.55 mm</quantity>
|
||
; female: length
|
||
<quantity box="[228,330,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="7.06" pageId="5" pageNumber="6" unit="mm" value="7.06">7.06 mm</quantity>
|
||
, width
|
||
<quantity box="[413,517,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="3.89" pageId="5" pageNumber="6" unit="mm" value="3.89">3.89 mm</quantity>
|
||
), subparallel-sided, widening slightly towards the posterior third, not covering the last two tergites; only seven striae visible, the first six well-marked, the seventh faint, all slightly punctured and not reaching the base and the apex; intervals slightly convex, coarse setiferous punctures in two rows on 2
|
||
<superScript attach="none" box="[1379,1395,1410,1424]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
|
||
–7
|
||
<superScript attach="none" box="[1424,1437,1410,1424]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
intervals and one row on 1
|
||
<superScript attach="left" box="[458,469,1447,1461]" fontSize="6" pageId="5" pageNumber="6">st</superScript>
|
||
interval, decumbent seta brown-yellowish sub-erected and directed posteriad; lateral margin rounded in the shoulders, ending in deep furrow densely punctuated towards the scutellum. Scutellar striae hardly visible through the deep punctures in the basal region of first interval. Basal pore present. Just behind the shoulders 3 pores on the very margin. Umbilicate series with about 22 to 29 pores, not interrupted in the middle although pores are more separated from each other in that area; 1 subapical pore shifted inward in the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[355,368,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
interval; apex with 4 extra pores, in the lengthening of the imaginary 7
|
||
<superScript attach="left" box="[1157,1170,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
striae and 7
|
||
<superScript attach="none" box="[1304,1317,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
, 6
|
||
<superScript attach="left" box="[1344,1357,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
and 4
|
||
<superScript attach="none" box="[1424,1437,1626,1640]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
intervals; apical transparent membrane extending from sutural angle to the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[1325,1338,1662,1676]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
interval. The longitudinal humeral orange band extends from the base to the median part, extending from the 2
|
||
<superScript attach="left" box="[1284,1300,1698,1712]" fontSize="6" pageId="5" pageNumber="6">nd</superScript>
|
||
striae to the lateral margin; the median dark brown transverse band stops very near the lateral margin, or reaches it, extending backwards to the ending of the imaginary 8
|
||
<superScript attach="left" box="[636,649,1770,1784]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
striae; the oval sub transverse orange band in the last third reaches the apex and part of lateral margins; the dark brown basal marking reaches the basal margin towards the 5
|
||
<superScript attach="left" box="[1322,1335,1806,1820]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
|
||
interval, extending, diamond-shaped, backwards, but not reaching the apex (
|
||
<figureCitation box="[907,987,1843,1869]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="5" pageNumber="6">Fig. 1c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="6">
|
||
<emphasis box="[199,260,1879,1904]" italics="true" pageId="5" pageNumber="6">Legs.</emphasis>
|
||
Reddish-brown coloured; all tarsomeres pubescent, first three fore tarsomeres not dilated, fourth tarsomere slightly triangulate; fore tibiae on the outer edge with two blunt spines.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,1365]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<emphasis box="[199,367,1951,1977]" italics="true" pageId="5" pageNumber="6">Ventral surface</emphasis>
|
||
.Almost all underside orange-brownish; whole underside, including abdominal sternites coarsely punctured and pubescent (yellowish color), except gula, ventral and lateral sides of neck. Pubescence of prosternum and proepisternum slightly erected and directed anteriad; pubescence of the remaining ventral sclerites dressed and directed posteriad. Elytral epipleura moderately punctured, pubescence dressed and directed posteriad.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="7">
|
||
<emphasis box="[199,364,223,249]" italics="true" pageId="6" pageNumber="7">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[379,462,223,249]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="6" pageNumber="7">Fig. 3b</figureCitation>
|
||
). Median lobe dilated in the middle of ventral area, apical lamella short, wide and subacuminated at apex (dorsal view), downward (lateral view), ostium with one pair of thin sclerotized stripes in the dorsal median region, bending laterally downwards; endophallus with several minor sclerites just behind the dilated region; left paramere longer than wide, straight, rounded at apex, a tiny tooth in postero-dorsal edge.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="7">
|
||
<emphasis box="[199,389,367,393]" italics="true" pageId="6" pageNumber="7">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[403,485,367,393]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="6" pageNumber="7">Fig. 4b</figureCitation>
|
||
). Gonocoxite 1 subtriangular; gonocoxite 2 slender sickle shaped, apex slightly sharp; dorsal surface concave, strongly setose, four setae on dorsal half basal inner side and other four on dorsal half subapical outer side, several minor setae in a slight submarginal ridge along the half outer side; ventral surface slightly convex, smooth with some minor pubescence.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,472,510,536]" pageId="6" pageNumber="7">Intraspecific variation.</emphasis>
|
||
The range of variability observed in
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[882,1069,511,537]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="philippeorum" status="sp. n.">
|
||
<emphasis box="[882,1069,511,537]" italics="true" pageId="6" pageNumber="7">M. philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1076,1135,511,537]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<specimenCount box="[1150,1288,511,537]" pageId="6" pageNumber="7" type="generic">8 specimens</specimenCount>
|
||
) includes the darkness of the general colour of head+pronotum, the lateral extension of the median transverse dark brown band of elytra, reaching, or not, the lateral margin, extending more backwards, or not, or even in its middle width. There is some variability in the number of anterior submarginal setae of clypeus and in the number of umbilicate elytral pores (22 to 29). These values seem to be correlated with the elytral length (longer or shorter lengths correspond to larger or smaller values, respectively). There is also some variation in the location of the 4 apical extra pores, sometimes in the lengthening of the imaginary 8
|
||
<superScript attach="left" box="[693,706,726,740]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
|
||
interval or 7
|
||
<superScript attach="left" box="[849,862,726,740]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
|
||
striae and 7
|
||
<superScript attach="none" box="[998,1011,726,740]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
|
||
, 6
|
||
<superScript attach="left" box="[1039,1052,726,740]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
|
||
and 3
|
||
<superScript attach="left" box="[1120,1134,726,740]" fontSize="6" pageId="6" pageNumber="7">rd</superScript>
|
||
intervals or even 3
|
||
<superScript attach="left" box="[1348,1362,726,740]" fontSize="6" pageId="6" pageNumber="7">rd</superScript>
|
||
striae. The medium lobe of aedeagus is very conservative, but in
|
||
<specimenCount box="[812,914,763,789]" pageId="6" pageNumber="7" type="male">one male</specimenCount>
|
||
specimen there is an unusual asymmetry concerning the dorsal apical area of the left strip, having a sclerotized expansion towards left, not present in the other
|
||
<specimenCount box="[151,260,835,861]" pageId="6" pageNumber="7" type="male">six males</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="7" type="etymology">
|
||
<paragraph blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,334,871,897]" pageId="6" pageNumber="7">Etymology.</emphasis>
|
||
In a modest homage, this species is named in honour of two French entomologists namely Philippe Moretto (Association Catharsius,
|
||
<collectingCountry box="[526,607,907,933]" name="France" pageId="6" pageNumber="7">France</collectingCountry>
|
||
), a well-known expert on African Scarabaeinae, who collected the
|
||
<collectingCountry name="Burkina Faso" pageId="6" pageNumber="7">Burkina Faso</collectingCountry>
|
||
specimens and other
|
||
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[451,604,943,969]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[451,604,943,969]" italics="true" pageId="6" pageNumber="7">Macrocheilus</emphasis>
|
||
</taxonomicName>
|
||
specimens here studied and the late Philippe Annoyer (1967–2020) (Muséum de Toulouse), who organized the Expedition Sangha 2012 to the
|
||
<collectingCountry box="[941,1227,979,1005]" name="Central African Republic" pageId="6" pageNumber="7">Central African Republic</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,315,1015,1041]" pageId="6" pageNumber="7">Remarks.</emphasis>
|
||
The new species is more akin to
|
||
<taxonomicName authority="Basilewsky, 1970" authorityName="Basilewsky" authorityYear="1970" box="[689,1164,1015,1041]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="persimilis">
|
||
<emphasis box="[689,959,1015,1041]" italics="true" pageId="6" pageNumber="7">Macrocheilus persimilis</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[967,1164,1015,1041]" pageId="6" pageNumber="7" pagination="337 - 348" refId="ref25250" refString="Basilewsky, P. (1970) Coleopteres Carabidae africains nouveaux. XVIII. Revue de Zoologie et Botanique Africaines, 81 (3 - 4), 337 - 348." type="journal article" year="1970">Basilewsky, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Peringuey, 1904" authorityName="Peringuey" authorityYear="1904" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">
|
||
<emphasis box="[1219,1374,1015,1041]" italics="true" pageId="6" pageNumber="7">M. longicollis</emphasis>
|
||
<bibRefCitation author="Peringuey, L." pageId="6" pageNumber="7" pagination="167 - 206" refId="ref25855" refString="Peringuey, L. (1904). VII. Sixth contribution to the South Africa coleopterous fauna. Description of new species of Coleoptera in the collection of the South African Museum. Annals of the South African Museum, 3, 167 - 206." type="journal article" year="1904">Péringuey, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
than to the other African species by the elytral colour pattern (cf.
|
||
<figureCitation box="[1021,1098,1051,1077]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="6" pageNumber="7">Fig. 1c</figureCitation>
|
||
<emphasis box="[1105,1128,1051,1076]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
|
||
<figureCitation box="[1135,1308,1051,1077]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="6" pageNumber="7">Figs. 9b and 9c</figureCitation>
|
||
). However,
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[151,464,1087,1113]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
|
||
<emphasis box="[151,464,1087,1113]" italics="true" pageId="6" pageNumber="7">Macrocheilus philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[473,538,1087,1113]" pageId="6" pageNumber="7">
|
||
<taxonomicNameLabel box="[473,538,1087,1113]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from the former species by the less transverse pronotum (cf.
|
||
<figureCitation box="[151,241,1123,1149]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="6" pageNumber="7">Figs. 1c</figureCitation>
|
||
and
|
||
<figureCitation box="[298,327,1123,1149]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="6" pageNumber="7">9e</figureCitation>
|
||
), the less acuminate apex of median lobe (cf.
|
||
<figureCitation box="[849,940,1123,1149]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="6" pageNumber="7">Figs. 3b</figureCitation>
|
||
and
|
||
<figureCitation box="[997,1039,1123,1149]" captionStart="FIGURE 13" captionStartId="27.[152,255,872,896]" captionTargetBox="[370,1262,159,835]" captionTargetId="figure-117@27.[365,1272,151,843]" captionTargetPageId="27" captionText="FIGURE 13. Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. persimilis, ZAM (TAC); b, M. scapularis (MRAC); c, M. spectandus, ZAM (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745311" httpUri="https://zenodo.org/record/7745311/files/figure.png" pageId="6" pageNumber="7">13a</figureCitation>
|
||
) and different shape of labrum (cf.
|
||
<figureCitation box="[151,239,1159,1185]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="6" pageNumber="7">Figs. 2c</figureCitation>
|
||
and
|
||
<figureCitation box="[292,323,1159,1185]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="6" pageNumber="7">6b</figureCitation>
|
||
), and from the latter species by the less slender body shape, the bigger size, the more transverse pronotum (cf.
|
||
<figureCitation box="[274,363,1195,1221]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="6" pageNumber="7">Figs. 1c</figureCitation>
|
||
and
|
||
<figureCitation box="[417,448,1195,1221]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="6" pageNumber="7">9b</figureCitation>
|
||
), the less securiform last maxillary palpomere, the narrower transverse dark brown band (cf.
|
||
<figureCitation box="[196,285,1231,1257]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="6" pageNumber="7">Figs. 1c</figureCitation>
|
||
and
|
||
<figureCitation box="[340,369,1231,1257]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="6" pageNumber="7">9b</figureCitation>
|
||
) and different shapes of median lobe of aedeagus (cf.
|
||
<figureCitation box="[978,1068,1231,1257]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="6" pageNumber="7">Figs. 3b</figureCitation>
|
||
and
|
||
<figureCitation box="[1124,1152,1231,1257]" captionStart="FIGURE 7" captionStartId="19.[152,255,1531,1555]" captionTargetBox="[163,1424,429,1485]" captionTargetId="figure-63@19.[151,1436,421,1492]" captionTargetPageId="19" captionText="FIGURE 7.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. dorsalis, ZAM (TAC); b, M. elegantulus, DOC? (MRAC); c, M. labrosus, RCA (PSC); d, M. longicollis, MOZ (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745297" httpUri="https://zenodo.org/record/7745297/files/figure.png" pageId="6" pageNumber="7">7d</figureCitation>
|
||
) and labrum (cf.
|
||
<figureCitation box="[1347,1436,1231,1257]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="6" pageNumber="7">Figs. 2c</figureCitation>
|
||
and
|
||
<figureCitation box="[199,239,1267,1293]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="6" pageNumber="7">10a</figureCitation>
|
||
). The new species differs from the remaining African species by elytra colour pattern and the median lobe shape, among other external morphological characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,741,1338,1365]" pageId="6" pageNumber="7" type="distribution">
|
||
<paragraph blockId="6.[151,1437,151,1365]" box="[199,741,1338,1365]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[199,345,1338,1364]" pageId="6" pageNumber="7">Distribution</emphasis>
|
||
.
|
||
<collectingCountry box="[359,450,1339,1365]" name="Senegal" pageId="6" pageNumber="7">Senegal</collectingCountry>
|
||
,
|
||
<collectingCountry box="[462,612,1339,1365]" name="Burkina Faso" pageId="6" pageNumber="7">Burkina Faso</collectingCountry>
|
||
and
|
||
<collectingCountry box="[667,738,1339,1365]" name="Benin" pageId="6" pageNumber="7">Benin</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |